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<mods:title id="37FCB4A0F270E759779ED769444C71E9">Orbitolina Daviesi Hofker, 1966: A Forgotten Orbitolinidae From The Thanetian Of Pakistan And Its Taxonomic Status</mods:title>
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<treatment id="038C87C90749FF9A46B4D0E244D2FD02" LSID="urn:lsid:plazi:treatment:038C87C90749FF9A46B4D0E244D2FD02" httpUri="http://treatment.plazi.org/id/038C87C90749FF9A46B4D0E244D2FD02" lastPageNumber="46" pageId="5" pageNumber="46">
<subSubSection id="C33F65540749FF9A46B4D0E24524FBB6" box="[151,618,1018,1043]" pageId="5" pageNumber="46" type="nomenclature">
<paragraph id="8B9A36DF0749FF9A46B4D0E24524FBB6" blockId="5.[151,771,687,1043]" box="[151,618,1018,1043]" pageId="5" pageNumber="46">
<taxonomicName id="4C254D5C0749FF9A46B4D0E246A2FBB6" ID-CoL="82SBR" authority="(Hofker, 1966)" authorityName="SCHLAGINTWEIT" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT" baseAuthorityYear="1966" box="[151,492,1018,1043]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="daviesi" status="comb. nov.">
<emphasis id="B951EACD0749FF9A46B4D0E2460BFBB7" bold="true" box="[151,325,1019,1043]" italics="true" pageId="5" pageNumber="46">Karsella daviesi</emphasis>
(
<bibRefCitation id="EFB44B2E0749FF9A4770D0E346AAFBB6" author="Hofker, J. Jr." box="[339,484,1018,1042]" pageId="5" pageNumber="46" pagination="1 - 34" refId="ref5015" refString="Hofker, J. Jr., 1966. Studies on the family Orbitolinidae. Palaeontographica, Abt. A, 126: 1 - 34." type="journal article" year="1966">Hofker, 1966</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26257B60749FF9A47D1D0E34524FBB6" box="[498,618,1018,1042]" pageId="5" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C33F65540749FF9A46B4D725466BF96D" pageId="5" pageNumber="46" type="reference_group">
<paragraph id="8B9A36DF0749FF9A46B4D7254656FB33" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
1941
<taxonomicName id="4C254D5C0749FF9A46F7D7254627FBF0" box="[212,361,1084,1108]" class="Globothalamea" family="Orbitolinidae" genus="Orbitolina" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B951EACD0749FF9A46F7D725460AFBF0" box="[212,324,1084,1108]" italics="true" pageId="5" pageNumber="46">Orbitolina</emphasis>
sp.
</taxonomicName>
Davies, p. 317 (fide
<bibRefCitation id="EFB44B2E0749FF9A447ED72545A2FBF0" author="Hofker, J. Jr." box="[605,748,1084,1108]" pageId="5" pageNumber="46" pagination="1 - 34" refId="ref5015" refString="Hofker, J. Jr., 1966. Studies on the family Orbitolinidae. Palaeontographica, Abt. A, 126: 1 - 34." type="journal article" year="1966">Hofker, 1966</bibRefCitation>
). 1966
<taxonomicName id="4C254D5C0749FF9A46F6D74446D6FBD1" ID-CoL="835D4" authority="SCHLAGINTWEIT, 2021" authorityName="SCHLAGINTWEIT" authorityYear="2021" box="[213,408,1117,1141]" class="Globothalamea" family="Orbitolinidae" genus="Orbitolina" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="daviesi" status="sp. nov.">
<emphasis id="B951EACD0749FF9A46F6D74446D6FBD1" box="[213,408,1117,1141]" italics="true" pageId="5" pageNumber="46">Orbitolina daviesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257B60749FF9A4783D7474694FBD1" box="[416,474,1118,1141]" pageId="5" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
Hofker, p. 12, pl. 2, figs. 10-14, pl. 3.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D7B9468CFB7D" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
1997
<taxonomicName id="4C254D5C0749FF9A46F7D7B946EEFB1C" authority="SCHLAGINTWEIT, 2021" authorityName="SCHLAGINTWEIT" authorityYear="2021" box="[212,416,1184,1208]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="hottingeri" status="n. gen., n. sp.">
<emphasis id="B951EACD0749FF9A46F7D7B946EEFB1C" box="[212,416,1184,1208]" italics="true" pageId="5" pageNumber="46">Karsella hottingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257B60749FF9A478BD7B94577FB13" box="[424,569,1184,1208]" pageId="5" pageNumber="46" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Sirel, p. 210, pl. 1, figs. 1-10, pl. 2, figs. 1-4.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D7FA4787FA98" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
Non 2005
<taxonomicName id="4C254D5C0749FF9A472AD7FB46C4FB5E" box="[265,394,1250,1274]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B951EACD0749FF9A472AD7FB462AFB5E" box="[265,356,1250,1274]" italics="true" pageId="5" pageNumber="46">Karsella</emphasis>
sp.
</taxonomicName>
Tentor and Venturini, fig. 8, 9?, 10-12 [
<taxonomicName id="4C254D5C0749FF9A46CFD61A47F0FA98" authority="(Hottinger &amp; Drobne)" baseAuthorityName="Hottinger &amp; Drobne" baseAuthorityYear="1980" class="Globothalamea" family="Orbitolinidae" genus="Schroedericonus" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="turriculus">
<emphasis id="B951EACD0749FF9A46CFD61A4558FABF" box="[236,534,1283,1307]" italics="true" pageId="5" pageNumber="46">Schroedericonus turriculus</emphasis>
(Hottinger &amp; Drobne)
</taxonomicName>
].
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D65C45AEFAF9" blockId="5.[151,772,1084,1737]" box="[151,736,1349,1373]" pageId="5" pageNumber="46">
2006
<taxonomicName id="4C254D5C0749FF9A46F7D65C4696FAF9" authority="Sirel" authorityName="Sirel" authorityYear="1999" box="[212,472,1349,1373]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="hottingeri">
<emphasis id="B951EACD0749FF9A46F7D65C46D1FAF9" box="[212,415,1349,1373]" italics="true" pageId="5" pageNumber="46">Karsella hottingeri</emphasis>
Sirel
</taxonomicName>
Hottinger, fig. 41K-L.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D67F46DEFA3B" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
2010
<taxonomicName id="4C254D5C0749FF9A46FAD67F46A7FADA" authority="Sirel" authorityName="Sirel" authorityYear="1999" box="[217,489,1382,1406]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="hottingeri">
<emphasis id="B951EACD0749FF9A46FAD67F46E4FADA" box="[217,426,1382,1406]" italics="true" pageId="5" pageNumber="46">Karsella hottingeri</emphasis>
Sirel
</taxonomicName>
Afzal, p. 115, pl. 14, figs. 5-10,?11-13?, 14.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D6B04601FA46" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
2014 “
<taxonomicName id="4C254D5C0749FF9A46C0D6B0456BFA65" authority="J. Hofker" authorityName="J. Hofker" authorityYear="1966" box="[227,549,1449,1473]" class="Globothalamea" family="Orbitolinidae" genus="Orbitolina" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="daviesi">
<emphasis id="B951EACD0749FF9A46C0D6B0461DFA65" box="[227,339,1449,1473]" italics="true" pageId="5" pageNumber="46">Orbitolina</emphasis>
<emphasis id="B951EACD0749FF9A474BD6B046FAFA65" box="[360,436,1449,1473]" italics="true" pageId="5" pageNumber="46">daviesi</emphasis>
J. Hofker
</taxonomicName>
jun. Hottinger, p. 51, pl. 4.9, fig. 2.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D6F24550F980" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
2018
<taxonomicName id="4C254D5C0749FF9A46FBD6F246A8F9A7" authority="Sirel" authorityName="Sirel" authorityYear="1999" box="[216,486,1515,1539]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="hottingeri">
<emphasis id="B951EACD0749FF9A46FBD6F246E6F9A7" box="[216,424,1515,1539]" italics="true" pageId="5" pageNumber="46">Karsella hottingeri</emphasis>
Sirel
</taxonomicName>
Cruz-Abad, p. 84, fig. 78A-E (reproduction of Sirel, 1997).
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D53447FFF9C2" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
2018 “
<taxonomicName id="4C254D5C0749FF9A46C3D534454DF9E1" authority="Hofker" authorityName="J. Hofker" authorityYear="1966" box="[224,515,1581,1605]" class="Globothalamea" family="Orbitolinidae" genus="Orbitolina" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="daviesi">
<emphasis id="B951EACD0749FF9A46C3D534461EF9E1" box="[224,336,1581,1605]" italics="true" pageId="5" pageNumber="46">Orbitolina</emphasis>
<emphasis id="B951EACD0749FF9A4746D53446FEF9E1" box="[357,432,1581,1605]" italics="true" pageId="5" pageNumber="46">daviesi</emphasis>
Hofker
</taxonomicName>
Cruz-Abad, fig. 78F-H.
</paragraph>
<paragraph id="8B9A36DF0749FF9A46B4D576466BF96D" blockId="5.[151,772,1084,1737]" pageId="5" pageNumber="46">
2020
<taxonomicName id="4C254D5C0749FF9A46F4D57646AAF923" ID-CoL="82SBS" authority="Sirel" authorityName="Sirel" authorityYear="1999" box="[215,484,1647,1671]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="hottingeri">
<emphasis id="B951EACD0749FF9A46F4D57646E9F923" box="[215,423,1647,1671]" italics="true" pageId="5" pageNumber="46">Karsella hottingeri</emphasis>
Sirel
</taxonomicName>
Schlagintweit, figs. 1, 2A-B (reproductions of Sirel, 1997,
<bibRefCitation id="EFB44B2E0749FF9A4400D589459EF90C" author="Hottinger, L." box="[547,720,1680,1704]" pageId="5" pageNumber="46" pagination="2006" refId="ref5042" refString="Hottinger, L., 2006. Illustrated glossary of terms used in foraminiferal research. Carnets de Geologie / Notebooks on Geology, Memoir 2006 / 02 (CG 2006 _ M 02)." type="journal article" year="2006">Hottinger, 2006</bibRefCitation>
, and
<bibRefCitation id="EFB44B2E0749FF9A46B4D5A84654F96D" author="Afzal, J." box="[151,282,1713,1737]" pageId="5" pageNumber="46" pagination="1 - 291" refId="ref4381" refString="Afzal, J., 2010. Evolution of Larger Benthic Foraminifera during the Paleocene-Early Eocene interval in the East Tethys (Indus Basin, Pakistan). PhD Thesis University of Leicester: 1 - 291. https: // core. ac. uk / display / 2796424" type="book chapter" year="2010">Afzal, 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F65540749FF9A46B4D5ED43AEFDA5" pageId="5" pageNumber="46" type="description">
<paragraph id="8B9A36DF0749FF9A46B4D5ED43AEFDA5" blockId="5.[151,772,1780,2068]" lastBlockId="5.[816,1437,163,513]" pageId="5" pageNumber="46">
<emphasis id="B951EACD0749FF9A46B4D5ED466AF8A8" bold="true" box="[151,292,1780,1804]" pageId="5" pageNumber="46">Description:</emphasis>
Test large, high-conical to slightly cylindroconical with moderately convex base. The megalospheric embryo is situated in a central position at the rounded apex and presents a diameter of up to 0.5 mm. It consists of an irregular ellpisoidal proloculus, a large deuteroconch with partitions (supraembryonic zone) and a few broad peri-embryonic chambers (“subembryonic zone”) that are sporadically subdivided. The neanic stage consists of up to 27 uniserial chambers subdivided into marginal, broad radial, and reduced central zones. There are 5-7 chambers per the last millimeter of the axial cone length. The marginal zone is complex, displaying several orders of beams and rafters. The main partitions (septula), are triangular in axial section, increase in thickeness towards the cone centre and alternate in position between subsequent chambers. In transverse sections, the septula display a zig-zag pattern with associated radially arranged foramina piercing the septa obliquely. The septula form an irregular network displaying oblique stolon axes in the reduced central zone. Wall agglutinated.
</paragraph>
</subSubSection>
<subSubSection id="C33F65540749FF9A4513D13244D2FD02" pageId="5" pageNumber="46" type="distribution">
<paragraph id="8B9A36DF0749FF9A4513D13244D2FD02" blockId="5.[816,1436,555,678]" pageId="5" pageNumber="46">
<emphasis id="B951EACD0749FF9A4513D1324487FDE7" bold="true" box="[816,969,555,579]" pageId="5" pageNumber="46">Occurrences:</emphasis>
<taxonomicName id="4C254D5C0749FF9A45F5D13243BAFDE7" authority="(Hofker)" authorityName="SCHLAGINTWEIT" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT" baseAuthorityYear="1966" box="[982,1268,555,579]" class="Globothalamea" family="Orbitolinidae" genus="Karsella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="46" phylum="Foraminifera" rank="species" species="daviesi" status="comb. nov.">
<emphasis id="B951EACD0749FF9A45F5D13243C4FDE7" box="[982,1162,555,579]" italics="true" pageId="5" pageNumber="46">Karsella daviesi</emphasis>
(Hofker)
</taxonomicName>
<taxonomicNameLabel id="A26257B60749FF9A4322D1324231FDE7" box="[1281,1407,555,579]" pageId="5" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
is known from the Thanetian of
<collectingCountry id="F332764F0749FF9A4293D15543B3FDC0" box="[1200,1277,588,612]" name="Turkey" pageId="5" pageNumber="46">Turkey</collectingCountry>
(Sirel, 1997), and
<collectingCountry id="F332764F0749FF9A45ABD17444ACFD21" box="[904,994,621,645]" name="Pakistan" pageId="5" pageNumber="46">Pakistan</collectingCountry>
(
<bibRefCitation id="EFB44B2E0749FF9A45D5D17443C2FD21" author="Hofker, J. Jr." box="[1014,1164,621,645]" pageId="5" pageNumber="46" pagination="1 - 34" refId="ref5015" refString="Hofker, J. Jr., 1966. Studies on the family Orbitolinidae. Palaeontographica, Abt. A, 126: 1 - 34." type="journal article" year="1966">Hofker, 1966</bibRefCitation>
;
<bibRefCitation id="EFB44B2E0749FF9A42B8D1744206FD21" author="Hottinger, L." box="[1179,1352,621,645]" pageId="5" pageNumber="46" pagination="2006" refId="ref5042" refString="Hottinger, L., 2006. Illustrated glossary of terms used in foraminiferal research. Carnets de Geologie / Notebooks on Geology, Memoir 2006 / 02 (CG 2006 _ M 02)." type="journal article" year="2006">Hottinger, 2006</bibRefCitation>
;
<bibRefCitation id="EFB44B2E0749FF9A4379D17444C5FD02" author="Afzal, J." pageId="5" pageNumber="46" pagination="1 - 291" refId="ref4381" refString="Afzal, J., 2010. Evolution of Larger Benthic Foraminifera during the Paleocene-Early Eocene interval in the East Tethys (Indus Basin, Pakistan). PhD Thesis University of Leicester: 1 - 291. https: // core. ac. uk / display / 2796424" type="book chapter" year="2010">Afzal, 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>