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<mods:title id="B01EDA6E7CDD58A79FBB0F19B2A3656F">Three new synonyms within the flower chafer genus Goliathopsis Janson, 1881 (Coleoptera: Scarabaeidae: Cetoniinae) from China</mods:title>
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<taxonomicName id="4C254D52CE00FFBAFF0BF8E7FD3DF8DC" authority="(Westwood, 1873)" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[151,655,1805,1832]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFF0BF8E7FD3DF8DC" bold="true" box="[151,655,1805,1832]" pageId="7" pageNumber="98">
<emphasis id="B951EAC3CE00FFBAFF0BF8E7FE18F8D3" bold="true" box="[151,426,1805,1832]" italics="true" pageId="7" pageNumber="98">Goliathopsis despectus</emphasis>
(
<bibRefCitation id="EFB44B20CE00FFBAFE25F8E7FD34F8DC" author="Westwood, J. O." box="[441,646,1805,1831]" pageId="7" pageNumber="98" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
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<paragraph id="8B9A36D1CE00FFBAFF0BF8DBFDC2F8B7" blockId="7.[151,655,1805,1868]" box="[151,624,1840,1868]" pageId="7" pageNumber="98">
(
<figureCitation id="131E2A54CE00FFBAFF03F8DBFF5DF8B0" box="[159,239,1841,1867]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="7" pageNumber="98">Figs. 6</figureCitation>
,
<figureCitation id="131E2A54CE00FFBAFF67F8DBFEABF8B0" box="[251,281,1841,1867]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="7" pageNumber="98">14</figureCitation>
,
<figureCitation id="131E2A54CE00FFBAFEB9F8DAFEDFF8B0" box="[293,365,1840,1867]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="7" pageNumber="98">2425</figureCitation>
,
<figureCitation id="131E2A54CE00FFBAFEE5F8DBFE70F8B0" box="[377,450,1841,1867]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="7" pageNumber="98">4849</figureCitation>
,
<figureCitation id="131E2A54CE00FFBAFE51F8DBFD83F8B0" box="[461,561,1841,1867]" captionStart-0="FIGURES 109114" captionStart-1="FIGURES 115121" captionStartId-0="25.[151,264,1140,1165]" captionStartId-1="26.[151,264,1159,1184]" captionTargetBox-0="[157,1430,185,1105]" captionTargetBox-1="[155,1422,187,1125]" captionTargetId-0="figure@25.[151,1435,181,1115]" captionTargetId-1="figure@26.[151,1435,181,1133]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 109114. Goliathopsis despectus (Westwood, 1873). 109111, female paralectotype in OXUM and labels (photograph by Amoret Spooner); 112114, non-type female in MHNH from Combodia and labels." captionText-1="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi-0="http://doi.org/10.5281/zenodo.3884757" figureDoi-1="http://doi.org/10.5281/zenodo.3884759" httpUri-0="https://zenodo.org/record/3884757/files/figure.png" httpUri-1="https://zenodo.org/record/3884759/files/figure.png" pageId="7" pageNumber="98">109121</figureCitation>
,
<figureCitation id="131E2A54CE00FFBAFDA1F8DBFDDAF8B7" box="[573,616,1841,1868]" captionStart="FIGURE 164" captionStartId="33.[151,250,1616,1641]" captionTargetBox="[159,1427,209,1579]" captionTargetId="figure@33.[151,1436,181,1591]" captionTargetPageId="33" captionText="FIGURE 164. Known distribution of Goliathopsis Janson, 1881 species. Syncretic markers indicate identical or very close localities; type localities of the known species are marked, that of G. despectus (Westwood, 1873), G. cervus Janson, 1881, and G. esquiroli Pouillaude, 1913 are missing because the precise type localities are unknown." figureDoi="http://doi.org/10.5281/zenodo.3884777" httpUri="https://zenodo.org/record/3884777/files/figure.png" pageId="7" pageNumber="98">164</figureCitation>
)
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<taxonomicName id="4C254D52CE00FFBAFF0BF885FDF2F87C" authority="Westwood, 1873: 32" authorityName="Westwood" authorityPageNumber="32" authorityYear="1873" box="[151,576,1902,1927]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFF0BF885FED8F87C" box="[151,362,1903,1927]" italics="true" pageId="7" pageNumber="98">Pilinurgus despectus</emphasis>
<bibRefCitation id="EFB44B20CE00FFBAFEECF884FDF2F87C" author="Westwood, J. O." box="[368,576,1902,1927]" pageId="7" pageNumber="98" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873: 32</bibRefCitation>
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(
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locality:
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[=
<collectingCountry id="F3327641CE00FFBAFCB6F884FC3EF87C" box="[810,908,1902,1927]" name="Thailand" pageId="7" pageNumber="98">Thailand</collectingCountry>
]), plate IX, fig.
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;
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<bibRefCitation id="EFB44B20CE00FFBAFBC3F885FAB8F87C" author="Berge, A." box="[1119,1290,1902,1927]" pageId="7" pageNumber="98" pagination="113 - 163" refId="ref19490" refString="Berge, A. (1884) Enumeration des Cetonides decrits depuis la publication du Catalogue de MM. Gemminger &amp; de Harold. An- nales de la Societe Entomologique de Belgique, 28, 113 - 163." type="journal article" year="1884">Bergé 1884: 158</bibRefCitation>
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.
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<treatmentCitation id="0A8410C0CE00FFBAFF0BF864FDF3F85D" author="Janson, O. E." box="[151,577,1933,1958]" page="610" pageId="7" pageNumber="98" year="1881">
<taxonomicName id="4C254D52CE00FFBAFF0BF864FDF3F85D" authority=": Janson 1881: 610" authorityName=": Janson" authorityPageNumber="610" authorityYear="1881" box="[151,577,1933,1958]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFF0BF864FE32F85D" box="[151,384,1934,1958]" italics="true" pageId="7" pageNumber="98">Goliathopsis despectus</emphasis>
:
<bibRefCitation id="EFB44B20CE00FFBAFE12F864FDF3F85D" author="Janson, O. E." box="[398,577,1933,1958]" pageId="7" pageNumber="98" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson 1881: 610</bibRefCitation>
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;
<treatmentCitation id="0A8410C0CE00FFBAFDD7F867FCA3F85D" author="Schoch, G." box="[587,785,1933,1958]" page="395" pageId="7" pageNumber="98" year="1896">
<bibRefCitation id="EFB44B20CE00FFBAFDD7F867FCA3F85D" author="Schoch, G." box="[587,785,1933,1958]" pageId="7" pageNumber="98" pagination="378 - 395" refId="ref21766" refString="Schoch, G. (1896 b) Nachtrag II zu Schoch: Die Genera und Species meiner Cetoniden-sammlung. Mitteilungen der Schweizeri- schen Entomologischen Gesellschaft, 9 (9), 378 - 395." type="journal article" year="1896">Schoch 1896b: 395</bibRefCitation>
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;
<treatmentCitation id="0A8410C0CE00FFBAFC85F864FC7AF85D" author="Arrow, G. J." box="[793,968,1933,1958]" page="206" pageId="7" pageNumber="98" year="1910">
<bibRefCitation id="EFB44B20CE00FFBAFC85F864FC7AF85D" author="Arrow, G. J." box="[793,968,1933,1958]" pageId="7" pageNumber="98" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow 1910: 206</bibRefCitation>
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, plate II, figs.
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,
<specimenCount id="9D23FD58CE00FFBAFB16F864FB03F85E" box="[1162,1201,1934,1958]" pageId="7" pageNumber="98" type="female">3 ♀</specimenCount>
;
<treatmentCitation id="0A8410C0CE00FFBAFB21F864FADEF85D" author="Janson, O. E." box="[1213,1388,1934,1958]" page="110" pageId="7" pageNumber="98" year="1917">
<bibRefCitation id="EFB44B20CE00FFBAFB21F864FADEF85D" author="Janson, O. E." box="[1213,1388,1934,1958]" pageId="7" pageNumber="98" pagination="102 - 111" refId="ref20489" refString="Janson, O. E. (1917) Additions to the knowledge of the Cetoniinae of British India. Transactions of the Entomological Society of London, 65 (1), 102 - 111. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01404. x" type="journal article" year="1917">Janson 1917: 110</bibRefCitation>
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(ho- lotype
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in OXUM);
<treatmentCitation id="0A8410C0CE00FFBAFE3DF846FD33F83E" author="Schenkling, S." box="[417,641,1964,1989]" page="364" pageId="7" pageNumber="98" year="1921">
<bibRefCitation id="EFB44B20CE00FFBAFE3DF846FD33F83E" author="Schenkling, S." box="[417,641,1964,1989]" pageId="7" pageNumber="98" pagination="1 - 431" refId="ref21671" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, pp. 1 - 431 pp. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1921">Schenkling 1921: 364</bibRefCitation>
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;
<treatmentCitation id="0A8410C0CE00FFBAFD16F846FC9CF83F" author="Antoine, P." box="[650,814,1964,1989]" page="9" pageId="7" pageNumber="98" year="1991">
<bibRefCitation id="EFB44B20CE00FFBAFD16F846FC9CF83F" author="Antoine, P." box="[650,814,1964,1989]" pageId="7" pageNumber="98" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine 1991: 9</bibRefCitation>
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(in key), plate I, fig. 2 (parameres), plate III, fig.
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;
<treatmentCitation id="0A8410C0CE00FFBAFACCF846FE91F818" author="Krajcik, M." page="39" pageId="7" pageNumber="98" year="1999">
<bibRefCitation id="EFB44B20CE00FFBAFACCF846FE91F818" author="Krajcik, M." pageId="7" pageNumber="98" refId="ref20545" refString="Krajcik, M. (1999) Cetoniidae of the World Catalogue. Part II. Typos Studio, Most, 72 pp." type="book" year="1999">Krajčík 1999: 39</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410C0CE00FFBAFEB2F820FE5AF818" author="Krajcik, M." box="[302,488,1994,2019]" page="116" pageId="7" pageNumber="98" year="2012">
<bibRefCitation id="EFB44B20CE00FFBAFEB2F820FE5AF818" author="Krajcik, M." box="[302,488,1994,2019]" pageId="7" pageNumber="98" pagination="1 - 278" refId="ref20603" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 116</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36D1CE0FFFB5FF0BFF73FA2EFF34" blockId="8.[151,1436,152,244]" pageId="8" pageNumber="99">
<treatmentCitationGroup id="AB3511FFCE0FFFB5FF0BFF73FA2EFF34" pageId="8" pageNumber="99">
<treatmentCitation id="0A8410C0CE0FFFB5FF0BFF73FDF0FF4A" author="Gestro, R." box="[151,578,153,177]" page="118" pageId="8" pageNumber="99" year="1888">
<taxonomicName id="4C254D52CE0FFFB5FF0BFF73FDF0FF4A" authority="Gestro, 1888: 118" authorityName="Gestro" authorityPageNumber="118" authorityYear="1888" box="[151,578,153,177]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0FFFB5FF0BFF73FE31FF4A" box="[151,387,153,177]" italics="true" pageId="8" pageNumber="99">Goliathopsis capreolus</emphasis>
<bibRefCitation id="EFB44B20CE0FFFB5FE17FF73FDF0FF4A" author="Gestro, R." box="[395,578,153,177]" pageId="8" pageNumber="99" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro, 1888: 118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(type locality: Farm Caves, Moulmein,
<collectingCountry id="F3327641CE0FFFB5FC43FF73FBF4FF4A" box="[991,1094,153,177]" name="Myanmar" pageId="8" pageNumber="99">Myanmar</collectingCountry>
), fig.
<specimenCount id="9D23FD58CE0FFFB5FB1EFF73FB27FF4B" box="[1154,1173,153,176]" pageId="8" pageNumber="99" type="male"></specimenCount>
(horn);
<treatmentCitation id="0A8410C0CE0FFFB5FB76FF73FA2EFF4A" author="Gestro, R." box="[1258,1436,152,177]" page="853" pageId="8" pageNumber="99" year="1891">
<bibRefCitation id="EFB44B20CE0FFFB5FB76FF73FA2EFF4A" author="Gestro, R." box="[1258,1436,152,177]" pageId="8" pageNumber="99" pagination="835 - 876" refId="ref20201" refString="Gestro, R. (1891) Viaggio di Leonardo Fea in Birmania regione vicine. XXXVII. Enumerazione delle Cetonie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 10, 835 - 876." type="journal article" year="1891">Gestro 1891: 853</bibRefCitation>
</treatmentCitation>
(visiting flower);
<treatmentCitation id="0A8410C0CE0FFFB5FEE3FF5DFDEDFF34" author="Pouillaude, I." box="[383,607,183,208]" page="421" pageId="8" pageNumber="99" year="1913">
<bibRefCitation id="EFB44B20CE0FFFB5FEE3FF5DFDEDFF34" author="Pouillaude, I." box="[383,607,183,208]" pageId="8" pageNumber="99" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude 1913: 421</bibRefCitation>
</treatmentCitation>
, fig. 2 (cephalic horns);
<treatmentCitation id="0A8410C0CE0FFFB5FCFFFF5DFB9AFF34" author="Krikken, J." box="[867,1064,183,208]" page="314" pageId="8" pageNumber="99" year="1977">
<bibRefCitation id="EFB44B20CE0FFFB5FCFFFF5DFB9AFF34" author="Krikken, J." box="[867,1064,183,208]" pageId="8" pageNumber="99" pagination="309 - 320" refId="ref20674" refString="Krikken, J. (1977). Notes on Asian cremastochiliform genera, with descriptions of two new species (Coleoptera: Cetoniidae). Zoologische Mededelingen, 50 (21), 309 - 320." type="journal article" year="1977">Krikken 1977: 314</bibRefCitation>
</treatmentCitation>
(three female
<typeStatus id="549E8873CE0FFFB5FB5FFF52FAAEFF2B" box="[1219,1308,184,208]" pageId="8" pageNumber="99" type="syntype">syntypes</typeStatus>
in RMNH);
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36D1CE0FFFB5FF5BFF31FD9FFF0F" blockId="8.[151,1436,152,244]" box="[199,557,219,244]" pageId="8" pageNumber="99">
Synonymized by
<bibRefCitation id="EFB44B20CE0FFFB5FEEBFF36FD9BFF0F" author="Arrow, G. J." box="[375,553,219,244]" pageId="8" pageNumber="99" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow 1910: 206</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5A6659CE0FFFB5FF0BF884FAE1F87C" ID-DOI="http://doi.org/10.5281/zenodo.3884735" ID-Zenodo-Dep="3884735" box="[151,1363,1902,1927]" httpUri="https://zenodo.org/record/3884735/files/figure.png" pageId="8" pageNumber="99" startId="8.[151,264,1902,1927]" targetBox="[186,1367,278,1868]" targetPageId="8">
<paragraph id="8B9A36D1CE0FFFB5FF0BF884FAE1F87C" blockId="8.[151,1363,1902,1927]" box="[151,1363,1902,1927]" pageId="8" pageNumber="99">
<emphasis id="B951EAC3CE0FFFB5FF0BF884FEE6F87C" bold="true" box="[151,340,1902,1927]" pageId="8" pageNumber="99">FIGURES 5457.</emphasis>
Habitus of
<taxonomicName id="4C254D52CE0FFFB5FE57F885FC92F87C" authority="Janson, 1881" authorityName="in Janson" authorityYear="1881" box="[459,800,1903,1927]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE0FFFB5FE57F885FD27F87C" box="[459,661,1903,1927]" italics="true" pageId="8" pageNumber="99">Goliathopsis cervus</emphasis>
<bibRefCitation id="EFB44B20CE0FFFB5FD07F885FC92F87C" author="Janson, O. E." box="[667,800,1903,1927]" pageId="8" pageNumber="99" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
</taxonomicName>
(dorsal and ventral view). 5455, male; 5657, female.
</paragraph>
</caption>
<subSubSection id="C33F655ACE0EFFB7FF0BFF73FEB1FCEC" lastPageId="10" lastPageNumber="101" pageId="9" pageNumber="100" type="materials_examined">
<paragraph id="8B9A36D1CE0EFFB4FF0BFF73FDEBFCEC" blockId="9.[151,1437,152,2015]" pageId="9" pageNumber="100">
<emphasis id="B951EAC3CE0EFFB4FF0BFF73FEF4FF48" bold="true" box="[151,326,152,179]" pageId="9" pageNumber="100">Type material.</emphasis>
When
<bibRefCitation id="EFB44B20CE0EFFB4FE04FF73FDD2FF48" author="Westwood, J. O." box="[408,608,153,179]" pageId="9" pageNumber="100" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood (1873)</bibRefCitation>
described
<taxonomicName id="4C254D52CE0EFFB4FD40FF73FC52FF48" authorityName=": Janson" authorityYear="1881" box="[732,992,153,179]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FD40FF73FC52FF48" box="[732,992,153,179]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
</taxonomicName>
, no cephalic horns were mentioned for the specimens, and these specimens were deposited in three type repositories (
<emphasis id="B951EAC3CE0EFFB4FB99FF54FB9AFF2C" box="[1029,1064,190,215]" italics="true" pageId="9" pageNumber="100">i.e.</emphasis>
Mus. Britann., Oxoniae
<emphasis id="B951EAC3CE0EFFB4FAA1FF54FAE3FF2C" box="[1341,1361,190,215]" italics="true" pageId="9" pageNumber="100">et</emphasis>
W. W. Saunders). It is implied that at least three
<typeStatus id="549E8873CE0EFFB4FDC2FF0BFD72FF00" box="[606,704,225,251]" pageId="9" pageNumber="100" type="syntype">syntypes</typeStatus>
should exist and they are all female. During our visiting in Europe museums, we attempted to confirm all of the three type repositories for this species. Whithin which,
<collectionCode id="ED34AE14CE0EFFB4FA8BFEEFFAD9FEE4" box="[1303,1387,261,287]" pageId="9" pageNumber="100">BMNH</collectionCode>
and
<collectionCode id="ED34AE14CE0EFFB4FF0BFEC3FF5FFEB8" box="[151,237,297,323]" pageId="9" pageNumber="100">OXUM</collectionCode>
each stores one female specimen, both specimens have the type label. Whereas the
<typeStatus id="549E8873CE0EFFB4FB2EFEC3FABCFEB8" box="[1202,1294,297,323]" pageId="9" pageNumber="100" type="syntype">syntype</typeStatus>
(s) originally deposited in Saunders Collection are hitherto unknown. In
<collectionCode id="ED34AE14CE0EFFB4FCB6FEA7FC32FE9C" box="[810,896,333,359]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
a female attached with label data “Sharp collection” and “Oberthür collection” was found; these labels agree with the transfer of
<taxonomicName id="4C254D52CE0EFFB4FBBCFE9BFB2BFE70" authorityName="Leach" authorityYear="1815" box="[1056,1177,369,395]" class="Insecta" family="Cetoniinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="family">Cetoniinae</taxonomicName>
specimens in Saunders Collection according to
<bibRefCitation id="EFB44B20CE0EFFB4FE3AFE7FFDC0FE54" author="Horn, W. &amp; Kahle, I. &amp; Friese, G. &amp; Gaedike, R." box="[422,626,405,431]" pageId="9" pageNumber="100" refId="ref20345" refString="Horn, W., Kahle, I., Friese, G. &amp; Gaedike, R. (1990) Collectiones entomologicae. Ein Kompendium uber den Verbleib ento- mologischer Sammlungen der Welt bis 1960. Teil II: L bis Z. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 353 pp." type="book" year="1990">
Horn
<emphasis id="B951EAC3CE0EFFB4FE7BFE7CFD92FE54" box="[487,544,405,431]" italics="true" pageId="9" pageNumber="100">et al.</emphasis>
(1990)
</bibRefCitation>
and
<bibRefCitation id="EFB44B20CE0EFFB4FD35FE7FFCC4FE4B" author="Cambefort, Y." box="[681,886,405,432]" pageId="9" pageNumber="100" refId="ref19726" refString="Cambefort, Y. (2006) Des Coleopteres, des Collections et des Hommes. Collection Archives, Museum National d'Histoire Naturelle, Paris, 375 pp. https: // doi. org / 10.4000 / books. mnhn. 2265" type="book" year="2006">Cambefort (2006)</bibRefCitation>
. Furthermore, the locality “
<collectingCountry id="F3327641CE0EFFB4FB36FE7FFB5AFE54" box="[1194,1256,405,431]" name="Thailand" pageId="9" pageNumber="100">Siam</collectingCountry>
” on the label of this specimen is accordant with the type locality indicated by
<bibRefCitation id="EFB44B20CE0EFFB4FCDBFE53FBA0FE28" author="Westwood, J. O." box="[839,1042,441,467]" pageId="9" pageNumber="100" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood (1873)</bibRefCitation>
. Hence, we considered the third female specimen as part of the type series. Mouthparts of
<taxonomicName id="4C254D52CE0EFFB4FC9DFE37FC22FE0C" authorityName=": Janson" authorityYear="1881" box="[769,912,477,503]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FC9DFE37FC22FE0C" box="[769,912,477,503]" italics="true" pageId="9" pageNumber="100">G. despectus</emphasis>
</taxonomicName>
were illustrated in the original description, and only the specimen in
<collectionCode id="ED34AE14CE0EFFB4FE1BFDEBFE6FFDE0" box="[391,477,513,539]" pageId="9" pageNumber="100">OXUM</collectionCode>
was dissected and mouthparts mounted on a card (
<figureCitation id="131E2A54CE0EFFB4FB8BFDEBFBC5FDE0" box="[1047,1143,513,539]" captionStart="FIGURES 109114" captionStartId="25.[151,264,1140,1165]" captionTargetBox="[157,1430,185,1105]" captionTargetId="figure@25.[151,1435,181,1115]" captionTargetPageId="25" captionText="FIGURES 109114. Goliathopsis despectus (Westwood, 1873). 109111, female paralectotype in OXUM and labels (photograph by Amoret Spooner); 112114, non-type female in MHNH from Combodia and labels." figureDoi="http://doi.org/10.5281/zenodo.3884757" httpUri="https://zenodo.org/record/3884757/files/figure.png" pageId="9" pageNumber="100">Fig. 111</figureCitation>
). As
<bibRefCitation id="EFB44B20CE0EFFB4FB33FDEBFAFFFDE0" author="Janson, O. E." box="[1199,1357,513,539]" pageId="9" pageNumber="100" pagination="102 - 111" refId="ref20489" refString="Janson, O. E. (1917) Additions to the knowledge of the Cetoniinae of British India. Transactions of the Entomological Society of London, 65 (1), 102 - 111. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01404. x" type="journal article" year="1917">Janson (1917)</bibRefCitation>
stated, this is usual for Westwoods type. Therefore, this female specimen is undoubted a
<typeStatus id="549E8873CE0EFFB4FBB3FDCFFB3BFDC4" box="[1071,1161,549,575]" pageId="9" pageNumber="100" type="syntype">syntype</typeStatus>
, although the locality on label reads “
<collectingCountry id="F3327641CE0EFFB4FEBFFDA3FE2EFD98" box="[291,412,585,611]" name="Cambodia" pageId="9" pageNumber="100">Cambodia</collectingCountry>
” which does not agree with the type locality. Besides, two more females also collected by Henri Mouhot from
<collectingCountry id="F3327641CE0EFFB4FEE7FD87FE5CFD7C" box="[379,494,621,647]" name="Cambodia" pageId="9" pageNumber="100">Cambodia</collectingCountry>
and
<collectingCountry id="F3327641CE0EFFB4FDB8FD87FDECFD7C" box="[548,606,621,647]" name="Thailand" pageId="9" pageNumber="100">Siam</collectingCountry>
were found in
<collectionCode id="ED34AE14CE0EFFB4FC9BFD87FCEFFD7C" box="[775,861,621,647]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
bearing label “Ex. Musaeo Parry” (
<figureCitation id="131E2A54CE0EFFB4FB71FD87FA23FD7C" box="[1261,1425,621,647]" captionStart="FIGURES 109114" captionStartId="25.[151,264,1140,1165]" captionTargetBox="[157,1430,185,1105]" captionTargetId="figure@25.[151,1435,181,1115]" captionTargetPageId="25" captionText="FIGURES 109114. Goliathopsis despectus (Westwood, 1873). 109111, female paralectotype in OXUM and labels (photograph by Amoret Spooner); 112114, non-type female in MHNH from Combodia and labels." figureDoi="http://doi.org/10.5281/zenodo.3884757" httpUri="https://zenodo.org/record/3884757/files/figure.png" pageId="9" pageNumber="100">Figs. 112114</figureCitation>
). Many species described in
<bibRefCitation id="EFB44B20CE0EFFB4FE51FD7BFD24FD50" author="Westwood, J. O." box="[461,662,657,683]" pageId="9" pageNumber="100" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood (1873)</bibRefCitation>
were definitely referred to Parry Collection, but this is not the case with
<taxonomicName id="4C254D52CE0EFFB4FF4DFD5FFED3FD34" authorityName=": Janson" authorityYear="1881" box="[209,353,693,719]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FF4DFD5FFED3FD34" box="[209,353,693,719]" italics="true" pageId="9" pageNumber="100">G. despectus</emphasis>
</taxonomicName>
. It is suggested that the two females came to Parry Collection later than the year 1873. Furthermore, there is no convincing evidence to prove that these two specimens are purchased from Saunders Collection, and thus none can be treated as
<typeStatus id="549E8873CE0EFFB4FE67FD17FDE4FCEC" box="[507,598,765,791]" pageId="9" pageNumber="100" type="syntype">syntype</typeStatus>
.
</paragraph>
<paragraph id="8B9A36D1CE0EFFB4FF5BFCCBFD8EFB84" blockId="9.[151,1437,152,2015]" pageId="9" pageNumber="100">
To preserve the stability of nomenclature, the female in
<collectionCode id="ED34AE14CE0EFFB4FCB3FCCBFC31FCC0" box="[815,899,801,827]" pageId="9" pageNumber="100">BMNH</collectionCode>
is
<emphasis id="B951EAC3CE0EFFB4FC3EFCCBFBEEFCC0" bold="true" box="[930,1116,800,827]" pageId="9" pageNumber="100">here designated</emphasis>
as
<emphasis id="B951EAC3CE0EFFB4FBE3FCCBFB58FCC0" bold="true" box="[1151,1258,801,827]" pageId="9" pageNumber="100">
<typeStatus id="549E8873CE0EFFB4FBE3FCCBFB58FCC0" box="[1151,1258,801,827]" pageId="9" pageNumber="100" type="lectotype">lectotype</typeStatus>
</emphasis>
:
<collectingCountry id="F3327641CE0EFFB4FB64FCCBFA80FCC0" box="[1272,1330,801,827]" name="Thailand" pageId="9" pageNumber="100">Siam</collectingCountry>
[rounded label] //
<taxonomicName id="4C254D52CE0EFFB4FF69FCAFFD85FCA4" authority="Westw." authorityName="Westwood" authorityYear="1873" box="[245,567,837,863]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FF69FCAFFE6DFCA4" box="[245,479,837,863]" italics="true" pageId="9" pageNumber="100">Pilinurgus despectus</emphasis>
Westw.
</taxonomicName>
(type) [Waterhouses handwriting] // Bowring, 63·47* // Type [rounded label with red border]. Two
<emphasis id="B951EAC3CE0EFFB4FE0FFC83FDF3FC78" bold="true" box="[403,577,873,899]" pageId="9" pageNumber="100">
<typeStatus id="549E8873CE0EFFB4FE0FFC83FDF3FC78" box="[403,577,873,899]" pageId="9" pageNumber="100" type="paralectotype">paralectotypes</typeStatus>
</emphasis>
labeled:
<specimenCount id="9D23FD58CE0EFFB4FD34FC83FD79FC78" box="[680,715,873,899]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FD46FC83FC86FC78" box="[730,820,873,899]" pageId="9" pageNumber="100">OXUM</collectionCode>
,
<figureCitation id="131E2A54CE0EFFB4FCA3FC83FC50FC78" box="[831,994,873,899]" captionStart="FIGURES 109114" captionStartId="25.[151,264,1140,1165]" captionTargetBox="[157,1430,185,1105]" captionTargetId="figure@25.[151,1435,181,1115]" captionTargetPageId="25" captionText="FIGURES 109114. Goliathopsis despectus (Westwood, 1873). 109111, female paralectotype in OXUM and labels (photograph by Amoret Spooner); 112114, non-type female in MHNH from Combodia and labels." figureDoi="http://doi.org/10.5281/zenodo.3884757" httpUri="https://zenodo.org/record/3884757/files/figure.png" pageId="9" pageNumber="100">Figs. 109111</figureCitation>
),
<taxonomicName id="4C254D52CE0EFFB4FC69FC83FA8BFC78" authority="Westw." authorityName="Westwood" authorityYear="1873" box="[1013,1337,873,899]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FC69FC83FB6CFC78" box="[1013,1246,873,899]" italics="true" pageId="9" pageNumber="100">Pilinurgus despectus</emphasis>
Westw.
</taxonomicName>
, Mon. pl 9, f. 3,
<collectingCountry id="F3327641CE0EFFB4FF7BFC67FEEFFC5C" box="[231,349,909,935]" name="Cambodia" pageId="9" pageNumber="100">Cambodia</collectingCountry>
, Mouhot, Steven 1861 [Westwoods handwriting on blue label] // Co-type, Westwood, Thes. Ent. p. 32, T. 9, f. 3, Coll. Hope. Oxon. //
<taxonomicName id="4C254D52CE0EFFB4FDDDFC5BFC2CFC30" authority="Westw." authorityName="Westw." box="[577,926,945,971]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FDDDFC5BFCF7FC30" box="[577,837,945,971]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
Westw.
</taxonomicName>
Type, test O.E.Janson, Tr. Ent. Soc. 1917, p. 110 [Jansons handwriting] //
<taxonomicName id="4C254D52CE0EFFB4FE7BFC3FFCF6FC14" authority="Westw." authorityName="Westw." box="[487,836,981,1007]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FE7BFC3FFD59FC14" box="[487,747,981,1007]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
Westw.
</taxonomicName>
Type, test O.E.Janson, Trans. Ent. Soc. 1917, p. 110,
<specimenCount id="9D23FD58CE0EFFB4FF0BFC13FF1EFBE8" box="[151,172,1017,1043]" pageId="9" pageNumber="100" type="female"></specimenCount>
[Jansons handwriting] // t.
<date id="FF9B1011CE0EFFB4FE79FC13FDD0FBE8" box="[485,610,1017,1043]" pageId="9" pageNumber="100" value="1912" valueMax="1920">1912-1920</date>
. O.E.Janson. //
<taxonomicName id="4C254D52CE0EFFB4FC8CFC13FBD8FBE8" authority="Westw." authorityName="Westw." box="[784,1130,1017,1043]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FC8CFC13FBA0FBE8" box="[784,1042,1017,1043]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
Westw.
</taxonomicName>
t. G.J.Arrow. in B.M. 1907 // TYPE COL: 850,
<taxonomicName id="4C254D52CE0EFFB4FEE5FBF7FD53FBCC" authority="Westwood" authorityName="Westwood" authorityYear="1873" box="[377,737,1053,1079]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FEE5FBF7FDD0FBCC" box="[377,610,1053,1079]" italics="true" pageId="9" pageNumber="100">Pilinurgus despectus</emphasis>
Westwood
</taxonomicName>
, HOPE DEPT. OXFORD;
<specimenCount id="9D23FD58CE0EFFB4FB93FBF7FB80FBCC" box="[1039,1074,1053,1079]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FBDDFBF7FB2EFBCC" box="[1089,1180,1053,1079]" pageId="9" pageNumber="100">MHNH</collectionCode>
),
<collectingCountry id="F3327641CE0EFFB4FB33FBF7FB59FBCC" box="[1199,1259,1053,1079]" name="Thailand" pageId="9" pageNumber="100">Siam</collectingCountry>
, Mouhot// MU- SEUM PARIS, 1952, Coll. R. Oberthür // Ex. Musaeo D. Sharp 1890 //
<taxonomicName id="4C254D52CE0EFFB4FC5FFBABFABAFBA0" authority="Westw." authorityName="Westwood" authorityYear="1873" box="[963,1288,1089,1115]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FC5FFBABFB1FFBA0" box="[963,1197,1089,1115]" italics="true" pageId="9" pageNumber="100">Pilinurgus despectus</emphasis>
Westw.
</taxonomicName>
, nov., Pl9 f3, (pinguidine obscurata) [handwriting].
</paragraph>
<paragraph id="8B9A36D1CE0EFFB4FF5BFB63FBE5F9D4" blockId="9.[151,1437,152,2015]" pageId="9" pageNumber="100">
<materialsCitation id="3B4D3C8CCE0EFFB4FF5BFB63FBE1F9D4" collectingDate="1852" collectingDateMax="2018-03" collectingDateMin="1852" collectionCode="MSNG, TYPUS, BMNH, MFNB, MHNG, MNHN, IRSNB" collectorName="Mr. Leonardo Fea" country="Myanmar" location="Myanmar" pageId="9" pageNumber="100" specimenCount="17" specimenCount-female="6" specimenCount-male="11" typeStatus="syntype">
All the six specimens (
<specimenCount id="9D23FD58CE0EFFB4FE5DFB63FDA8FB58" box="[449,538,1161,1187]" pageId="9" pageNumber="100" type="male">4 males</specimenCount>
,
<specimenCount id="9D23FD58CE0EFFB4FDB8FB63FD23FB5F" box="[548,657,1161,1188]" pageId="9" pageNumber="100" type="female">2 females</specimenCount>
) of
<taxonomicName id="4C254D52CE0EFFB4FD27FB63FC0EFB58" box="[699,956,1161,1187]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FD27FB63FC0EFB58" box="[699,956,1161,1187]" italics="true" pageId="9" pageNumber="100">Goliathopsis capreolus</emphasis>
</taxonomicName>
in
<collectionCode id="ED34AE14CE0EFFB4FC42FB63FB9DFB58" box="[990,1071,1161,1187]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
bear an identical printed date-locality label and conform to the original description (
<bibRefCitation id="EFB44B20CE0EFFB4FD45FB47FCDBFB3C" author="Gestro, R." box="[729,873,1197,1223]" pageId="9" pageNumber="100" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro 1888</bibRefCitation>
): they were all collected from Moulmein, southern
<collectingCountry id="F3327641CE0EFFB4FF0BFB3BFEB6FB10" box="[151,260,1233,1259]" name="Myanmar" pageId="9" pageNumber="100">Myanmar</collectingCountry>
by
<collectorName id="26D05307CE0EFFB4FEB3FB3BFE4AFB10" box="[303,504,1233,1259]" pageId="9" pageNumber="100">Mr. Leonardo Fea</collectorName>
(
<date id="FF9B1011CE0EFFB4FD94FB3AFD3BFB10" box="[520,649,1232,1259]" pageId="9" pageNumber="100" value="1852" valueMax="1903">
<collectingDate id="EFDFE9F9CE0EFFB4FD94FB3AFD3BFB10" box="[520,649,1232,1259]" pageId="9" pageNumber="100" value="1852" valueMax="1903">18521903</collectingDate>
</date>
) in
<date id="FF9B1011CE0EFFB4FD29FB3BFC99FB10" box="[693,811,1233,1259]" pageId="9" pageNumber="100" value="1887-05">
<collectingDate id="EFDFE9F9CE0EFFB4FD29FB3BFC99FB10" box="[693,811,1233,1259]" pageId="9" pageNumber="100" value="1887-05">May 1887</collectingDate>
</date>
. Although a printed type label (
<collectionCode id="ED34AE14CE0EFFB4FB11FB3BFB5BFB10" box="[1165,1257,1233,1259]" pageId="9" pageNumber="100">TYPUS</collectionCode>
) alongside only one specimen of each sex (
<figureCitation id="131E2A54CE0EFFB4FE56FB1FFDDDFAF4" box="[458,623,1269,1295]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="9" pageNumber="100">Figs. 118, 121</figureCitation>
), all of them must be regarded as
<typeStatus id="549E8873CE0EFFB4FC6FFB1FFBEBFAF4" box="[1011,1113,1269,1295]" pageId="9" pageNumber="100" type="syntype">syntypes</typeStatus>
. Additionally, the specimens in
<collectionCode id="ED34AE14CE0EFFB4FF2AFAF3FEB8FAC8" box="[182,266,1305,1331]" pageId="9" pageNumber="100">BMNH</collectionCode>
(
<specimenCount id="9D23FD58CE0EFFB4FE82FAF3FEDEFACF" box="[286,364,1305,1332]" pageId="9" pageNumber="100" type="male">1 male</specimenCount>
),
<collectionCode id="ED34AE14CE0EFFB4FE1EFAF3FE63FAC8" box="[386,465,1305,1331]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="100">MFNB</collectionCode>
(
<specimenCount id="9D23FD58CE0EFFB4FE7FFAF3FD8DFACF" box="[483,575,1305,1332]" pageId="9" pageNumber="100" type="male">2 males</specimenCount>
),
<collectionCode id="ED34AE14CE0EFFB4FDC8FAF3FD18FAC8" box="[596,682,1305,1331]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="100" type="Museum">MHNG</collectionCode>
(
<specimenCount id="9D23FD58CE0EFFB4FD22FAF3FC96FACF" box="[702,804,1305,1332]" pageId="9" pageNumber="100" type="female">1 female</specimenCount>
),
<collectionCode id="ED34AE14CE0EFFB4FCA4FAF3FC3CFAC8" box="[824,910,1305,1331]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
(
<specimenCount id="9D23FD58CE0EFFB4FC3DFAF3FC5EFAC8" box="[929,1004,1305,1331]" pageId="9" pageNumber="100" type="male">1 male</specimenCount>
and
<specimenCount id="9D23FD58CE0EFFB4FBBBFAF3FB3CFACF" box="[1063,1166,1305,1332]" pageId="9" pageNumber="100" type="female">1 female</specimenCount>
),
<collectionCode id="ED34AE14CE0EFFB4FB3FFAF3FB47FAC8" box="[1187,1269,1305,1331]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="9" pageNumber="100">IRSNB</collectionCode>
(
<specimenCount id="9D23FD58CE0EFFB4FA95FAF3FAE6FAC8" box="[1289,1364,1305,1331]" pageId="9" pageNumber="100" type="male">1 male</specimenCount>
and
<specimenCount id="9D23FD58CE0EFFB4FA12FAF3FF57FAA3" pageId="9" pageNumber="100" type="female">1 female</specimenCount>
) all attached the identical date-locality label; another three specimens (
<specimenCount id="9D23FD58CE0EFFB4FB92FAD7FBEAFAAC" box="[1038,1112,1341,1367]" pageId="9" pageNumber="100" type="male">1 male</specimenCount>
in
<collectionCode id="ED34AE14CE0EFFB4FBE2FAD7FB64FAAC" box="[1150,1238,1341,1367]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
,
<specimenCount id="9D23FD58CE0EFFB4FB7EFAD7FA9CFAAC" box="[1250,1326,1341,1367]" pageId="9" pageNumber="100" type="male">1 male</specimenCount>
and
<specimenCount id="9D23FD58CE0EFFB4FAFBFAD7FF7FFA80" pageId="9" pageNumber="100" type="female">1 female</specimenCount>
in
<collectionCode id="ED34AE14CE0EFFB4FF6FFA8BFEF6FA80" box="[243,324,1377,1403]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="100">MFNB</collectionCode>
) with handwritten label reflecting “collected from Burma by Fea in
<date id="FF9B1011CE0EFFB4FBD6FA8BFB0DFA80" box="[1098,1215,1377,1403]" pageId="9" pageNumber="100" value="1887-05">
<collectingDate id="EFDFE9F9CE0EFFB4FBD6FA8BFB0DFA80" box="[1098,1215,1377,1403]" pageId="9" pageNumber="100" value="1887-05">May 1887</collectingDate>
</date>
” are also treated as
<typeStatus id="549E8873CE0EFFB4FF0BFA6FFF4EFA64" box="[151,252,1413,1439]" pageId="9" pageNumber="100" type="syntype">syntypes</typeStatus>
, because the labeling indicates that those specimens were part of the type series originally preserved in
<collectionCode id="ED34AE14CE0EFFB4FF0BFA43FF5AFA38" box="[151,232,1449,1475]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
and were exchanged after
<taxonomicName id="4C254D52CE0EFFB4FD93FA43FD2FFA38" box="[527,669,1449,1475]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FD93FA43FD2FFA38" box="[527,669,1449,1475]" italics="true" pageId="9" pageNumber="100">G. capreolus</emphasis>
</taxonomicName>
been named (Roberto Poggi, personal communication,
<date id="FF9B1011CE0EFFB4FA99FA43FA23FA38" box="[1285,1425,1449,1475]" pageId="9" pageNumber="100" value="2018-03">
<collectingDate id="EFDFE9F9CE0EFFB4FA99FA43FA23FA38" box="[1285,1425,1449,1475]" pageId="9" pageNumber="100" value="2018-03">March 2018</collectingDate>
</date>
). More potential
<typeStatus id="549E8873CE0EFFB4FEDFFA27FE17FA1C" box="[323,421,1485,1511]" pageId="9" pageNumber="100" type="syntype">syntypes</typeStatus>
perhaps exist in other European museums we have not visited. While, the female specimen bearing a handwritten label of data “Moulmein” in
<collectionCode id="ED34AE14CE0EFFB4FD4DFA1BFC95F9F0" box="[721,807,1521,1547]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
was not regarded as
<typeStatus id="549E8873CE0EFFB4FB8CFA1BFBD5F9F0" box="[1040,1127,1521,1547]" pageId="9" pageNumber="100" type="syntype">syntype</typeStatus>
due to the label style is different and no additional information to confirm it belonged to the original type series
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36D1CE0EFFB7FF5BF9D3FB00FE70" blockId="9.[151,1437,152,2015]" lastBlockId="10.[151,1437,152,1835]" lastPageId="10" lastPageNumber="101" pageId="9" pageNumber="100">
To preserve nomenclatural stability, the male (
<figureCitation id="131E2A54CE0EFFB4FD40F9D3FCCDF9A8" box="[732,895,1592,1619]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="9" pageNumber="100">Figs. 115118</figureCitation>
) in
<collectionCode id="ED34AE14CE0EFFB4FC33F9D3FBB2F9A8" box="[943,1024,1593,1619]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
bearing a handwritten type label of Gestro is
<emphasis id="B951EAC3CE0EFFB4FE99F9B7FE73F98C" bold="true" box="[261,449,1628,1655]" pageId="9" pageNumber="100">here designated</emphasis>
as
<emphasis id="B951EAC3CE0EFFB4FE75F9B7FDE6F98C" bold="true" box="[489,596,1629,1655]" pageId="9" pageNumber="100">
<typeStatus id="549E8873CE0EFFB4FE75F9B7FDE6F98C" box="[489,596,1629,1655]" pageId="9" pageNumber="100" type="lectotype">lectotype</typeStatus>
</emphasis>
: Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FC3AF9B7FB8CF98C" box="[934,1086,1629,1655]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
// F. caves [Gestros handwriting] // Museo Civ. Genova [orange label]// TYPUS [red ink]//
<taxonomicName id="4C254D52CE0EFFB4FCD7F96BFBBEF960" authority="Gestro" authorityName="Gestro" box="[843,1036,1665,1691]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FCD7F96BFC0AF960" box="[843,952,1665,1691]" italics="true" pageId="9" pageNumber="100">capreolus</emphasis>
Gestro
</taxonomicName>
,
<specimenCount id="9D23FD58CE0EFFB4FB8BF96BFB9EF960" box="[1047,1068,1665,1691]" pageId="9" pageNumber="100" type="male"></specimenCount>
<specimenCount id="9D23FD58CE0EFFB4FBAEF96BFBF5F960" box="[1074,1095,1665,1691]" pageId="9" pageNumber="100" type="female"></specimenCount>
[Gestros handwriting] //
<emphasis id="B951EAC3CE0EFFB4FAFDF96BFF46F944" italics="true" pageId="9" pageNumber="100">Goli- athopsis</emphasis>
<taxonomicName id="4C254D52CE0EFFB4FF67F94FFE0EF944" authority="Gestro" authorityName="Gestro" box="[251,444,1701,1727]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FF67F94FFEDAF944" box="[251,360,1701,1727]" italics="true" pageId="9" pageNumber="100">capreolus</emphasis>
Gestro
</taxonomicName>
, typus!
<specimenCount id="9D23FD58CE0EFFB4FD87F94FFD82F944" box="[539,560,1701,1727]" pageId="9" pageNumber="100" type="male"></specimenCount>
<specimenCount id="9D23FD58CE0EFFB4FDAAF94FFDF9F944" box="[566,587,1701,1727]" pageId="9" pageNumber="100" type="female"></specimenCount>
[Gestros handwriting]. Sixteen examined
<emphasis id="B951EAC3CE0EFFB4FBB7F94FFB6BF944" bold="true" box="[1067,1241,1701,1727]" pageId="9" pageNumber="100">
<typeStatus id="549E8873CE0EFFB4FBB7F94FFB6BF944" box="[1067,1241,1701,1727]" pageId="9" pageNumber="100" type="paralectotype">paralectotypes</typeStatus>
</emphasis>
(
<specimenCount id="9D23FD58CE0EFFB4FB75F94FFA80F944" box="[1257,1330,1701,1727]" pageId="9" pageNumber="100" type="male">10♂♂</specimenCount>
,
<specimenCount id="9D23FD58CE0EFFB4FAA1F94FFAC6F93B" box="[1341,1396,1701,1728]" pageId="9" pageNumber="100" type="female">6♀♀</specimenCount>
) of
<taxonomicName id="4C254D52CE0EFFB4FF0BF923FE2BF918" box="[151,409,1737,1763]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FF0BF923FE2BF918" box="[151,409,1737,1763]" italics="true" pageId="9" pageNumber="100">Goliathopsis capreolus</emphasis>
</taxonomicName>
:
<specimenCount id="9D23FD58CE0EFFB4FE30F923FE7DF918" box="[428,463,1737,1763]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FE40F923FD83F918" box="[476,561,1737,1763]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
,
<figureCitation id="131E2A54CE0EFFB4FDA7F923FD6EF918" box="[571,732,1737,1763]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="9" pageNumber="100">Figs. 119121</figureCitation>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FBB5F923FB0FF918" box="[1065,1213,1737,1763]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
// F. caves [Gestros handwriting] // Museo Civ. Genova [orange label]// TYPUS [red ink];
<specimenCount id="9D23FD58CE0EFFB4FC5DF907FC56F8FC" box="[961,996,1773,1799]" pageId="9" pageNumber="100" type="male">1♂</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FC69F907FBFEF8FC" box="[1013,1100,1773,1799]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FF0BF8FBFE9CF8D0" box="[151,302,1809,1835]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
// Farm. caves [Gestros handwriting] // Museo Civ. Genova [orange label]//
<taxonomicName id="4C254D52CE0EFFB4FB04F8FBFEDEF8B4" authority="Westw., G. J. Arrow" authorityName="Westw., G. J. Arrow" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FB04F8FBFA2EF8D0" box="[1176,1436,1809,1835]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
Westw., G.J.Arrow
</taxonomicName>
det. [Arrows handwriting];
<specimenCount id="9D23FD58CE0EFFB4FD2DF8DFFD59F8B4" box="[689,747,1845,1871]" pageId="9" pageNumber="100" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D23FD58CE0EFFB4FD6BF8DFFCA8F8B4" box="[759,794,1845,1871]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FCB5F8DFFC32F8B4" box="[809,896,1845,1871]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="9" pageNumber="100">MSNG</collectionCode>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FB4FF8DFFAD8F8B4" box="[1235,1386,1845,1871]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
// F. caves [Gestros handwriting] // Museo Civ. Genova [white label];
<specimenCount id="9D23FD58CE0EFFB4FCF3F8B3FC20F888" box="[879,914,1881,1907]" pageId="9" pageNumber="100" type="male">1♂</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FC03F8B3FC4BF888" box="[927,1017,1881,1907]" pageId="9" pageNumber="100">BMNH</collectionCode>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FAD9F8B3FF7DF86C" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
// Deuts. Ent. Mus. 1909-157 //
<taxonomicName id="4C254D52CE0EFFB4FDA2F897FC2BF86C" authority="Westw." authorityName="Westw." box="[574,921,1917,1943]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0EFFB4FDA2F897FCF3F86C" box="[574,833,1917,1943]" italics="true" pageId="9" pageNumber="100">Goliathopsis despectus</emphasis>
Westw.
</taxonomicName>
// G.J.Arrow det.;
<specimenCount id="9D23FD58CE0EFFB4FBFAF897FB2DF86C" box="[1126,1183,1917,1943]" pageId="9" pageNumber="100" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FB32F897FAB1F86C" box="[1198,1283,1917,1943]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="100">MFNB</collectionCode>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FED0F84BFE55F840" box="[332,487,1953,1979]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
;
<specimenCount id="9D23FD58CE0EFFB4FE68F84BFDAAF840" box="[500,536,1953,1979]" pageId="9" pageNumber="100" type="male">1♂</specimenCount>
,
<specimenCount id="9D23FD58CE0EFFB4FDBAF84BFDFBF840" box="[550,585,1953,1979]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FDC6F84BFD1DF840" box="[602,687,1953,1979]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="100">MFNB</collectionCode>
),
<collectingCountry id="F3327641CE0EFFB4FD5FF84BFCA0F840" box="[707,786,1953,1979]" name="Myanmar" pageId="9" pageNumber="100">Burma</collectingCountry>
, Maggio, Fea;
<specimenCount id="9D23FD58CE0EFFB4FC22F84BFC53F840" box="[958,993,1953,1979]" pageId="9" pageNumber="100" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FC6EF84BFBFFF840" box="[1010,1101,1953,1979]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="100" type="Museum">MHNG</collectionCode>
), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0EFFB4FF0BF82FFE83F824" box="[151,305,1989,2015]" pageId="9" pageNumber="100" value="1887-05">Maggio 1887</date>
;
<specimenCount id="9D23FD58CE0EFFB4FEA1F82FFED2F824" box="[317,352,1989,2015]" pageId="9" pageNumber="100" type="male">1♂</specimenCount>
(
<collectionCode id="ED34AE14CE0EFFB4FEECF82FFE79F824" box="[368,459,1989,2015]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="100">MNHN</collectionCode>
),
<taxonomicName id="4C254D52CE0EFFB4FE43F82FFC8AF824" authority="Gestro" authorityName="Gestro" box="[479,824,1989,2015]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0EFFB4FE43F82FFD51F824" box="[479,739,1989,2015]" italics="true" pageId="9" pageNumber="100">Goliathopsis capreolus</emphasis>
Gestro
</taxonomicName>
, typus! Moulmein, Fea [Gestros handwriting] //
<emphasis id="B951EAC3CE0EFFB7FAF6F82CFF6EFF48" italics="true" lastPageId="10" lastPageNumber="101" pageId="9" pageNumber="100">cap- reolus</emphasis>
gestro, moulmein [unidentified handwriting]// MUSEUM PARIS, 1952, Coll. R. Oberthür;
<specimenCount id="9D23FD58CE0DFFB7FB76FF73FABEFF48" box="[1258,1292,153,179]" pageId="10" pageNumber="101" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FA80FF73FAC4FF48" box="[1308,1398,153,179]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
, de Lisle coll.), Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0DFFB7FDC2FF57FD46FF2C" box="[606,756,189,215]" pageId="10" pageNumber="101" value="1887-05">Maggio 1887</date>
// F. caves [Gestros handwriting] //
<typeStatus id="549E8873CE0DFFB7FB12FF57FAABFF2C" box="[1166,1305,189,215]" pageId="10" pageNumber="101" type="paratype">PARATYPE</typeStatus>
[red label];
<specimenCount id="9D23FD58CE0DFFB7FF0BFF0BFF08FF00" box="[151,186,225,251]" pageId="10" pageNumber="101" type="male">1♂</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FF51FF0BFE9AFF00" box="[205,296,225,251]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
), Tenasserim, Moulmein, Fea.
<date id="FF9B1011CE0DFFB7FD13FF0BFC95FF00" box="[655,807,225,251]" pageId="10" pageNumber="101" value="1887-05">Maggio 1887</date>
//
<taxonomicName id="4C254D52CE0DFFB7FCDFFF0BFB10FF00" authority="Gestro" authorityName="Gestro" box="[835,1186,225,251]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FCDFFF0BFBF8FF00" box="[835,1098,225,251]" italics="true" pageId="10" pageNumber="101">Goliathopsis capreolus</emphasis>
Gestro
</taxonomicName>
, ex typ. [unidentified handwriting] // MUSEUM PARIS, 1936, DE LISLE [blue label]//
<taxonomicName id="4C254D52CE0DFFB7FC0AFEEFFABBFEE4" authority="(Westw.)" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[918,1289,261,287]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FC0AFEEFFB2EFEE4" box="[918,1180,261,287]" italics="true" pageId="10" pageNumber="101">Goliathopsis despectus</emphasis>
(Westw.)
</taxonomicName>
<emphasis id="B951EAC3CE0DFFB7FA8EFEECFA97FEE4" box="[1298,1317,262,287]" italics="true" pageId="10" pageNumber="101">=</emphasis>
<taxonomicName id="4C254D52CE0DFFB7FAB3FEEFFEA6FEB8" authority="Gestro, Ph." authorityName="Gestro, Ph." class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FAB3FEEFFA2EFEE4" box="[1327,1436,261,287]" italics="true" pageId="10" pageNumber="101">capreolus</emphasis>
Gestro, Ph.
</taxonomicName>
ANTOINE det. 1990 [white label]//
<typeStatus id="549E8873CE0DFFB7FD36FEC3FC9AFEB8" box="[682,808,297,323]" pageId="10" pageNumber="101" type="syntype">SYNTYPE</typeStatus>
[red label]//
<typeStatus id="549E8873CE0DFFB7FC2BFEC3FB8AFEB8" box="[951,1080,297,323]" pageId="10" pageNumber="101" type="syntype">SYNTYPE</typeStatus>
,
<taxonomicName id="4C254D52CE0DFFB7FBDEFEC3FF7DFE9C" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FBDEFEC3FAF6FEB8" box="[1090,1348,297,323]" italics="true" pageId="10" pageNumber="101">Goliathopsis capreolus</emphasis>
<bibRefCitation id="EFB44B20CE0DFFB7FAD7FEC3FF7DFE9C" author="Gestro, R." pageId="10" pageNumber="101" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro, 1888</bibRefCitation>
</taxonomicName>
[white label]//
<collectionCode id="ED34AE14CE0DFFB7FEE7FEA7FE61FE9C" box="[379,467,333,359]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
,
<collectionCode id="ED34AE14CE0DFFB7FE42FEA7FDB0FE9C" box="[478,514,333,359]" pageId="10" pageNumber="101">EC</collectionCode>
7184 [white label];
<specimenCount id="9D23FD58CE0DFFB7FD41FEA7FCB0FE9C" box="[733,770,333,359]" pageId="10" pageNumber="101" type="male">1♂</specimenCount>
,
<specimenCount id="9D23FD58CE0DFFB7FC92FEA7FC83FE9C" box="[782,817,333,359]" pageId="10" pageNumber="101" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FCDCFEA7FC2AFE9C" box="[832,920,333,359]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="101">IRSNB</collectionCode>
), Coll. R. I. Sc. N. B., Tenasserim, Moulmein, Fea
<date id="FF9B1011CE0DFFB7FF5BFE9BFEE9FE70" box="[199,347,369,395]" pageId="10" pageNumber="101" value="1887-05">Maggio 1887</date>
//
<taxonomicName id="4C254D52CE0DFFB7FEEFFE9BFD79FE70" authority="Gestro" authorityName="Gestro" box="[371,715,369,395]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FEEFFE9BFDC4FE70" box="[371,630,369,395]" italics="true" pageId="10" pageNumber="101">Goliathopsis capreolus</emphasis>
Gestro
</taxonomicName>
, det.
<bibRefCitation id="EFB44B20CE0DFFB7FC9BFE9BFC22FE70" author="Gestro, R." box="[775,912,369,395]" pageId="10" pageNumber="101" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro 1888</bibRefCitation>
[handwriting]//
<typeStatus id="549E8873CE0DFFB7FBD4FE9BFB1CFE70" box="[1096,1198,369,395]" pageId="10" pageNumber="101" type="paratype">Paratype</typeStatus>
.
</paragraph>
<paragraph id="8B9A36D1CE0DFFB7FF5BFE7FFC27FD98" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
Except for the printed date-locality label, a handwritten label “F. caves” attached to each
<typeStatus id="549E8873CE0DFFB7FB5AFE7FFAAFFE54" box="[1222,1309,405,431]" pageId="10" pageNumber="101" type="syntype">syntype</typeStatus>
in
<collectionCode id="ED34AE14CE0DFFB7FAD8FE7FFA2BFE54" box="[1348,1433,405,431]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="10" pageNumber="101">MSNG</collectionCode>
. According to the collecting routes of Mr. Leonardo Fea presented in
<bibRefCitation id="EFB44B20CE0DFFB7FC30FE53FBFEFE28" author="Gestro, R." box="[940,1100,441,467]" pageId="10" pageNumber="101" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro (1888)</bibRefCitation>
, Fea had been Farm caves of Moulmein in
<date id="FF9B1011CE0DFFB7FEB3FE37FE11FE0C" box="[303,419,477,503]" pageId="10" pageNumber="101" value="1887-05">May 1887</date>
. Gestro probably added the label of precise site by his own handwriting, and the type locality of
<taxonomicName id="4C254D52CE0DFFB7FF47FDEBFE0CFDE0" authority="Gestro" authorityName="Gestro" box="[219,446,513,539]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FF47FDEBFED9FDE0" box="[219,363,513,539]" italics="true" pageId="10" pageNumber="101">G. capreolus</emphasis>
Gestro
</taxonomicName>
is thus considered as Farm Caves. The caves (N 16°3200.08, E 97°4252.41, alt.
<quantity id="4CDD9B34CE0DFFB7FA1CFDEBFF1CFDC4" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="101" unit="m" value="80.0">80 m</quantity>
) also called Kayon Caves or Kha-Yone Cave, which are situated at Kayon Hill about
<quantity id="4CDD9B34CE0DFFB7FBF0FDCFFB01FDC4" box="[1132,1203,549,575]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="101" unit="km" value="13.0">13 km</quantity>
northeast of Mawlamyine (formerly Moulmein) in southern
<collectingCountry id="F3327641CE0DFFB7FDFCFDA3FD7FFD98" box="[608,717,585,611]" name="Myanmar" pageId="10" pageNumber="101">Myanmar</collectingCountry>
(
<bibRefCitation id="EFB44B20CE0DFFB7FD47FDA3FC3AFD98" author="Chhibber, H. L." box="[731,904,585,611]" pageId="10" pageNumber="101" pagination="124 - 131" refId="ref19807" refString="Chhibber, H. L. (1928) A short note on the Limestone Caves in the neighbourhood of Nyaungbinzeik, Kyaikmaraw Township, Amherst District, Lower Burma. Journal of the Burma Research Society, 18 (3), 124 - 131." type="journal article" year="1928">Chhibber 1928</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A36D1CE0DFFB7FF5BFD86FEB1FCEC" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
<emphasis id="B951EAC3CE0DFFB7FF5BFD86FE4DFD7C" bold="true" box="[199,511,620,647]" pageId="10" pageNumber="101">Other material examined</emphasis>
(
<specimenCount id="9D23FD58CE0DFFB7FD8BFD87FDFCFD7C" box="[535,590,621,647]" pageId="10" pageNumber="101" type="female">3♀♀</specimenCount>
).
<emphasis id="B951EAC3CE0DFFB7FDF6FD86FCA0FD7C" bold="true" box="[618,786,620,647]" pageId="10" pageNumber="101">
<collectingCountry id="F3327641CE0DFFB7FDF6FD86FCBFFD7C" box="[618,781,620,647]" name="Cambodia" pageId="10" pageNumber="101">CAMBODIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D23FD58CE0DFFB7FC83FD87FCF0FD7C" box="[799,834,621,647]" pageId="10" pageNumber="101" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FCCAFD87FC03FD7C" box="[854,945,621,647]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
),
<taxonomicName id="4C254D52CE0DFFB7FC56FD87FAABFD7C" authority="Westw." authorityName="Westwood" authorityYear="1873" box="[970,1305,621,647]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FC56FD87FB0BFD7C" box="[970,1209,621,647]" italics="true" pageId="10" pageNumber="101">Pilinurgus despectus</emphasis>
Westw.
</taxonomicName>
, Cambodia [Parrys handwriting] // Ex. Musaeo Parry // MUSEUM PARIS, 1952, Coll. R. Oberthür //
<taxonomicName id="4C254D52CE0DFFB7FB0EFD7BFAEAFD50" authority="Westw." authorityName="Westwood" authorityYear="1873" box="[1170,1368,657,683]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FB0EFD7BFB4DFD50" box="[1170,1279,657,683]" italics="true" pageId="10" pageNumber="101">despectus</emphasis>
Westw.
</taxonomicName>
Cambodia.
<emphasis id="B951EAC3CE0DFFB7FF7AFD5EFECFFD34" bold="true" box="[230,381,692,719]" pageId="10" pageNumber="101">
<collectingCountry id="F3327641CE0DFFB7FF7AFD5EFECFFD34" box="[230,381,692,719]" name="Thailand" pageId="10" pageNumber="101">THAILAND</collectingCountry>
</emphasis>
:
<specimenCount id="9D23FD58CE0DFFB7FE13FD5FFE00FD34" box="[399,434,693,719]" pageId="10" pageNumber="101" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FE5FFD5FFDACFD34" box="[451,542,693,719]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
), Siam, Mouhot // MUSEUM PARIS, 1952, Coll. R. Oberthür // Ex. Musaeo Parry.
<emphasis id="B951EAC3CE0DFFB7FF7DFD32FECAFD08" bold="true" box="[225,376,728,755]" pageId="10" pageNumber="101">
<collectingCountry id="F3327641CE0DFFB7FF7DFD32FECAFD08" box="[225,376,728,755]" name="Myanmar" pageId="10" pageNumber="101">MYANMAR</collectingCountry>
</emphasis>
:
<specimenCount id="9D23FD58CE0DFFB7FE1BFD33FE18FD08" box="[391,426,729,755]" pageId="10" pageNumber="101" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE0DFFB7FE25FD33FDA6FD08" box="[441,532,729,755]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="101">MNHN</collectionCode>
), Moulmein // Ex. Musaeo H.W.Bates 1892 // MUSEUM PARIS, 1952, Coll. R. Oberthür.
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE0DFFB7FF5BFCCAFB13FB33" pageId="10" pageNumber="101" type="diagnosis">
<paragraph id="8B9A36D1CE0DFFB7FF5BFCCAFE82FBA0" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
<emphasis id="B951EAC3CE0DFFB7FF5BFCCAFEF0FCC0" bold="true" box="[199,322,800,827]" pageId="10" pageNumber="101">Diagnosis.</emphasis>
Body length 12.013.0 mm; width 6.57.0 mm. Dorsal surface and pygidium clad with brown tomentum (
<figureCitation id="131E2A54CE0DFFB7FE9FFCAFFE6CFCA4" box="[259,478,836,863]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="10" pageNumber="101">Figs. 115, 117, 119</figureCitation>
). Midline on pronotum and scutellum yellow, distinct. Elytron usually with four light brown maculae: a small one on posthumeral area (
<figureCitation id="131E2A54CE0DFFB7FD5BFC83FC95FC78" box="[711,807,873,899]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="10" pageNumber="101">Fig. 117</figureCitation>
), sometimes absent; a small one on the mediodiscal area near sutural costa; a large one on the median portion of lateral declivity; a large one on the postdiscal areas near sutural costa (
<figureCitation id="131E2A54CE0DFFB7FEAFFC5BFE21FC30" box="[307,403,945,971]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="10" pageNumber="101">Fig. 119</figureCitation>
), sometimes split up (
<figureCitation id="131E2A54CE0DFFB7FD14FC5BFD5BFC30" box="[648,745,944,971]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="10" pageNumber="101">Fig. 115</figureCitation>
). Disc of elytron with U- or drop shaped, setiferous punctures (
<figureCitation id="131E2A54CE0DFFB7FF3CFC3FFF54FC0B" box="[160,230,981,1008]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="10" pageNumber="101">Fig. 6</figureCitation>
); setae long. Pygidium with rounded, setiferous punctures (
<figureCitation id="131E2A54CE0DFFB7FC0DFC3FFBAFFC14" box="[913,1053,980,1007]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="10" pageNumber="101">Figs. 2425</figureCitation>
). Cephalic horns of male slender, without tomentum, not constricted at base (
<figureCitation id="131E2A54CE0DFFB7FD1FFC13FD6AFBE8" box="[643,728,1017,1043]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="10" pageNumber="101">Fig. 14</figureCitation>
). Male genitalia small (
<figureCitation id="131E2A54CE0DFFB7FC78FC13FBDCFBE8" box="[996,1134,1017,1043]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="10" pageNumber="101">Figs. 4849</figureCitation>
), proximal and distal parts of parameres expanded in apical view, medially constricted, apex sharp; parameres near flat in lateral view, basal piece slender.
</paragraph>
<paragraph id="8B9A36D1CE0DFFB7FF5BFB8FFB13FB33" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
This species is close to
<taxonomicName id="4C254D52CE0DFFB7FE4AFB8FFD71FB84" authorityName="Antoine" authorityYear="1991" box="[470,707,1125,1151]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE0DFFB7FE4AFB8FFD71FB84" box="[470,707,1125,1151]" italics="true" pageId="10" pageNumber="101">Goliathopsis ferreroi</emphasis>
</taxonomicName>
, but setae on elytra shorter (
<figureCitation id="131E2A54CE0DFFB7FB96FB8FFBDEFB84" box="[1034,1132,1125,1151]" captionStart="FIGURES 115121" captionStartId="26.[151,264,1159,1184]" captionTargetBox="[155,1422,187,1125]" captionTargetId="figure@26.[151,1435,181,1133]" captionTargetPageId="26" captionText="FIGURES 115121. Type specimens of Goliathopsis capreolus Gestro, 1888 in MSNG. 115118, male lectotype and labels; 119121, female paralectotype and labels." figureDoi="http://doi.org/10.5281/zenodo.3884759" httpUri="https://zenodo.org/record/3884759/files/figure.png" pageId="10" pageNumber="101">Fig. 117</figureCitation>
; longer in
<taxonomicName id="4C254D52CE0DFFB7FB74FB8FFAD2FB84" authorityName="Antoine" authorityYear="1991" box="[1256,1376,1125,1151]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE0DFFB7FB74FB8FFAD2FB84" box="[1256,1376,1125,1151]" italics="true" pageId="10" pageNumber="101">G. ferreroi</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE0DFFB7FAECFB8FFF76FB58" captionStart="FIGURES 138143" captionStartId="29.[151,264,1140,1165]" captionTargetBox="[156,1407,197,1096]" captionTargetId="figure@29.[151,1435,181,1115]" captionTargetPageId="29" captionText="FIGURES 138143. Type specimens of Goliathopsis ferreroi Antoine, 1991 in MNHN. 138141, male holotype and labels; 142143, female paratype (allotype) and labels." figureDoi="http://doi.org/10.5281/zenodo.3884767" httpUri="https://zenodo.org/record/3884767/files/figure.png" pageId="10" pageNumber="101">Fig. 140</figureCitation>
), punctures on pygidium larger and sparser (
<figureCitation id="131E2A54CE0DFFB7FD24FB63FCF2FB58" box="[696,832,1160,1187]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="10" pageNumber="101">Figs. 2425</figureCitation>
; smaller and denser in
<taxonomicName id="4C254D52CE0DFFB7FBDEFB63FB0BFB58" authorityName="Antoine" authorityYear="1991" box="[1090,1209,1161,1187]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE0DFFB7FBDEFB63FB0BFB58" box="[1090,1209,1161,1187]" italics="true" pageId="10" pageNumber="101">G. ferreroi</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE0DFFB7FB5BFB63FAFCFB58" box="[1223,1358,1161,1187]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="10" pageNumber="101">Figs. 3031</figureCitation>
), outer line of parameres distal part evenly arcuate (
<figureCitation id="131E2A54CE0DFFB7FD15FB47FD6EFB3C" box="[649,732,1197,1223]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="10" pageNumber="101">Fig. 48</figureCitation>
; obtusely angled in
<taxonomicName id="4C254D52CE0DFFB7FC20FB47FB81FB3C" authorityName="Antoine" authorityYear="1991" box="[956,1075,1197,1223]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE0DFFB7FC20FB47FB81FB3C" box="[956,1075,1197,1223]" italics="true" pageId="10" pageNumber="101">G. ferreroi</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE0DFFB7FBDDFB47FB27FB33" box="[1089,1173,1197,1224]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="10" pageNumber="101">Fig. 46</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE0DFFB7FF5BFB3AFEFEFAAC" pageId="10" pageNumber="101" type="distribution">
<paragraph id="8B9A36D1CE0DFFB7FF5BFB3AFD7FFB10" blockId="10.[151,1437,152,1835]" box="[199,717,1232,1259]" pageId="10" pageNumber="101">
<emphasis id="B951EAC3CE0DFFB7FF5BFB3AFED3FB10" bold="true" box="[199,353,1232,1259]" pageId="10" pageNumber="101">Distribution.</emphasis>
<collectingCountry id="F3327641CE0DFFB7FEFBFB3BFE64FB10" box="[359,470,1233,1259]" name="Myanmar" pageId="10" pageNumber="101">Myanmar</collectingCountry>
,
<collectingCountry id="F3327641CE0DFFB7FE7DFB3BFDF5FB10" box="[481,583,1233,1259]" name="Thailand" pageId="10" pageNumber="101">Thailand</collectingCountry>
,
<collectingCountry id="F3327641CE0DFFB7FDCEFB3BFD7BFB10" box="[594,713,1233,1259]" name="Cambodia" pageId="10" pageNumber="101">Cambodia</collectingCountry>
.
</paragraph>
<paragraph id="8B9A36D1CE0DFFB7FF5BFB1FFEFEFAAC" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
<emphasis id="B951EAC3CE0DFFB7FF5BFB1FFEC5FAF4" bold="true" box="[199,375,1269,1295]" pageId="10" pageNumber="101">Nomenclature.</emphasis>
Following article 21.5 (
<bibRefCitation id="EFB44B20CE0DFFB7FD0EFB1FFCAEFAF4" author="ICZN" box="[658,796,1269,1295]" pageId="10" pageNumber="101" refId="ref20403" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London, 306 pp. Available from: https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / (accessed 9 March 2020)" type="book" year="1999">ICZN 1999</bibRefCitation>
), the date of
<taxonomicName id="4C254D52CE0DFFB7FC26FB1FFB72FAF4" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[954,1216,1269,1295]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FC26FB1FFB72FAF4" box="[954,1216,1269,1295]" italics="true" pageId="10" pageNumber="101">Goliathopsis despectus</emphasis>
</taxonomicName>
should be cited as 1873, because Westwoods
<taxonomicName id="4C254D52CE0DFFB7FE4EFAF3FCCCFAC8" authority="Entomologicus Oxoniensis" authorityName="Entomologicus Oxoniensis" box="[466,894,1305,1331]" class="Cyanophyceae" family="Thyssagaetidae" genus="Thesaurus" higherTaxonomySource="GBIF" kingdom="Bacteria" order="Anagliphonida" pageId="10" pageNumber="101" phylum="Cyanobacteria" rank="genus">
<emphasis id="B951EAC3CE0DFFB7FE4EFAF3FCCCFAC8" box="[466,894,1305,1331]" italics="true" pageId="10" pageNumber="101">Thesaurus Entomologicus Oxoniensis</emphasis>
</taxonomicName>
was published in four parts with different date (
<bibRefCitation id="EFB44B20CE0DFFB7FF03FAD7FE8DFAAC" author="Krikken, J." box="[159,319,1341,1367]" pageId="10" pageNumber="101" pagination="1 - 75" refId="ref20712" refString="Krikken, J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera. Zoologische Verhandelingen, 210, 1 - 75." type="journal article" year="1984">Krikken 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE0DFFB7FF5BFA8BFEF2F8D0" pageId="10" pageNumber="101" type="discussion">
<paragraph id="8B9A36D1CE0DFFB7FF5BFA8BFB18F98C" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
<emphasis id="B951EAC3CE0DFFB7FF5BFA8BFE89FA80" bold="true" box="[199,315,1377,1403]" pageId="10" pageNumber="101">Remarks.</emphasis>
When
<taxonomicName id="4C254D52CE0DFFB7FE0DFA8BFD24FA80" box="[401,662,1377,1403]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FE0DFA8BFD24FA80" box="[401,662,1377,1403]" italics="true" pageId="10" pageNumber="101">Goliathopsis capreolus</emphasis>
</taxonomicName>
was described,
<bibRefCitation id="EFB44B20CE0DFFB7FCCCFA8BFC5DFA80" author="Gestro, R." box="[848,1007,1377,1403]" pageId="10" pageNumber="101" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro (1888)</bibRefCitation>
has not realized that his new species actually is the third named taxon of the genus, and only compared it with
<taxonomicName id="4C254D52CE0DFFB7FC41FA6FFBFBFA64" authorityName="in Janson" authorityYear="1881" box="[989,1097,1413,1439]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE0DFFB7FC41FA6FFBFBFA64" box="[989,1097,1413,1439]" italics="true" pageId="10" pageNumber="101">G. cervus</emphasis>
</taxonomicName>
. Based on the examination of
<typeStatus id="549E8873CE0DFFB7FF0BFA43FF67FA38" box="[151,213,1449,1475]" pageId="10" pageNumber="101">types</typeStatus>
,
<bibRefCitation id="EFB44B20CE0DFFB7FF7CFA43FEC9FA38" author="Arrow, G. J." box="[224,379,1449,1475]" pageId="10" pageNumber="101" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
later regarded
<taxonomicName id="4C254D52CE0DFFB7FDBAFA43FD05FA38" box="[550,695,1449,1475]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FDBAFA43FD05FA38" box="[550,695,1449,1475]" italics="true" pageId="10" pageNumber="101">G. capreolus</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="4C254D52CE0DFFB7FC1CFA43FBA2FA38" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[896,1040,1449,1475]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FC1CFA43FBA2FA38" box="[896,1040,1449,1475]" italics="true" pageId="10" pageNumber="101">G. despectus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C254D52CE0DFFB7FB83FA43FA91FA38" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[1055,1315,1449,1475]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FB83FA43FA91FA38" box="[1055,1315,1449,1475]" italics="true" pageId="10" pageNumber="101">Goliathopsis despectus</emphasis>
</taxonomicName>
was originally described based on greasy females, and the tomentum on the body surface is indiscernible, while the tomentum of most
<typeStatus id="549E8873CE0DFFB7FF6DFA1BFE93F9F0" box="[241,289,1521,1547]" pageId="10" pageNumber="101">type</typeStatus>
specimens of
<taxonomicName id="4C254D52CE0DFFB7FE23FA1BFDFFF9F0" box="[447,589,1521,1547]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE0DFFB7FE23FA1BFDFFF9F0" box="[447,589,1521,1547]" italics="true" pageId="10" pageNumber="101">G. capreolus</emphasis>
</taxonomicName>
is in good condition. Nevertheless, the shape of setiferous punctures and the length of setae on the dorsal surface of the
<typeStatus id="549E8873CE0DFFB7FDE6F9FFFD07F9D4" box="[634,693,1557,1583]" pageId="10" pageNumber="101">types</typeStatus>
indicate that they are conspecific, and the appearance of the
<typeStatus id="549E8873CE0DFFB7FAFDF9FFFA2EF9D4" box="[1377,1436,1557,1583]" pageId="10" pageNumber="101">types</typeStatus>
of
<taxonomicName id="4C254D52CE0DFFB7FF29F9D3FEF6F9A8" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[181,324,1593,1619]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FF29F9D3FEF6F9A8" box="[181,324,1593,1619]" italics="true" pageId="10" pageNumber="101">G. despectus</emphasis>
</taxonomicName>
is obviously identical to the two greasy
<taxonomicName id="4C254D52CE0DFFB7FC94F9D3FC25F9A8" authorityName="Janson" authorityYear="1881" box="[776,919,1593,1619]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAC3CE0DFFB7FC94F9D3FC25F9A8" box="[776,919,1593,1619]" italics="true" pageId="10" pageNumber="101">Goliathopsis</emphasis>
</taxonomicName>
specimens in MFNB which were collected by Fea from
<collectingCountry id="F3327641CE0DFFB7FE98F9B7FEC1F98C" box="[260,371,1629,1655]" name="Myanmar" pageId="10" pageNumber="101">Myanmar</collectingCountry>
. Thus these
<typeStatus id="549E8873CE0DFFB7FE60F9B7FD9EF98C" box="[508,556,1629,1655]" pageId="10" pageNumber="101">type</typeStatus>
specimens of the two species are confirmed conspecific.
</paragraph>
<paragraph id="8B9A36D1CE0DFFB7FF5BF96BFEF2F8D0" blockId="10.[151,1437,152,1835]" pageId="10" pageNumber="101">
Apart from the specimens from the historical collections, no further records for
<taxonomicName id="4C254D52CE0DFFB7FBC9F96BFAE8F960" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[1109,1370,1665,1691]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE0DFFB7FBC9F96BFAE8F960" box="[1109,1370,1665,1691]" italics="true" pageId="10" pageNumber="101">Goliathopsis despectus</emphasis>
</taxonomicName>
up to now, and the actual distribution of this species is still unclear. According to his exploration route (
<bibRefCitation id="EFB44B20CE0DFFB7FB4BF94FFEB3F91F" author="Mouhot, M. H. &amp; Hodgkin, T." pageId="10" pageNumber="101" pagination="142 - 163" refId="ref21128" refString="Mouhot, M. H. &amp; Hodgkin, T. (1862) Notes on Cambodia, the Lao Country, &amp; c. The Journal of the Royal Geographical Society of London, 32, 142 - 163. https: // doi. org / 10.2307 / 1798407" type="journal article" year="1862">Mouhot &amp; Hodgkin 1862</bibRefCitation>
), the Cambodian specimens collected by Henri Mouhot (
<date id="FF9B1011CE0DFFB7FC09F923FBA7F91F" box="[917,1045,1737,1764]" pageId="10" pageNumber="101" value="1826" valueMax="1861">18261861</date>
) probably from the northern part, while the collecting site of the specimens from
<collectingCountry id="F3327641CE0DFFB7FD35F907FCBFF8FC" box="[681,781,1773,1799]" name="Thailand" pageId="10" pageNumber="101">Thailand</collectingCountry>
cannot be presumed due to Mouhot explored several parts of this country.
</paragraph>
</subSubSection>
</treatment>
</document>