treatments-xml/data/03/8C/87/038C87BF206C1904FEB48BF7D8DEFB63.xml
2024-06-21 12:22:17 +02:00

729 lines
108 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6CAC8F312824A28A51CD141EDC4DCD12" ID-CLB-Dataset="152185" ID-DOI="10.18590/euscorpius.2014.vol2014.iss191.1" ID-GBIF-Dataset="a6381f1d-22e1-4629-b369-07f5ab1d7834" ID-ISSN="1536-9307" ID-Zenodo-Dep="6505622" ID-ZooBank="E467B3C0-D693-4EAF-B5F0-759D8C63FE35" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638245227176" checkinUser="felipe" docAuthor="Lowe, Graeme, Yağmur, Ersen Aydın &amp; Kovařík, František" docDate="2014" docId="038C87BF206C1904FEB48BF7D8DEFB63" docLanguage="en" docName="Euscorpius.191.1-129.pdf" docOrigin="Euscorpius 191" docSource="https://doi.org/10.18590/euscorpius.2014.vol2014.iss191.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Leiurus heberti Lowe &amp; Yağmur &amp; Kovařík 2014, sp. n." docType="treatment" docUuid="2E5906DA-1108-4669-B2B7-8831B60EA4D1" docUuidSource="ZooBank" docVersion="14" lastPageNumber="73" masterDocId="FFB5FFC72052194EFFD78D2ADD40FF87" masterDocTitle="A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula" masterLastPageNumber="129" masterPageNumber="1" pageNumber="61" updateTime="1699471136939" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="1C452C71A368C1AEDABE618D84BE928F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E08CFEFF0B3BA46A71836D9385600A54">
<mods:title id="02EE79E721F2C0B7277D1B497B339DB7">A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula</mods:title>
</mods:titleInfo>
<mods:name id="38BBD831FA5430577D7DFBD62F9A9EA3" type="personal">
<mods:role id="D0BEFDCCBA63A928B18B9E305A528E9B">
<mods:roleTerm id="9902C043F94E90EA89963F52E8793BF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8DC2FFFA96761D89536E77BB4979759">Lowe, Graeme</mods:namePart>
<mods:affiliation id="795CFC4D072D18BC42DD562C67DD9AC8">Monell Chemical Senses Center, 3500 Market St, Philadelphia, USA</mods:affiliation>
</mods:name>
<mods:name id="3CCC756B56BC106C8F23DBC95105BABC" type="personal">
<mods:role id="B3D7013D512F81D886FD653D5991211E">
<mods:roleTerm id="357E108E03CDDB1950148AFAE64F612E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD0BC3E28B19431BEC906A1C5C5A6A9B">Yağmur, Ersen Aydın</mods:namePart>
<mods:affiliation id="777C784C8400EAE27A026BAB380B4155">Alaşehir Vocational School, Celal Bayar University, Manisa, Turkey</mods:affiliation>
</mods:name>
<mods:name id="B84365CC45D740863D4DE3351F8766E7" type="personal">
<mods:role id="633E521CF7D8B34967A89B361EE3F166">
<mods:roleTerm id="58401C035D557B8C2327FB0548080336">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2A28C87C3CEC46D1F676F5244CFB633">Kovařík, František</mods:namePart>
<mods:affiliation id="BDA16C92D69FA84F893E23560607940E">P. O. Box 27, CZ - 145 01, Praha 45, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="148956F83BF8ACA81295E6429E4F64DF" type="URL">www.kovarex.com/scorpio</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="43F41FDDC749CCCC71C74D9F8404743B">text</mods:typeOfResource>
<mods:relatedItem id="6798F3E8BD6AAE938A30C299E9A86C56" type="host">
<mods:titleInfo id="6E6982D91F124F0741C82D7298798DFC">
<mods:title id="6518A2542E68508CE1EF5F93BD2C5B6D">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="6523D70A6CD51720C3C631F962FEC35F">
<mods:date id="4D031B8B1E4E0AA7F2C08AFBED8278BC">2014</mods:date>
<mods:detail id="B4370D63F7BAAA6362492391BDF46AD5" type="pubDate">
<mods:number id="642C6259FE4698124DA1F1B354EC4129">2014-12-31</mods:number>
</mods:detail>
<mods:detail id="FD4828EEC5A764465B52926FADB154F6" type="volume">
<mods:number id="5594E0732BBCB0F627E2EBC2449BE22A">191</mods:number>
</mods:detail>
<mods:extent id="8CCBE64D67FB3262EDA783633632107D" unit="page">
<mods:start id="C7D60F49C59E31C119CDB500290104B8">1</mods:start>
<mods:end id="03E88511A78A7859C10F07A640506178">129</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="77C929238553F4EE10A85F4D2A2C7DF7">
<mods:url id="880BCFE168DD00B70A6DACC1777E20A5">https://doi.org/10.18590/euscorpius.2014.vol2014.iss191.1</mods:url>
</mods:location>
<mods:classification id="69349F931483E43A152C0B65D11ABA95">journal article</mods:classification>
<mods:identifier id="290DC07861B91FC02CBBF18748FDF80F" type="CLB-Dataset">152185</mods:identifier>
<mods:identifier id="FEB2543D989B064CC74D3542FB21F9D6" type="DOI">10.18590/euscorpius.2014.vol2014.iss191.1</mods:identifier>
<mods:identifier id="6F085C7678E26D36D7F32D226DA9B1D4" type="GBIF-Dataset">a6381f1d-22e1-4629-b369-07f5ab1d7834</mods:identifier>
<mods:identifier id="09B5D0F16D1A8DDCB0752112A44DAD5B" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="40CBA801E92FBA14E6D19536A7360E22" type="Zenodo-Dep">6505622</mods:identifier>
<mods:identifier id="23AEFC18459D2F40CD19B5C851C9929C" type="ZooBank">E467B3C0-D693-4EAF-B5F0-759D8C63FE35</mods:identifier>
</mods:mods>
<treatment id="038C87BF206C1904FEB48BF7D8DEFB63" ID-DOI="http://doi.org/10.5281/zenodo.7117239" ID-GBIF-Taxon="201133255" ID-Zenodo-Dep="7117239" LSID="urn:lsid:zoobank.org:act:2E5906DA-1108-4669-B2B7-8831B60EA4D1" httpUri="http://treatment.plazi.org/id/038C87BF206C1904FEB48BF7D8DEFB63" lastPageId="74" lastPageNumber="73" pageId="62" pageNumber="61">
<subSubSection id="C33F6522206C1970FEB48BF7DF3EF97D" box="[355,638,1757,1786]" pageId="62" pageNumber="61" type="nomenclature">
<paragraph id="8B9A36A9206C1970FEB48BF7DF3EF97D" blockId="62.[198,795,1757,1908]" box="[355,638,1757,1786]" pageId="62" pageNumber="61">
<heading id="D0D281C5206C1970FEB48BF7DF3EF97D" box="[355,638,1757,1786]" centered="true" fontSize="11" level="2" pageId="62" pageNumber="61" reason="2">
<emphasis id="B951EABB206C1970FEB48BF7DF3EF97D" bold="true" box="[355,638,1757,1786]" pageId="62" pageNumber="61">
<taxonomicName id="4C254D2A206C1970FEB48BF7DF6EF97D" ID-CoL="6P795" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[355,558,1757,1786]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="62" pageNumber="61" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB206C1970FEB48BF7DF6EF97D" bold="true" box="[355,558,1757,1786]" italics="true" pageId="62" pageNumber="61">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206C1970FDE08BF7DF3EF97D" box="[567,638,1757,1786]" pageId="62" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6522206C1970FF058BD5DF85F8F3" pageId="62" pageNumber="61" type="description">
<paragraph id="8B9A36A9206C1970FF058BD5DF85F8F3" blockId="62.[198,795,1757,1908]" pageId="62" pageNumber="61">
(
<figureCitation id="131E2A2C206C1970FF0D8BD5DC1BF890" box="[218,347,1791,1815]" captionStart-0="Figure 47" captionStart-1="Figure 48" captionStart-10="Figure 57" captionStart-2="Figure 49" captionStart-3="Figure 50" captionStart-4="Figure 51" captionStart-5="Figure 52" captionStart-6="Figure 53" captionStart-7="Figure 54" captionStart-8="Figure 55" captionStart-9="Figure 56" captionStartId-0="64.[192,267,1880,1904]" captionStartId-1="65.[192,267,1878,1902]" captionStartId-10="74.[192,267,944,968]" captionStartId-2="66.[192,267,1295,1319]" captionStartId-3="67.[192,267,1672,1696]" captionStartId-4="68.[192,267,1379,1403]" captionStartId-5="69.[192,267,1351,1375]" captionStartId-6="70.[192,267,1279,1303]" captionStartId-7="71.[192,267,1593,1617]" captionStartId-8="72.[192,267,1790,1814]" captionStartId-9="73.[192,267,899,923]" captionTargetBox-0="[333,1297,216,1849]" captionTargetBox-1="[321,1285,215,1846]" captionTargetBox-10="[223,766,203,893]" captionTargetBox-2="[204,1427,219,1253]" captionTargetBox-3="[191,1440,170,1641]" captionTargetBox-4="[191,1437,191,1349]" captionTargetBox-5="[191,1440,191,1321]" captionTargetBox-6="[204,1426,207,1238]" captionTargetBox-7="[191,1439,167,1563]" captionTargetBox-8="[286,1333,215,1760]" captionTargetBox-9="[191,1440,191,869]" captionTargetId-0="figure-25@64.[334,1296,217,1848]" captionTargetId-1="figure-21@65.[322,1284,216,1845]" captionTargetId-10="figure-746@74.[179,803,193,915]" captionTargetId-2="figure-317@66.[192,1439,206,1265]" captionTargetId-3="figure-132@67.[192,1439,183,1640]" captionTargetId-4="figure-277@68.[192,1436,192,1348]" captionTargetId-5="figure-299@69.[192,1439,192,1320]" captionTargetId-6="figure-350@70.[192,1438,192,1250]" captionTargetId-7="figure-156@71.[192,1438,169,1563]" captionTargetId-8="figure-45@72.[287,1332,216,1759]" captionTargetId-9="figure-574@73.[192,1439,192,868]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-10="74" captionTargetPageId-2="66" captionTargetPageId-3="67" captionTargetPageId-4="68" captionTargetPageId-5="69" captionTargetPageId-6="70" captionTargetPageId-7="71" captionTargetPageId-8="72" captionTargetPageId-9="73" captionText-0="Figure 47: Leiurus heberti sp. n., holotype male. Habitus, dorsal aspect. Wadi Andur, Oman." captionText-1="Figure 48: Leiurus heberti sp. n., holotype male. Habitus, ventral aspect. Wadi Andur, Oman." captionText-10="Figure 57: Leiurus heberti sp. n., holotype male. Trichobothrial map of pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, external aspect. E. Chela, ventral aspect. Wadi Andur, Oman." captionText-2="Figure 49: Leiurus heberti sp. n., holotype male. A. Carapace and tergites. B. Coxosternal area and sternites. Wadi Andur, Oman." captionText-3="Figure 50: Leiurus heberti sp. n., holotype male. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Fixed finger dentition. G. Movable finger dentition. H. Chela, external aspect. Wadi Andur, Oman." captionText-4="Figure 51: Leiurus heberti sp. n., paratype female. Habitus, dorsal aspect. Jabal Samhan, Oman." captionText-5="Figure 52: Leiurus heberti sp. n., paratype female. Habitus, ventral aspect. Jabal Samhan, Oman." captionText-6="Figure 53: Leiurus heberti sp. n., paratype female. A. Carapace and tergites. B. Coxosternal area and sternites. Jabal Samhan, Oman." captionText-7="Figure 54: Leiurus heberti sp. n., paratype female. Right pedipalp. A. Femur, dorsal aspect. B. Patella, dorsal aspect. C. Patella, external aspect. D. Chela, ventral aspect. E. Chela, dorsal aspect. F. Chela, external aspect. Jabal Samhan, Oman." captionText-8="Figure 55: Leiurus heberti sp. n. Metasoma. A. Male, lateral aspect. B. Male, ventral aspect. C. Female, lateral aspect. D. Female, ventral aspect. Male holotype, Wadi Andur, Oman. Female paratype, Jabal Samhan, Oman." captionText-9="Figure 56: Leiurus heberti sp. n., holotype male. Tarsi. A-D. Left telotarsus and distal basitarsus, ventral aspect. A. Leg I. B. Leg II. C. Leg III. D. Leg IV. E. Right basitarsus III retrolateral aspect. Upper scale bar: AD, lower scale bar: E. Wadi Andur, Oman." figureDoi-0="http://doi.org/10.5281/zenodo.6505723" figureDoi-1="http://doi.org/10.5281/zenodo.6505725" figureDoi-10="http://doi.org/10.5281/zenodo.6505743" figureDoi-2="http://doi.org/10.5281/zenodo.6505727" figureDoi-3="http://doi.org/10.5281/zenodo.6505729" figureDoi-4="http://doi.org/10.5281/zenodo.6505731" figureDoi-5="http://doi.org/10.5281/zenodo.6505733" figureDoi-6="http://doi.org/10.5281/zenodo.6505735" figureDoi-7="http://doi.org/10.5281/zenodo.6505737" figureDoi-8="http://doi.org/10.5281/zenodo.6505739" figureDoi-9="http://doi.org/10.5281/zenodo.6505741" httpUri-0="https://zenodo.org/record/6505723/files/figure.png" httpUri-1="https://zenodo.org/record/6505725/files/figure.png" httpUri-10="https://zenodo.org/record/6505743/files/figure.png" httpUri-2="https://zenodo.org/record/6505727/files/figure.png" httpUri-3="https://zenodo.org/record/6505729/files/figure.png" httpUri-4="https://zenodo.org/record/6505731/files/figure.png" httpUri-5="https://zenodo.org/record/6505733/files/figure.png" httpUri-6="https://zenodo.org/record/6505735/files/figure.png" httpUri-7="https://zenodo.org/record/6505737/files/figure.png" httpUri-8="https://zenodo.org/record/6505739/files/figure.png" httpUri-9="https://zenodo.org/record/6505741/files/figure.png" pageId="62" pageNumber="61">Figs. 4757</figureCitation>
,
<figureCitation id="131E2A2C206C1970FEB18BD5DCD5F890" box="[358,405,1791,1815]" captionStart="Figure 58" captionStartId="76.[192,267,1207,1231]" captionTargetBox="[191,1437,213,1177]" captionTargetId="figure-403@76.[192,1436,215,1177]" captionTargetPageId="76" captionText="Figure 58: Hemispermatophores of new Leiurus species. A. L. macroctenus sp. n., paratype male, Duhai, Oman. B. L. heberti sp. n., holotype male, Wadi Andur, Oman. C. L. haenggii sp. n., paratype male, Jabal Qara, Oman. Distal portions hemispermatophores illustrated to show flagellum and lobes (right hemispermatophores, convex aspect). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6505745" httpUri="https://zenodo.org/record/6505745/files/figure.png" pageId="62" pageNumber="61">58B</figureCitation>
,
<figureCitation id="131E2A2C206C1970FE778BD5DCB1F890" box="[416,497,1791,1815]" captionStart="Figure 59" captionStartId="77.[192,267,1302,1326]" captionTargetBox="[200,1431,198,1264]" captionTargetId="figure-354@77.[192,1439,192,1272]" captionTargetPageId="77" captionText="Figure 59: Chelicerae of Leiurus species. A, B. L. macroctenus sp. n., paratype male, Thumrait, Oman. Right chelicera, ventral (A) and dorsal (B) aspect. C, D. L. haenggii sp. n., paratype male, Jabal Qara, Oman. Right chelicera, ventral (C) and dorsal (D) aspect. E, F. L. brachycentrus (Ehrenberg, 1829), female, Al Mansuriah, Yemen. Right chelicera, ventral (E) and dorsal (F) aspect. G, H. L. heberti sp. n., holotype male, Wadi Andur, Oman. Right chelicera, ventral (G) and dorsal (H) aspect. Scale bar in A: 1 mm (also applies to BH)." figureDoi="http://doi.org/10.5281/zenodo.6505747" httpUri="https://zenodo.org/record/6505747/files/figure.png" pageId="62" pageNumber="61">59GH</figureCitation>
,
<figureCitation id="131E2A2C206C1970FE2B8BD5DF69F890" box="[508,553,1791,1815]" captionStart="Figure 87" captionStartId="107.[192,267,1798,1822]" captionTargetBox="[343,1333,196,1766]" captionTargetId="figure-90@107.[344,1333,197,1765]" captionTargetPageId="107" captionText="Figure 87: Metasoma V and telson of Leiurus spp. Males, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505806" httpUri="https://zenodo.org/record/6505806/files/figure.png" pageId="62" pageNumber="61">87E</figureCitation>
,
<figureCitation id="131E2A2C206C1970FDE38BD5DF21F890" box="[564,609,1791,1815]" captionStart="Figure 88" captionStartId="108.[192,267,1816,1840]" captionTargetBox="[318,1309,215,1785]" captionTargetId="figure-95@108.[319,1308,216,1784]" captionTargetPageId="108" captionText="Figure 88: Metasoma V and telson of Leiurus spp. Males, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n.. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n.. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n.. G. L. macroctenus sp. n.. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505808" httpUri="https://zenodo.org/record/6505808/files/figure.png" pageId="62" pageNumber="61">88E</figureCitation>
,
<figureCitation id="131E2A2C206C1970FDBB8BD5DFDAF890" box="[620,666,1791,1815]" captionStart="Figure 89" captionStartId="109.[192,267,1809,1833]" captionTargetBox="[345,1336,208,1778]" captionTargetId="figure-94@109.[346,1335,209,1777]" captionTargetPageId="109" captionText="Figure 89: Metasoma V and telson of Leiurus spp. Females, lateral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505810" httpUri="https://zenodo.org/record/6505810/files/figure.png" pageId="62" pageNumber="61">89E</figureCitation>
,
<figureCitation id="131E2A2C206C1970FD738BD5DF91F890" box="[676,721,1791,1815]" captionStart="Figure 90" captionStartId="110.[192,267,1808,1832]" captionTargetBox="[318,1309,207,1777]" captionTargetId="figure-95@110.[319,1308,208,1776]" captionTargetPageId="110" captionText="Figure 90: Metasoma V and telson of Leiurus spp. Females, ventral aspect. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009. B. L. arabicus sp. n. C. L. brachycentrus (Ehrenberg, 1829) stat. n.. D. L. haenggii sp. n. E. L. heberti sp. n. F. L. hebraeus (Birula, 1908) stat. n. G. L. macroctenus sp. n. H. L. quinquestriatus (Ehrenberg, 1828). Locality data as in Figs. 9, 20, 31, 43, 55, 67, 79. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505812" httpUri="https://zenodo.org/record/6505812/files/figure.png" pageId="62" pageNumber="61">90E</figureCitation>
,
<figureCitation id="131E2A2C206C1970FD0B8BD5DE4CF890" box="[732,780,1791,1815]" captionStart="Figure 91" captionStartId="111.[192,267,1766,1790]" captionTargetBox="[303,1321,191,1734]" captionTargetId="figure-124@111.[304,1320,192,1733]" captionTargetPageId="111" captionText="Figure 91: Pigmentation patterns of metasoma IV, V of Leiurus species, ventral aspect. A. L. haenggii sp. n., holotype female, Taif, Saudi Arabia (NHMB 17k). B. L. macroctenus sp. n., holotype male, Thumrait, Oman. C. L. heberti sp. n., holotype male, Wadi Andur, Oman. D. L. quinquestriatus (Ehrenberg, 1828), female, Cairo, Egypt (NHMB 17d). E. L. brachycentrus (Ehrenberg, 1829) stat. n., male, Ad Darb, Saudi Arabia (NHMB 17ag). F. L. arabicus sp. n., holotype female, Kushm Dibi, Saudi Arabia (NHMB 17aq). G. L. hebraeus (Birula, 1908) stat. n., female, Israel (NHMB 17a). H. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, male, Eski Sarkaya Village, Turkey. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6505814" httpUri="https://zenodo.org/record/6505814/files/figure.png" pageId="62" pageNumber="61">91C</figureCitation>
,
<figureCitation id="131E2A2C206C1970FECE8A34DC0AF8B1" box="[281,330,1822,1846]" captionStart="Figure 92" captionStartId="112.[192,267,1486,1510]" captionTargetBox="[224,1390,157,1461]" captionTargetId="figure-274@112.[212,1402,145,1473]" captionTargetPageId="112" captionText="Figure 92: Sternite III of Leiurus spp., female. A. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. B. L. arabicus sp. n., holotype, Kushm Dibi, Saudi Arabia (NHMB 17aq). C. L. arabicus sp. n., paratype, Al Amar, Bahrain (NHMB 17bg). D. L. heberti sp. n., paratype, Jabal Samhan. Oman. E. L. haenggii sp. n., paratype, Wadi Turabah, Saudi Arabia (NHMB 17al). F. L. haenggii sp. n., paratype, Adnan, Saudi Arabia (NHMB 17am). G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. macroctenus sp. n., paratype, Thumrait, Oman. J. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). K. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505816" httpUri="https://zenodo.org/record/6505816/files/figure.png" pageId="62" pageNumber="61">92D</figureCitation>
,
<figureCitation id="131E2A2C206C1970FE828A34DCC2F8B1" box="[341,386,1822,1846]" captionStart="Figure 93" captionStartId="113.[192,267,1572,1596]" captionTargetBox="[224,1390,195,1538]" captionTargetId="figure-226@113.[212,1402,183,1550]" captionTargetPageId="113" captionText="Figure 93: Medial intercarinal areas of tergite III of female Leiurus spp. A. L. arabicus sp. n., paratype, Jeddah, Saudi Arabia. B. L. arabicus sp. n., paratype, Wadi Khumrah, Saudi Arabia (NHMB 17bj). C. L. haenggii sp. n., paratype, Yemen. D. L. haenggii sp. n., paratype, Wadi Maraba, Saudi Arabia (NHMB 17al). E. L. heberti sp. n., paratype, Jabal Samhan. Oman. F. L. abdullahbayrami Yağmur, Koç et Kunt, 2009, Caybasi Village, Turkey. G. L. brachycentrus (Ehrenberg, 1829) stat. n., Al Mansuriah, Yemen. H. L. macroctenus sp. n., paratype, Masirah Island, Oman. I. L. hebraeus (Birula, 1908) stat. n., Israel (NHMB 17a). J. L. hebraeus Birula, 1908, Kurayyima, Jordan. K. L. quinquestriatus (Ehrenberg, 1828), Kharga Oasis, Egypt (NHMB 17i). L. L. jordanensis Lourenço, Modry et Amr, 2002, al-Tawil, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.6505818" httpUri="https://zenodo.org/record/6505818/files/figure.png" pageId="62" pageNumber="61">93E</figureCitation>
,
<figureCitation id="131E2A2C206C1970FE5A8A34DC9CF8B1" box="[397,476,1822,1846]" captionStart="Figure 95" captionStartId="118.[192,260,1217,1238]" captionTargetBox="[214,1409,159,1149]" captionTargetId="figure-476@118.[192,1437,128,1181]" captionTargetPageId="118" captionText="Figure 95: Variation in key diagnostic biometrics of adult Leiurus spp. A. Cumulative distributions of position of trichobothrium db relative to est on pedipalp fixed finger for: L. abdullahbayrami (53), L. brachycentrus stat. n. (11), L. hebraeus stat. n. (38), L. macroctenus sp. n. (176), L. quinquestriatus (61), L. haenggii sp. n. (158), L. arabicus sp. n. (87) (number of fingers measured in parentheses). Data for males and females were pooled. The dbest distances were normalized to pedipalp movable finger length (chord length from finger tip to external articular condyle), with positive values indicating db distal to est, negative values indicating db proximal to est. Distributions of L. haenggii (µ = 0.06367) and L. arabicus sp. n. (µ = 0.06671) were not significantly different (P = 0.405148, t test), indicating a close relationship between these species. Distribution of L. quinquestriatus (µ = 0.047313) was significantly different from those of L. haenggii and L. arabicus (P = 0.000102 and 0.000030 respectively). B. Scatter plot of morphometric ratios of mid-pectine sensillar margin L (MPSM, indicated in inset) to metasoma I W, and to carapace L, showing differences in relative pectinal tooth size in females of nine species of Leiurus. C. A subset of the female data in Figure 95B, plotted as cumulative distributions of morphometric ratio of mid-pectine sensillar margin L to metasoma I W. Distributions of L. haenggii (µ = 0.102268) and L. arabicus (µ = 0.10432) were not significantly different (P&gt; 0.1, K-S test). D. Scatter plot of morphometric ratios of mid-pectine sensillar margin L to metasoma I W (relative size of pectine teeth), and metasoma II L/W (slenderness of metasoma II) for females of nine species of Leiurus. A, C, D: symbol key as in Figure 95B." figureDoi="http://doi.org/10.5281/zenodo.6505822" httpUri="https://zenodo.org/record/6505822/files/figure.png" pageId="62" pageNumber="61">95BD</figureCitation>
,
<figureCitation id="131E2A2C206C1970FE308A34DF79F8B1" box="[487,569,1822,1846]" captionStart-0="Figure 98" captionStart-1="Figure 99" captionStart-2="Figure 100" captionStartId-0="121.[192,267,1251,1275]" captionStartId-1="122.[192,267,1014,1038]" captionStartId-2="123.[192,267,1084,1108]" captionTargetBox-0="[215,1377,224,1220]" captionTargetBox-1="[191,1438,222,978]" captionTargetBox-2="[191,1441,177,1051]" captionTargetId-0="figure-474@121.[192,1437,206,1246]" captionTargetId-1="figure-479@122.[192,1437,223,978]" captionTargetId-2="figure-335@123.[192,1440,178,1050]" captionTargetPageId-0="121" captionTargetPageId-1="122" captionTargetPageId-2="123" captionText-0="Figure 98: Selected morphometrics of Leiurus, Cicileiurus, Cicileus and Compsobuthus compared to other Buthus group scorpions. AB. Scatter plots of the fraction of fixed finger length distal to db (A) and est (B) vs. the ratio of movable finger length to carapace length. Each point represents one sex of one species. Larger ordinate values correspond to more basal positions of the trichobothria, and larger abscissa values to longer pedipalp fingers. There was a significant inverse correlation between relative length of the portion of the fixed finger distal to db and est (R = -0.6534, -0.5488, respectively), and the relative length of the movable finger (the latter being a measure of elongation of both fixed and movable fingers). Highlighted symbols show that Leiurus (light magenta circles), Cicileiurus (red triangle), Cicileus (green squares) and Compsobuthus (yellow circles) are located in the lower right halves of the plots, i.e. all have relatively elongated fingers and more distal placement of both db and est. Gray circles are data from other Buthus group species. C. Scatter plot of the fraction of fixed finger length distal to db vs. the fraction distal to est. The strong positive correlation (R = 0.8052) indicates a tendency for db and est to move together towards more distal locations as the fixed finger becomes more elongated. Species above the diagonal (solid blue) have db proximal to est, and those below have db distal to est. Solid gray lines in AC are fits by least squares regression through all points. D. Scatter plot of pedipalp femur L/W (a measure of pedipalp elongation) vs. carapace L (a measure of body size). These two variables were not significantly correlated (R = 0.094). Data were compiled from the literature and specimens in the authors collections for 38 genera and 203 species representing the majority of taxa in the Buthus group, including both males (N = 124) and females (N = 97). Genera (and number of species) included: Afghanobuthus (1), Androctonus (11), Apistobuthus (2), Baloorthochirus (1), Birulatus (2), Buthacus (13), Butheoloides (11), Butheolus (5), Buthiscus (1), Buthus (26), Cicileiurus (1), Cicileus (2), Compsobuthus (33), Congobuthus (1), Darchenia (1), Gint (1), Hemibuthus (1), Hottentotta (31), Leiurus (10), Liobuthus (1), Lissothus (3), Mesobuthus (7), Neobuthus (2), Odontobuthus (6), Orthochirus (12), Pantobuthus (1), Pectinibuthus (1), Plesiobuthus (1), Polisius (1), Razianus (3), Saharobuthus (1), Somalibuthus (1), Vachoniolus (4), Vachonus (1)." captionText-1="Figure 99: Geographic plot of locality data of material examined for nine species of Leiurus distributed over northeast Africa, the Levant and Arabian Peninsula. Map colored by elevation with shaded relief. Additional locality data for: L. abdullahbayrami from Yağmur et al., 2009; for L. brachycentrus Ehrenberg, 1829 stat. n., from Simon, 1882 (Buthus beccarii)." captionText-2="Figure 100: Geographic plot of locality data of material examined for five species of Leiurus from the Arabian Peninsula. Map colored by terrain with shaded relief, indicating major physiographic regions." figureDoi-0="http://doi.org/10.5281/zenodo.6505828" figureDoi-1="http://doi.org/10.5281/zenodo.6505832" figureDoi-2="http://doi.org/10.5281/zenodo.6505834" httpUri-0="https://zenodo.org/record/6505828/files/figure.png" httpUri-1="https://zenodo.org/record/6505832/files/figure.png" httpUri-2="https://zenodo.org/record/6505834/files/figure.png" pageId="62" pageNumber="61">98100</figureCitation>
,
<tableCitation id="C6A70312206C1970FD928A34DFD5F8B1" box="[581,661,1822,1846]" captionStart="Table 2" captionStartId="75.[192,257,682,706]" captionTargetBox="[319,1283,200,646]" captionText="Table 2: Variation in male morphometric ratios with increasing size in Leiurus heberti sp. n." httpUri="http://table.plazi.org/id/DF5A662120191905FF178F80D909FD46" pageId="62" pageNumber="61" tableUuid="DF5A662120191905FF178F80D909FD46">Tabs. 2</tableCitation>
,
<tableCitation id="C6A70312206C1970FD768A34DF80F8B1" box="[673,704,1822,1846]" captionStart="Table 3" captionStartId="105.[192,257,1349,1373]" captionTargetBox="[209,1890,232,1297]" captionTargetId="graphics-7@105.[194,1919,192,1298]" captionText="Table 3A: Summary of key characters separating four Leiurus species examined in this study (adult females unless otherwise specified). Numeric values indicate ranges with sample sizes in parentheses." httpUri="http://table.plazi.org/id/DF5A6621203B1927FF17886FDC8EFAF2" pageId="62" pageNumber="61" tableUuid="DF5A6621203B1927FF17886FDC8EFAF2">3B</tableCitation>
)
<emphasis id="B951EABB206C1970FF118A14DF85F8F3" bold="true" pageId="62" pageNumber="61">http://zoobank.org/urn:lsid:zoobank.org:act:2E5906 DA-1108-4669-B2B7-8831B60EA4D1</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F6522206C1970FCE88841D818F953" pageId="62" pageNumber="61" type="materials_examined">
<paragraph id="8B9A36A9206C1970FCE88841D821FA58" blockId="62.[831,1440,1387,1503]" pageId="62" pageNumber="61">
<materialsCitation id="3B4D3CF4206C1970FCE88841D821FA58" ID-GBIF-Occurrence="3913646427" collectingDate="1995-09-26" collectingMethod="UV detection" collectionCode="NHMB" collectorName="G. Lowe &amp; M. D. Gallagher" country="Oman" elevation="405" latitude="17.668" location="rocky boulder wadi, along base of cliff, east edge of wadi" longLatPrecision="13" longitude="54.6545" municipality="Wadi Andur" pageId="62" pageNumber="61" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549E880B206C1970FCE88841DE8FFA04" box="[831,975,1387,1411]" pageId="62" pageNumber="61" type="holotype">HOLOTYPE</typeStatus>
. Adult
<specimenCount id="9D23FD20206C1970FBF78841D976FA05" box="[1056,1078,1387,1410]" pageId="62" pageNumber="61" type="male"></specimenCount>
,
<emphasis id="B951EABB206C1970FB948841D9CAFA04" bold="true" box="[1091,1162,1387,1411]" pageId="62" pageNumber="61">
<collectingCountry id="F3327639206C1970FB948841D9CAFA04" box="[1091,1162,1387,1411]" name="Oman" pageId="62" pageNumber="61">Oman</collectingCountry>
</emphasis>
,
<collectingMunicipality id="6BFEACD3206C1970FB4D8841D863FA04" box="[1178,1315,1387,1411]" pageId="62" pageNumber="61">Wadi Andur</collectingMunicipality>
,
<collectingMethod id="52644EBE206C1970FAF88841DE2BFA26" pageId="62" pageNumber="61">UV detection</collectingMethod>
,
<location id="8EFA6072206C1970FCA288A3DE35FA47" LSID="urn:lsid:plazi:treatment:038C87BF206C1904FEB48BF7D8DEFB63:8EFA6072206C1970FCA288A3DE35FA47" country="Oman" latitude="17.668" longLatPrecision="13" longitude="54.6545" municipality="Wadi Andur" name="rocky boulder wadi, along base of cliff, east edge of wadi" pageId="62" pageNumber="61">rocky boulder wadi, along base of cliff, east edge of wadi</location>
,
<geoCoordinate id="EE11506E206C1970FC548882DEBDFA47" box="[899,1021,1448,1472]" degrees="17" direction="north" minutes="40.08" orientation="latitude" pageId="62" pageNumber="61" precision="9" value="17.668">17°40.08'N</geoCoordinate>
<geoCoordinate id="EE11506E206C1970FBD18882D93EFA47" box="[1030,1150,1448,1472]" degrees="54" direction="east" minutes="39.27" orientation="longitude" pageId="62" pageNumber="61" precision="9" value="54.6545">54°39.27'E</geoCoordinate>
,
<elevation id="0008D19A206C1970FB5A8882D84FFA47" box="[1165,1295,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="62" pageNumber="61" unit="m" value="405.0">
<quantity id="4CDD9B4C206C1970FB5A8882D993FA47" box="[1165,1235,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="62" pageNumber="61" unit="m" value="405.0">405 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B1069206C1970FAC88882D8DAFA47" box="[1311,1434,1448,1472]" pageId="62" pageNumber="61" value="1995-09-26">
<collectingDate id="EFDFE981206C1970FAC88882D8DAFA47" box="[1311,1434,1448,1472]" pageId="62" pageNumber="61" value="1995-09-26">26.IX.1995</collectingDate>
</date>
, 22:50 h, leg.
<collectorName id="26D0537F206C1970FC1C88EDD967FA59" box="[971,1063,1479,1503]" pageId="62" pageNumber="61">G. Lowe</collectorName>
&amp;
<collectorName id="26D0537F206C1970FB9E88EDD9B2FA58" box="[1097,1266,1479,1503]" pageId="62" pageNumber="61">M.D. Gallagher</collectorName>
(
<collectionCode id="ED34AE6C206C1970FAD788EDD816FA58" box="[1280,1366,1479,1503]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="62" pageNumber="61">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A36A9206C1970FCE88B2ED818F953" blockId="62.[831,1440,1540,1748]" pageId="62" pageNumber="61">
<typeStatus id="549E880B206C1970FCE88B2EDE9BF99B" box="[831,987,1540,1564]" pageId="62" pageNumber="61" type="paratype">PARATYPES</typeStatus>
.
<materialsCitation id="3B4D3CF4206C1970FC3F8B2ED964F910" ID-GBIF-Occurrence="3913646305" collectingDate="1995-09-26" collectingMethod="UV detection" collectionCode="NHMB" collectorName="G. Lowe &amp; M. D. Gallagher" country="Oman" elevation="405" latitude="17.668" location="rocky boulder wadi, along base of cliff, east edge of wadi" longLatPrecision="13" longitude="54.6545" municipality="Wadi Andur" pageId="62" pageNumber="61" specimenCount="2" typeStatus="paratype">
<emphasis id="B951EABB206C1970FC3F8B2ED977F99B" bold="true" box="[1000,1079,1540,1564]" pageId="62" pageNumber="61">
<collectingCountry id="F3327639206C1970FC3F8B2ED973F99B" box="[1000,1075,1540,1564]" name="Oman" pageId="62" pageNumber="61">Oman</collectingCountry>
:
</emphasis>
<specimenCount id="9D23FD20206C1970FB968B2ED9EDF99B" box="[1089,1197,1540,1564]" pageId="62" pageNumber="61">1 adult ♂</specimenCount>
,
<specimenCount id="9D23FD20206C1970FB6C8B2ED817F99B" box="[1211,1367,1540,1564]" pageId="62" pageNumber="61">1 immature ♂</specimenCount>
,
<collectingMunicipality id="6BFEACD3206C1970FAB18B2EDEC6F9BC" country="Oman" name="Wadi Andur" pageId="62" pageNumber="61">Wadi Andur</collectingMunicipality>
,
<collectingMethod id="52644EBE206C1970FC468B09D964F9BC" box="[913,1060,1571,1595]" pageId="62" pageNumber="61">UV detection</collectingMethod>
,
<location id="8EFA6072206C1970FBF88B09D901F9DE" LSID="urn:lsid:plazi:treatment:038C87BF206C1904FEB48BF7D8DEFB63:8EFA6072206C1970FBF88B09D901F9DE" country="Oman" latitude="17.668" longLatPrecision="13" longitude="54.6545" municipality="Wadi Andur" name="rocky boulder wadi, along base of cliff, east edge of wadi" pageId="62" pageNumber="61">rocky boulder wadi, along base of cliff, east edge of wadi</location>
,
<geoCoordinate id="EE11506E206C1970FB988B6BD989F9DE" box="[1103,1225,1601,1625]" degrees="17" direction="north" minutes="40.08" orientation="latitude" pageId="62" pageNumber="61" precision="9" value="17.668">17°40.08'N</geoCoordinate>
<geoCoordinate id="EE11506E206C1970FB058B6BD80BF9DE" box="[1234,1355,1601,1625]" degrees="54" direction="east" minutes="39.27" orientation="longitude" pageId="62" pageNumber="61" precision="9" value="54.6545">54°39.27'E</geoCoordinate>
,
<elevation id="0008D19A206C1970FA8E8B6BDE31F9FF" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="62" pageNumber="61" unit="m" value="405.0">
<quantity id="4CDD9B4C206C1970FA8E8B6BD8DFF9DE" box="[1369,1439,1601,1625]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="62" pageNumber="61" unit="m" value="405.0">405 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B1069206C1970FC5F8B4AD945F9FF" box="[904,1029,1632,1656]" pageId="62" pageNumber="61" value="1995-09-26">
<collectingDate id="EFDFE981206C1970FC5F8B4AD945F9FF" box="[904,1029,1632,1656]" pageId="62" pageNumber="61" value="1995-09-26">26.IX.1995</collectingDate>
</date>
, 22:50 h, leg.
<collectorName id="26D0537F206C1970FB108B4AD86FF9FF" box="[1223,1327,1632,1656]" pageId="62" pageNumber="61">G. Lowe</collectorName>
&amp;
<collectorName id="26D0537F206C1970FAB08B4BDEE8F910" pageId="62" pageNumber="61">M.D. Gallagher</collectorName>
(
<collectionCode id="ED34AE6C206C1970FC148B55D959F910" box="[963,1049,1663,1687]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="62" pageNumber="61">NHMB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4D3CF4206C1970FBEF8B55D814F953" ID-GBIF-Occurrence="3913646510" collectingDate="2011-09-24" collectionCode="FKCP" collectorName="W. Grosser" country="Oman" elevation="380" latitude="17.18615" location="Jabal Samhan" longLatPrecision="1" longitude="54.94285" pageId="62" pageNumber="61" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D23FD20206C1970FBEF8B55D9CFF910" box="[1080,1167,1663,1687]" pageId="62" pageNumber="61" type="adult">1 adult</specimenCount>
or subadult
<specimenCount id="9D23FD20206C1970FAEC8BAAD80FF910" box="[1339,1359,1664,1687]" pageId="62" pageNumber="61" type="female"></specimenCount>
,
<location id="8EFA6072206C1970FABE8B55DED7F932" LSID="urn:lsid:plazi:treatment:038C87BF206C1904FEB48BF7D8DEFB63:8EFA6072206C1970FABE8B55DED7F932" country="Oman" latitude="17.18615" longLatPrecision="1" longitude="54.94285" name="Jabal Samhan" pageId="62" pageNumber="61">Jabal Samhan</location>
,
<locationDeviation id="9A282F8B206C1970FC718BB7D92FF932" box="[934,1135,1693,1717]" location="Jufa" pageId="62" pageNumber="61">
<quantity id="4CDD9B4C206C1970FC718BB7DEAFF932" box="[934,1007,1693,1717]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="62" pageNumber="61" unit="km" value="15.0">15 km</quantity>
W of Jufa
</locationDeviation>
,
<geoCoordinate id="EE11506E206C1970FBA88BB7D84AF932" box="[1151,1290,1693,1717]" degrees="17" direction="north" minutes="11.169" orientation="latitude" pageId="62" pageNumber="61" precision="1" value="17.18615">17°11.169N</geoCoordinate>
<geoCoordinate id="EE11506E206C1970FAC28BB7D8DBF932" box="[1301,1435,1693,1717]" degrees="54" direction="east" minutes="56.571" orientation="longitude" pageId="62" pageNumber="61" precision="1" value="54.94285">54°56.571'E</geoCoordinate>
,
<elevation id="0008D19A206C1970FCE88B96DEFBF953" box="[831,955,1724,1748]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="62" pageNumber="61" unit="m" value="380.0">
<quantity id="4CDD9B4C206C1970FCE88B96DEC2F953" box="[831,898,1724,1748]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="62" pageNumber="61" unit="m" value="380.0">380 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9B1069206C1970FC108B96D903F953" box="[967,1091,1724,1748]" pageId="62" pageNumber="61" value="2011-09-24">
<collectingDate id="EFDFE981206C1970FC108B96D903F953" box="[967,1091,1724,1748]" pageId="62" pageNumber="61" value="2011-09-24">24.IX.2011</collectingDate>
</date>
, leg.
<collectorName id="26D0537F206C1970FBAB8B97D9B5F953" box="[1148,1269,1724,1748]" pageId="62" pageNumber="61">W. Grosser</collectorName>
(
<collectionCode id="ED34AE6C206C1970FAD38B96D80DF953" box="[1284,1357,1724,1748]" pageId="62" pageNumber="61">FKCP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6522206C1971FCE88BD3D94BFF7B" lastPageId="63" lastPageNumber="62" pageId="62" pageNumber="61" type="diagnosis">
<paragraph id="8B9A36A9206C1971FCE88BD3DC34FBF1" blockId="62.[831,1441,1785,1901]" lastBlockId="63.[192,802,198,1142]" lastPageId="63" lastPageNumber="62" pageId="62" pageNumber="61">
DIAGNOSIS (adults). Medium to large
<taxonomicName id="4C254D2A206C1970FB2C8BD0D80CF896" authorityName=": Vachon" authorityYear="1949" box="[1275,1356,1786,1809]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="62" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABB206C1970FB2C8BD0D80CF896" box="[1275,1356,1786,1809]" italics="true" pageId="62" pageNumber="61">Leiurus</emphasis>
</taxonomicName>
,
<quantity id="4CDD9B4C206C1970FA8A8BD3DE28F8B7" metricMagnitude="-2" metricUnit="m" metricValue="7.8" metricValueMax="8.3" metricValueMin="7.3" pageId="62" pageNumber="61" unit="mm" value="78.0" valueMax="83.0" valueMin="73.0">7383 mm</quantity>
in length, carapace L
<quantity id="4CDD9B4C206C1970FBB18A32D9ADF8B7" box="[1126,1261,1816,1840]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="8.9" metricValueMin="8.4" pageId="62" pageNumber="61" unit="mm" value="8.65" valueMax="8.9" valueMin="8.4">8.48.9 mm</quantity>
; base color yellow, carapace darkly pigmented on interocular area and carinae, tergites with weak, variable fuscosity or pale; pace between anterior median carinae with sparse medium or fine granules; area between posterior median carinae with shallow median furrow flanked by lateral arcs of medium or fine granules; medial intercarinal surfaces of tergites IIIII sparsely granular, finely shagreened or smooth; posterior margin of coxa III smooth; metasoma very long, slender, metasoma II L/W
<specimenCount id="9D23FD20206D1971FD1E8C54DF9DFE12" box="[713,733,382,405]" pageId="63" pageNumber="62" type="male"></specimenCount>
1.91 1.95,
<specimenCount id="9D23FD20206D1971FED78CB7DC54FE33" box="[256,276,413,436]" pageId="63" pageNumber="62" type="female"></specimenCount>
1.74, metasoma III L/W
<specimenCount id="9D23FD20206D1971FDE48CB7DF07FE33" box="[563,583,413,436]" pageId="63" pageNumber="62" type="male"></specimenCount>
2.192.30,
<specimenCount id="9D23FD20206D1971FD198CB7DFA2FE33" box="[718,738,413,436]" pageId="63" pageNumber="62" type="female"></specimenCount>
1.92, metasoma IV L/W
<specimenCount id="9D23FD20206D1971FE408C96DCEBFE54" box="[407,427,444,467]" pageId="63" pageNumber="62" type="male"></specimenCount>
2.702.94,
<specimenCount id="9D23FD20206D1971FDE68C96DF05FE54" box="[561,581,444,467]" pageId="63" pageNumber="62" type="female"></specimenCount>
2.30; ventromedian carinae of metasoma II and III with 3646 (
<specimenCount id="9D23FD20206D1971FD4B8CF3DFF1FE76" box="[668,689,473,497]" pageId="63" pageNumber="62" type="male"></specimenCount>
) or 2025 (
<specimenCount id="9D23FD20206D1971FF1D8CD2DD9BFD97" box="[202,219,504,528]" pageId="63" pageNumber="62" type="female"></specimenCount>
) denticles; ventrolateral carinae of metasoma V with small triangular denticles in male, enlarged subtriangular or lobate denticles in female; pedipalps very slender, patella L/W
<specimenCount id="9D23FD20206D1971FE9D8F7FDC1EFDEB" box="[330,350,597,620]" pageId="63" pageNumber="62" type="male"></specimenCount>
4.284.48,
<specimenCount id="9D23FD20206D1971FE368F7FDCB5FDEB" box="[481,501,597,620]" pageId="63" pageNumber="62" type="female"></specimenCount>
3.09; leg III patella L/D
<specimenCount id="9D23FD20206D1971FCDA8F7FDE61FDEB" box="[781,801,597,620]" pageId="63" pageNumber="62" type="female"></specimenCount>
4.354.54,
<specimenCount id="9D23FD20206D1971FEE88F5EDC13FD0C" box="[319,339,628,651]" pageId="63" pageNumber="62" type="male"></specimenCount>
3.23; pectine teeth
<specimenCount id="9D23FD20206D1971FDE88F5EDF13FD0C" box="[575,595,628,651]" pageId="63" pageNumber="62" type="male"></specimenCount>
3637,
<specimenCount id="9D23FD20206D1971FD618F5EDF8AFD0C" box="[694,714,628,651]" pageId="63" pageNumber="62" type="female"></specimenCount>
3334; pectines long, narrow, pectine L/ carapace L
<specimenCount id="9D23FD20206D1971FD128FB8DF99FD2E" box="[709,729,658,681]" pageId="63" pageNumber="62" type="male"></specimenCount>
1.38 1.44,
<specimenCount id="9D23FD20206D1971FF2A8F9BDC51FD4F" box="[253,273,689,712]" pageId="63" pageNumber="62" type="female"></specimenCount>
1.05, mid- pectine sensillar margin L/metasoma I W
<specimenCount id="9D23FD20206D1971FF338FFADDB8FD60" box="[228,248,720,743]" pageId="63" pageNumber="62" type="male"></specimenCount>
<geoCoordinate id="EE11506E206D1971FED58FE5DC7EFD60" box="[258,318,719,743]" degrees="0.198" direction="north" orientation="latitude" pageId="63" pageNumber="62" precision="55" value="0.198">0.198</geoCoordinate>
<geoCoordinate id="EE11506E206D1971FEE98FFADCCAFD60" box="[318,394,719,743]" degrees="0.225" direction="west" orientation="longitude" pageId="63" pageNumber="62" precision="55" value="-0.225">0.225</geoCoordinate>
,
<specimenCount id="9D23FD20206D1971FE4F8FFADCECFD60" box="[408,428,720,743]" pageId="63" pageNumber="62" type="female"></specimenCount>
0.114; basal 13 pectine teeth of males overlap if anterior pectine margins aligned to posterior margins of coxae IV; fulcra mostly with 34 setae; pectine basal piece smooth except for anterior median patches of fine granules; leg III basitarsus with 811 retrosuperior setae; pedipalp chela fixed finger with trichobothrium
<emphasis id="B951EABB206D1971FE708EADDC82FC18" box="[423,450,903,927]" italics="true" pageId="63" pageNumber="62">db</emphasis>
distal to
<emphasis id="B951EABB206D1971FDE18EA2DF13FC18" box="[566,595,904,927]" italics="true" pageId="63" pageNumber="62">est</emphasis>
; sternite VII with area between median carinae smooth or very faintly shagreened anteriorly; sternite carination:
<emphasis id="B951EABB206D1971FD508EEEDF85FC5B" box="[647,709,964,988]" italics="true" pageId="63" pageNumber="62">males</emphasis>
, sternite III with median carinae strong, sternites IVV with lateral carinae strong, median carinae moderate to strong;
<emphasis id="B951EABB206D1971FEC2890ADC26FBBF" box="[277,358,1056,1080]" italics="true" pageId="63" pageNumber="62">females</emphasis>
, sternite III with median carinae obsolete, sternites IVV with lateral carinae weak, median carinae obsolete.
</paragraph>
<paragraph id="8B9A36A9206D1971FF1789B1D94BFF7B" blockId="63.[192,802,1179,1879]" lastBlockId="63.[831,1441,198,252]" pageId="63" pageNumber="62">
COMPARISONS.
<taxonomicName id="4C254D2A206D1971FE5B89B6DCB6FB34" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[396,502,1179,1203]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB206D1971FE5B89B6DCDBFB34" box="[396,411,1180,1203]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FE7D89B1DCB6FB34" box="[426,502,1179,1203]" italics="true" pageId="63" pageNumber="62">heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206D1971FE2989B6DF78FB34" box="[510,568,1180,1203]" pageId="63" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
appears most closely related to
<taxonomicName id="4C254D2A206D1971FEE78991DCEDFB55" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[304,429,1210,1234]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
<emphasis id="B951EABB206D1971FEE78991DC7FFB55" box="[304,319,1211,1234]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FE988990DCEDFB55" box="[335,429,1210,1234]" italics="true" pageId="63" pageNumber="62">haenggii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206D1971FE618990DCB1FB56" box="[438,497,1210,1233]" pageId="63" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
from
<collectingCountry id="F3327639206D1971FDEF8990DFC8FB56" box="[568,648,1210,1233]" name="Yemen" pageId="63" pageNumber="62">Yemen</collectingCountry>
, southwestern
<collectingCountry id="F3327639206D1971FF1789F2DC44FB77" box="[192,260,1240,1264]" name="Oman" pageId="63" pageNumber="62">Oman</collectingCountry>
, and western
<collectingCountry id="F3327639206D1971FE4F89F2DF69FB77" box="[408,553,1240,1264]" name="Saudi Arabia" pageId="63" pageNumber="62">Saudi Arabia</collectingCountry>
. The two species share similar carination and granulation of the carapace, tergites and sternites, with transverse arcs of granules between posterior median carinae on the carapace, and moderately developed basitarsal bristle combs.
<taxonomicName id="4C254D2A206D1971FCDB887EDC5DFA0D" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
<emphasis id="B951EABB206D1971FCDB887EDE5BFAEC" box="[780,795,1364,1387]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FF178858DC5DFA0D" box="[192,285,1394,1418]" italics="true" pageId="63" pageNumber="62">haenggii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206D1971FEF18858DC21FA0E" box="[294,353,1394,1417]" pageId="63" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
differs from
<taxonomicName id="4C254D2A206D1971FE228859DF1FFA0D" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[501,607,1394,1418]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB206D1971FE228859DF44FA0D" box="[501,516,1395,1418]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FDC48858DF1FFA0D" box="[531,607,1394,1418]" italics="true" pageId="63" pageNumber="62">heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206D1971FDB08858DFE3FA0E" box="[615,675,1394,1417]" pageId="63" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
as follows: (i) males with less elongated pedipalps, legs and metasoma IIV; (ii) carinae on metasoma IIV more robust with coarser granulation; (iii) males with larger, lobate dentition on ventrolateral carinae of metasoma V; (iv) dark pigmentation typically extends over posterior area of carapace and most of tergites; (v) lower range of pectinal tooth counts.
<taxonomicName id="4C254D2A206D1971FE7B8B63DC78F9F8" authority="Lourenco, Modry et Amr, 2002" authorityName="Lourenco, Modry et Amr" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="B951EABB206D1971FE7B8B63DCFBF9E7" box="[428,443,1609,1632]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FE1F8B62DF04F9E7" box="[456,580,1608,1632]" italics="true" pageId="63" pageNumber="62">jordanensis</emphasis>
<bibRefCitation id="EFB44B58206D1971FD9C8B63DC78F9F8" author="Lourenco, Modry et Amr" firstAuthor="Lourenco" journalOrPublisher="Revue suisse de Zoologie" pageId="63" pageNumber="62" pagination="635 - 642" part="109" refId="ref62388" refString="LOURENCO, W. R., D. MODRY &amp; Z. AMR. 2002. Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan. Revue suisse de Zoologie, 109 (3): 635 - 642." title="Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan" type="journal article" year="2002">Lourenço, Modrý et Amr, 2002</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F3327639206D1971FE578B4DDC87F9F8" box="[384,455,1639,1663]" name="Jordan" pageId="63" pageNumber="62">Jordan</collectingCountry>
and northern
<collectingCountry id="F3327639206D1971FDB68B4DDFAEF9F8" box="[609,750,1639,1663]" name="Saudi Arabia" pageId="63" pageNumber="62">Saudi Arabia</collectingCountry>
also has long, slender pedipalps and metasomal segments, but can be differentiated from
<taxonomicName id="4C254D2A206D1971FE098B8FDF06F93B" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[478,582,1700,1724]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB206D1971FE098B8FDCADF93B" box="[478,493,1701,1724]" italics="true" pageId="63" pageNumber="62">L</emphasis>
.
<emphasis id="B951EABB206D1971FE2D8B8EDF06F93B" box="[506,582,1700,1724]" italics="true" pageId="63" pageNumber="62">heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0206D1971FD9A8B8FDFC6F93B" box="[589,646,1701,1724]" pageId="63" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
as follows: (i) different color pattern, with dark brown carapace, pedipalps, tergites and metasoma, and contrasting yellow telson and pedipalp fingers; (ii) larger size, adult female carapace length up to
<quantity id="4CDD9B4C206D1971FE648A35DF57F8B0" box="[435,535,1823,1847]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="63" pageNumber="62" unit="mm" value="11.4">11.4 mm</quantity>
; (iii) telson with vesicle elongated, not bulbous; (iv) lower range of pectinal tooth counts:
<specimenCount id="9D23FD20206D1971FC458DECDEE6FF5A" box="[914,934,198,221]" pageId="63" pageNumber="62" type="male"></specimenCount>
3337,
<specimenCount id="9D23FD20206D1971FC298DECD952FF5A" box="[1022,1042,198,221]" pageId="63" pageNumber="62" type="female"></specimenCount>
2731; (v) basitarsi IIII with 1020 retrosuperior setae.
</paragraph>
</subSubSection>
<subSubSection id="C33F6522206D1971FCE88C0BD98DFEF0" pageId="63" pageNumber="62" type="etymology">
<paragraph id="8B9A36A9206D1971FCE88C0BD98DFEF0" blockId="63.[831,1441,289,375]" pageId="63" pageNumber="62">
ETYMOLOGY. A patronym in honor of Blaine Hébert,
<collectingRegion id="49E1F84B206D1971FCE88C6ADEEEFEDF" box="[831,942,320,344]" country="United States of America" name="California" pageId="63" pageNumber="62">California</collectingRegion>
, for his friendship and support of arachnology, science education and music.
</paragraph>
</subSubSection>
<subSubSection id="C33F6522206D1904FCE88C86D963FD60" lastPageId="74" lastPageNumber="73" pageId="63" pageNumber="62" type="description">
<paragraph id="8B9A36A9206D1971FCE88C86D9E3FE43" blockId="63.[830,1441,428,1894]" box="[831,1187,428,452]" pageId="63" pageNumber="62">
DESCRIPTION (
<typeStatus id="549E880B206D1971FC2D8C86D918FE43" box="[1018,1112,428,452]" pageId="63" pageNumber="62" type="holotype">holotype</typeStatus>
male).
</paragraph>
<paragraph id="8B9A36A9206D1971FCB88CE6D9DDFD1C" blockId="63.[830,1441,428,1894]" pageId="63" pageNumber="62">
<emphasis id="B951EABB206D1971FCB88CE6DEB1FE63" bold="true" box="[879,1009,460,484]" pageId="63" pageNumber="62">Coloration.</emphasis>
Base color pale orange-yellow; carapace with dark pigmentation on interocular triangle, and around median ocular tubercle; carinae of carapace and tergites darkened; faint intercarinal fuscosity on posteromedial area of carapace and medial areas of tergites; metasoma V black except for most distal portion; all appendages pale yellow.
</paragraph>
<paragraph id="8B9A36A9206D1971FCB88F88DEE0F9F7" blockId="63.[830,1441,428,1894]" pageId="63" pageNumber="62">
<emphasis id="B951EABB206D1971FCB88F88DEA3FD3D" bold="true" box="[879,995,674,698]" pageId="63" pageNumber="62">Carapace.</emphasis>
Subrectangular, W/L 1.03, with steeply sloped lateral flanks; upper surface with nearly flat posterior and medial plateau areas, well raised ocular tubercle; interocular triangle convex laterally, weakly depressed medially; anterior margin distinctly emarginate, microdenticulate except for lateral ends, bordered by row of large granules; 6 short macrosetae on anterior margin, carapace otherwise devoid of macrosetae; 5 lateral eyes (3 large, 2 small) on each side;
<emphasis id="B951EABB206D1971FAFD8EB2D8D9FC28" box="[1322,1433,920,943]" italics="true" pageId="63" pageNumber="62">carination</emphasis>
: anterior median, superciliary, central lateral, posterior median and posterior lateral carinae strong, coarsely granular; anterior median carinae separated by clear gap from anterior marginal granules; central lateral and posterior median carinae fused into lyre configuration; central median carinae coarsely granular, anterior and posterior portions nearly collinear, not strongly curved, with only slight inward bend on posterior ends; posterior lateral carinae strong, hind end with slight lateral bend, projecting past posterior margin of carapace; lateral ocular carinae weak with medium granules;
<emphasis id="B951EABB206D1971FACC89C2D8D9FA87" box="[1307,1433,1256,1280]" italics="true" pageId="63" pageNumber="62">granulation</emphasis>
: sparse patches of 810 medium or large granules on each anterolateral corner of interocular triangle, 89 small granules on area in front of lateral ocular carinae; surface between anterior median carinae smooth with 8 scattered small granules; lateral flanks almost smooth, with few sparse granules; other intercarinal surfaces smooth with few fine granules; posterior median furrow shallow, almost flat, with median line of several fine granules or microgranules, flanked by lateral arcs of small granules; posterior margin of carapace between posterior lateral carinae bordered by row of medium granules.
</paragraph>
<paragraph id="8B9A36A9206D190CFCB88B52DC05F9C5" blockId="63.[830,1441,428,1894]" lastBlockId="66.[192,802,1394,1909]" lastPageId="66" lastPageNumber="65" pageId="63" pageNumber="62">
<emphasis id="B951EABB206D1971FCB88B52DEA3F917" bold="true" box="[879,995,1656,1680]" pageId="63" pageNumber="62">Chelicera.</emphasis>
Dorsal surface of manus smooth, with 6 short, pale microsetae, 4 near apical margin, 2 subapical, each surrounded by clusters of granules; dorsointernal carina at base of fixed finger moderately strong, terminated anteriorly by large granules projecting over front of manus; single dark macroseta midway along dorsointernal carina; dorsal surface of movable finger smooth, with 4 pale microsetae; fingers with characteristic buthid dentition (
<bibRefCitation id="EFB44B582010190CFE1B8858DF28FA0D" author="Vachon" box="[460,616,1394,1418]" firstAuthor="Vachon" pageId="66" pageNumber="65" pagination="161 - 166" refId="ref64728" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
); movable finger dorsal margin with 5 teeth: dorsal distal tine, subdistal, median and 2 basal teeth fused in bicusp; ventral margin with 3 teeth: ventral distal tine, median and basal teeth; fixed finger margin with 4 teeth: distal tine, subdistal, median and basal teeth; ventral aspect of fixed finger with 2 teeth.
</paragraph>
<caption id="DF5A66212012190EFF178A72D90DF8E8" ID-DOI="http://doi.org/10.5281/zenodo.6505723" ID-Zenodo-Dep="6505723" box="[192,1101,1880,1904]" httpUri="https://zenodo.org/record/6505723/files/figure.png" pageId="64" pageNumber="63" startId="64.[192,267,1880,1904]" targetBox="[333,1297,216,1849]" targetPageId="64">
<paragraph id="8B9A36A92012190EFF178A72D90DF8E8" blockId="64.[192,1101,1880,1904]" box="[192,1101,1880,1904]" pageId="64" pageNumber="63">
<emphasis id="B951EABB2012190EFF178A72DC75F8F7" bold="true" box="[192,309,1880,1904]" pageId="64" pageNumber="63">Figure 47:</emphasis>
<taxonomicName id="4C254D2A2012190EFEEB8A70DC8FF8E8" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[316,463,1881,1903]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2012190EFEEB8A70DC8FF8E8" box="[316,463,1881,1903]" italics="true" pageId="64" pageNumber="63">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02012190EFE018A70DF4AF8E8" box="[470,522,1882,1903]" pageId="64" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Habitus, dorsal aspect. Wadi Andur, Oman.
</paragraph>
</caption>
<caption id="DF5A66212013190FFF178A7CD915F8EA" ID-DOI="http://doi.org/10.5281/zenodo.6505725" ID-Zenodo-Dep="6505725" box="[192,1109,1878,1902]" httpUri="https://zenodo.org/record/6505725/files/figure.png" pageId="65" pageNumber="64" startId="65.[192,267,1878,1902]" targetBox="[321,1285,215,1846]" targetPageId="65">
<paragraph id="8B9A36A92013190FFF178A7CD915F8EA" blockId="65.[192,1109,1878,1902]" box="[192,1109,1878,1902]" pageId="65" pageNumber="64">
<emphasis id="B951EABB2013190FFF178A7CDC75F8E9" bold="true" box="[192,309,1878,1902]" pageId="65" pageNumber="64">Figure 48:</emphasis>
<taxonomicName id="4C254D2A2013190FFEEB8A72DC8FF8EA" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[316,463,1879,1901]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="65" pageNumber="64" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2013190FFEEB8A72DC8FF8EA" box="[316,463,1879,1901]" italics="true" pageId="65" pageNumber="64">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02013190FFE018A72DF4AF8EA" box="[470,522,1880,1901]" pageId="65" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Habitus, ventral aspect. Wadi Andur, Oman.
</paragraph>
</caption>
<caption id="DF5A66212010190CFF178825DC42FAB8" ID-DOI="http://doi.org/10.5281/zenodo.6505727" ID-Zenodo-Dep="6505727" httpUri="https://zenodo.org/record/6505727/files/figure.png" pageId="66" pageNumber="65" startId="66.[192,267,1295,1319]" targetBox="[204,1427,219,1253]" targetPageId="66">
<paragraph id="8B9A36A92010190CFF178825DC42FAB8" blockId="66.[192,1440,1295,1343]" pageId="66" pageNumber="65">
<emphasis id="B951EABB2010190CFF178825DC79FAA0" bold="true" box="[192,313,1295,1319]" pageId="66" pageNumber="65">Figure 49:</emphasis>
<taxonomicName id="4C254D2A2010190CFE95883BDC99FAA1" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[322,473,1296,1318]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="66" pageNumber="65" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2010190CFE95883BDC99FAA1" box="[322,473,1296,1318]" italics="true" pageId="66" pageNumber="65">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02010190CFE35883BDF5AFAA1" box="[482,538,1297,1318]" pageId="66" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
, holotype male.
<emphasis id="B951EABB2010190CFD14883ADF9AFAA2" bold="true" box="[707,730,1296,1317]" pageId="66" pageNumber="65">A.</emphasis>
Carapace and tergites.
<emphasis id="B951EABB2010190CFC10883ADE9EFAA2" bold="true" box="[967,990,1296,1317]" pageId="66" pageNumber="65">B.</emphasis>
Coxosternal area and sternites. Wadi Andur, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A92010190CFF278B63D92CF9C5" blockId="66.[192,802,1394,1909]" lastBlockId="66.[831,1440,1394,1909]" pageId="66" pageNumber="65">
<emphasis id="B951EABB2010190CFF278B63DCFBF9E6" bold="true" box="[240,443,1609,1633]" pageId="66" pageNumber="65">Coxosternal area.</emphasis>
Coxa I smooth except for coarse granules on distal margin, II smooth with sparse fine granules along posterior margin, III smooth, IV smooth to faintly shagreened; coxal endite II with weak, finely granulated carina; coxae IIIII with medium to coarse granulation on anterior carinae, distal margins bearing coarse granules; proximal part of anterior carina of coxa III with fine denticulation; 3 macrosetae along anterior carinae of coxae IIIII; anterior carina of coxa IV with regular coarse granulation, becoming fine proximally, with single proximal macroseta; posterior margin of coxa IV with finely granulated carina on proximal half; sternum smooth, with anterior clusters of fine granules, scattered fine granules, one pair of macrosetae, and deep posteromedian furrow; genital opercula subtriangular with convex margins, sparse microgranulation in lateral areas, 56 short macrosetae.
</paragraph>
<paragraph id="8B9A36A92010190DFCB88B63D991F8BC" blockId="66.[831,1440,1394,1909]" lastBlockId="67.[831,1441,1796,1912]" lastPageId="67" lastPageNumber="66" pageId="66" pageNumber="65">
<emphasis id="B951EABB2010190CFCB88B63DE93F9E6" bold="true" box="[879,979,1609,1633]" pageId="66" pageNumber="65">Pectines.</emphasis>
Basal piece with deep anterior median pit, with patches of fine granules anteromedially, otherwise smooth; 6 short macrosetae along anterior margin, 3 more in center; pectines long, tips extending past proximal 7/8 of trochanter IV; 3 marginal lamellae, small accessory lamella distal to first marginal lamella, 11 middle lamellae, 3737 teeth; marginal lamellae, middle lamellae and fulcra with dense cover of short reddish macrosetae; fulcra with 34 setae; if anterior margins of left and right pectines aligned with posterior edges of coxae IV, only most basal pair of teeth slightly overlap, with wide gap between basal middle lamellae, and teeth contact only anterior half of median carinae of sternite III; pectine teeth of moderate size, mid-pectine sensillar margin L/ pectine L 0.080, mid-pectine sensillar margin L/ metasoma I
<geoCoordinate id="EE11506E2011190DFBBA8A09D98CF8BC" box="[1133,1228,1827,1851]" degrees="0.198" direction="west" orientation="longitude" pageId="67" pageNumber="66" precision="55" value="-0.198">W 0.198</geoCoordinate>
.
</paragraph>
<caption id="DF5A66212011190DFF178BA2DF24F956" ID-DOI="http://doi.org/10.5281/zenodo.6505729" ID-Zenodo-Dep="6505729" httpUri="https://zenodo.org/record/6505729/files/figure.png" pageId="67" pageNumber="66" startId="67.[192,267,1672,1696]" targetBox="[191,1440,170,1641]" targetPageId="67">
<paragraph id="8B9A36A92011190DFF178BA2DF24F956" blockId="67.[192,1441,1672,1745]" pageId="67" pageNumber="66">
<emphasis id="B951EABB2011190DFF178BA2DC76F927" bold="true" box="[192,310,1672,1696]" pageId="67" pageNumber="66">Figure 50:</emphasis>
<taxonomicName id="4C254D2A2011190DFEE98BA0DC93F918" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[318,467,1673,1695]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2011190DFEE98BA0DC93F918" box="[318,467,1673,1695]" italics="true" pageId="67" pageNumber="66">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02011190DFE0D8BA0DF50F918" box="[474,528,1674,1695]" pageId="67" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Right pedipalp.
<emphasis id="B951EABB2011190DFC988BA3DE26F919" bold="true" box="[847,870,1673,1694]" pageId="67" pageNumber="66">A.</emphasis>
Femur, dorsal aspect.
<emphasis id="B951EABB2011190DFB938BA3D91AF919" bold="true" box="[1092,1114,1673,1694]" pageId="67" pageNumber="66">B.</emphasis>
Patella, dorsal aspect.
<emphasis id="B951EABB2011190DFAED8BA3D812F919" bold="true" box="[1338,1362,1673,1694]" pageId="67" pageNumber="66">C.</emphasis>
Patella, external aspect.
<emphasis id="B951EABB2011190DFEB58B88DC3AF930" bold="true" box="[354,378,1698,1719]" pageId="67" pageNumber="66">D.</emphasis>
Chela, ventral aspect.
<emphasis id="B951EABB2011190DFDB58B88DF39F930" bold="true" box="[610,633,1698,1719]" pageId="67" pageNumber="66">E.</emphasis>
Chela, dorsal aspect.
<emphasis id="B951EABB2011190DFC8E8B88DE2EF930" bold="true" box="[857,878,1698,1719]" pageId="67" pageNumber="66">F.</emphasis>
Fixed finger dentition.
<emphasis id="B951EABB2011190DFB888B88D938F930" bold="true" box="[1119,1144,1698,1719]" pageId="67" pageNumber="66">G.</emphasis>
Movable finger dentition.
<emphasis id="B951EABB2011190DFA5F8B88D8E1F930" bold="true" box="[1416,1441,1698,1719]" pageId="67" pageNumber="66">H.</emphasis>
Chela, external aspect. Wadi Andur, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A92011190BFCB88A68D97AF906" blockId="67.[831,1441,1796,1912]" lastBlockId="69.[831,1440,1426,1910]" lastPageId="69" lastPageNumber="68" pageId="67" pageNumber="66">
<emphasis id="B951EABB2011190DFCB88A68DEADF8DD" bold="true" box="[879,1005,1858,1882]" pageId="67" pageNumber="66">Mesosoma.</emphasis>
<emphasis id="B951EABB2011190DFC228A68D90CF8DE" box="[1013,1100,1858,1881]" italics="true" pageId="67" pageNumber="66">Tergites</emphasis>
: pretergites smooth; tergites I II with 5 granular carinae; median and inner lateral carinae linear with medium granules; outer lateral carinae aligned with posterior lateral carinae of carapace, angled outward, strong with large, granules, hind ends extending past posterior margins of tergites without (on I) or with short (on II) lateral extensions; medial intercarinal surfaces smooth with few medium granules, and linear transverse series of fine granules on anterior margin; lateral flanks steeply sloped with sparse microgranulation; tergites IIIVI with 3 straight coarsely granulated carinae; medial intercarinal surfaces mostly smooth with traces of microgranulation; fine granules present in anterior median patch and short transverse strips on either side; lateral surfaces of IIIVI moderately sloped, with numerous medium to coarse granules, some arranged in short longitudinal rows; tergite VII with 5 strong, granular carinae; inner and outer lateral carinae joined anteriorly by transverse series of granules becoming more irregular laterally; medial intercarinal surface smooth, sparse anterior median and anterior lateral patches of microgranules; mediolateral surface smooth with 23 isolated small granules; lateral surfaces smooth; posterior margins of tergites IVI rimmed with linear rows of small to medium sized granules; posterior margin of tergite VII with 5 granules between inner lateral carinae, and 02 small granules laterally;
<emphasis id="B951EABB2016190AFBBC8BC8D986F97E" box="[1131,1222,1762,1785]" italics="true" pageId="68" pageNumber="67">sternites</emphasis>
: sternite III with median carinae strong, thick, densely finely granulated; sternites IVV with thick, densely granulated carinae, lateral carinae strong, median carinae moderate; sternite VI with median carinae strong, with dense fine granulation, lateral carinae strong with dense, fine granulation in anterior half, coarse serial granulation in posterior half; lateral margins of sternites IVVII armed with regular medium to large denticulate granules; medial intercarinal surface of III lightly shagreened anteriorly, heavily on posterior margin, of IV densely shagreened, smooth along posterior margin, of V densely shagreened anteriorly, with broad, smooth posterior margin, of VI densely shagreened anteriorly, nearly smooth towards posterior margin; mediolateral surfaces on IVVI heavily shagreened anteriorly, lightly in posterior area; lateral surfaces of IVVI smooth; sternite VII with 4 strong, narrow, coarsely granular carinae, intercarinal surfaces smooth except for lightly shagreened area anterior to median carinae;
<emphasis id="B951EABB2017190BFD118A6ADE5AF8D0" box="[710,794,1856,1879]" italics="true" pageId="69" pageNumber="68">setation</emphasis>
: sternite III with 4 macrosetae on median carinae, 4 along external margins; sternites IVVII with 2 paired macrosetae on median carinae, one in middle of sternite, other on posterior margin; lateral carinae on IVVI with single posterior marginal macroseta; intercarinal macrosetae: one pair of lateral submarginal setae, one pair of outer mediolateral setae on IVVI, one pair of lateral setae on VII; intercarinal posterior marginal macrosetae on IIIVII: 4, 3, 2, 0, 0.
</paragraph>
<caption id="DF5A66212016190AFF178849D92BFAFC" ID-DOI="http://doi.org/10.5281/zenodo.6505731" ID-Zenodo-Dep="6505731" box="[192,1131,1379,1403]" httpUri="https://zenodo.org/record/6505731/files/figure.png" pageId="68" pageNumber="67" startId="68.[192,267,1379,1403]" targetBox="[191,1437,191,1349]" targetPageId="68">
<paragraph id="8B9A36A92016190AFF178849D92BFAFC" blockId="68.[192,1131,1379,1403]" box="[192,1131,1379,1403]" pageId="68" pageNumber="67">
<emphasis id="B951EABB2016190AFF178849DC76FAFC" bold="true" box="[192,310,1379,1403]" pageId="68" pageNumber="67">Figure 51:</emphasis>
<taxonomicName id="4C254D2A2016190AFEEB884CDC8FFAFC" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[316,463,1381,1403]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="68" pageNumber="67" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2016190AFEEB884CDC8FFAFC" box="[316,463,1381,1403]" italics="true" pageId="68" pageNumber="67">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02016190AFE01884FDF4AFAFD" box="[470,522,1381,1402]" pageId="68" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
, paratype female. Habitus, dorsal aspect. Jabal Samhan, Oman.
</paragraph>
</caption>
<caption id="DF5A66212017190BFF17886DD934FAD8" ID-DOI="http://doi.org/10.5281/zenodo.6505733" ID-Zenodo-Dep="6505733" box="[192,1140,1351,1375]" httpUri="https://zenodo.org/record/6505733/files/figure.png" pageId="69" pageNumber="68" startId="69.[192,267,1351,1375]" targetBox="[191,1440,191,1321]" targetPageId="69">
<paragraph id="8B9A36A92017190BFF17886DD934FAD8" blockId="69.[192,1140,1351,1375]" box="[192,1140,1351,1375]" pageId="69" pageNumber="68">
<emphasis id="B951EABB2017190BFF17886DDC75FAD8" bold="true" box="[192,309,1351,1375]" pageId="69" pageNumber="68">Figure 52:</emphasis>
<taxonomicName id="4C254D2A2017190BFEEB8860DC8FFAD8" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[316,463,1353,1375]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="69" pageNumber="68" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB2017190BFEEB8860DC8FFAD8" box="[316,463,1353,1375]" italics="true" pageId="69" pageNumber="68">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C02017190BFE018860DF4AFAD8" box="[470,522,1354,1375]" pageId="69" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
, paratype female. Habitus, ventral aspect. Jabal Samhan, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A920171908FCB88BA2DCFDFA5D" blockId="69.[831,1440,1426,1910]" lastBlockId="70.[192,802,1378,1897]" lastPageId="70" lastPageNumber="69" pageId="69" pageNumber="68">
<emphasis id="B951EABB2017190BFCB88BA2D921F927" bold="true" box="[879,1121,1672,1696]" pageId="69" pageNumber="68">Hemispermatophore.</emphasis>
Trunk elongate, slender; flagellum long, filiform, pars recta 0.4 times length of trunk, pars reflecta 0.6 times length of trunk; inner lobe a broad lamina, gently tapering with blunt apex; median and outer lobes shorter, partially fused, sharply tapering to thin, flexed apical processes; basal lobe a prominent, digitate hook; measurements (topoparatype): trunk L (to base of flagellum)
<quantity id="4CDD9B4C2017190BFBC38A74D928F8F1" box="[1044,1128,1886,1910]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="69" pageNumber="68" unit="mm" value="7.6">7.6 mm</quantity>
, pars recta
<quantity id="4CDD9B4C2017190BFB308A74D87BF8F1" box="[1255,1339,1886,1910]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="69" pageNumber="68" unit="mm" value="2.9">2.9 mm</quantity>
, pars reflecta
<quantity id="4CDD9B4C20141908FED38848DC1CFAFD" box="[260,348,1378,1402]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="70" pageNumber="69" unit="mm" value="4.5">4.5 mm</quantity>
, inner lobe (from base of flagellum) 837 m, median lobe (from juncture with inner lobe) 694 m, outer lobe (from juncture with median lobe) 300 m, basal lobe 216 m.
</paragraph>
<caption id="DF5A662120141908FF1789D5DC42FAA8" ID-DOI="http://doi.org/10.5281/zenodo.6505735" ID-Zenodo-Dep="6505735" httpUri="https://zenodo.org/record/6505735/files/figure.png" pageId="70" pageNumber="69" startId="70.[192,267,1279,1303]" targetBox="[204,1426,207,1238]" targetPageId="70">
<paragraph id="8B9A36A920141908FF1789D5DC42FAA8" blockId="70.[192,1440,1279,1327]" pageId="70" pageNumber="69">
<emphasis id="B951EABB20141908FF1789D5DC77FA90" bold="true" box="[192,311,1279,1303]" pageId="70" pageNumber="69">Figure 53:</emphasis>
<taxonomicName id="4C254D2A20141908FEE88828DC94FA90" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[319,468,1281,1303]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="70" pageNumber="69" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB20141908FEE88828DC94FA90" box="[319,468,1281,1303]" italics="true" pageId="70" pageNumber="69">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C020141908FE0C8828DF52FA90" box="[475,530,1282,1303]" pageId="70" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
, paratype female.
<emphasis id="B951EABB20141908FD11882BDF9EFA91" bold="true" box="[710,734,1281,1302]" pageId="70" pageNumber="69">A.</emphasis>
Carapace and tergites.
<emphasis id="B951EABB20141908FC13882BDE9AFA91" bold="true" box="[964,986,1281,1302]" pageId="70" pageNumber="69">B.</emphasis>
Coxosternal area and sternites. Jabal Samhan, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A920141909FF2788CBDF48F953" blockId="70.[192,802,1378,1897]" lastBlockId="71.[192,802,1693,1902]" lastPageId="71" lastPageNumber="70" pageId="70" pageNumber="69">
<emphasis id="B951EABB20141908FF2788CBDC2DFA7E" bold="true" box="[240,365,1505,1529]" pageId="70" pageNumber="69">Metasoma.</emphasis>
Very long, slender, total metasoma and telson L/ carapace L 6.1;
<emphasis id="B951EABB20141908FE0C8B2BDF0AF99F" box="[475,586,1537,1560]" italics="true" pageId="70" pageNumber="69">carination</emphasis>
: segment I with 10 complete carinae; segments IIIII with 8 complete carinae, median lateral carinae restricted to posterior 1/4 of II, posterior 1/7 of III; metasoma IV with 8 carinae, V with 7 carinae; carinae on segments IIV finely crenulate-granulate; dorsosubmedian carinae moderately granulate on I, weakly granulate-crenulate on IIIII, with irregular fine granulation on posterior 2/3 of IV; dorsolateral carinae moderately serrate or crenulate on III, weakly granulate-crenulate on IIIIV, weak and finely granular on V; ventrolateral carinae moderate, finely granulate-crenulate on IIV; median lateral carinae moderate, granulate on IIII; ventromedian carinae moderate to strong on IIV, crenulations on IIIII uniform, not enlarged posteriorly; 4243 granules on ventromedian carinae of metasoma IIIII; metasoma V with dorsolateral carinae weak, finely granular; ventrolateral carinae strong, with small and large triangular dentate granules, larger posteriorly, forming scalloped margin; ventrosubmedian carinae on V weak, confined to anterior 2/3 of segment, marked by small and large dentate granules; ventromedian carina strong, with small to large dentate granules, increasing in size posteriorly; lateral anal margin with 3 large rounded lobes; ventral anal margin with 11 irregular, transverse crenulations;
<emphasis id="B951EABB20141908FCE88BD8D95FF88D" box="[831,1055,1778,1802]" italics="true" pageId="70" pageNumber="69">intercarinal surfaces</emphasis>
: smooth on segments IIII, smooth to minutely shagreened on IV, sparsely, finely shagreened on V;
<emphasis id="B951EABB20141908FC318A1AD97AF8C0" box="[998,1082,1840,1863]" italics="true" pageId="70" pageNumber="69">setation</emphasis>
: ventromedian and ventrolateral carinae on segments IIV each bearing 2 exterior macrosetae; metasoma V with 4 macrosetae on lateral surface, 2 on ventrosubmedian carinae.
</paragraph>
<caption id="DF5A662120151909FF178B13D842F9EE" ID-DOI="http://doi.org/10.5281/zenodo.6505737" ID-Zenodo-Dep="6505737" httpUri="https://zenodo.org/record/6505737/files/figure.png" pageId="71" pageNumber="70" startId="71.[192,267,1593,1617]" targetBox="[191,1439,167,1563]" targetPageId="71">
<paragraph id="8B9A36A920151909FF178B13D842F9EE" blockId="71.[192,1441,1593,1641]" pageId="71" pageNumber="70">
<emphasis id="B951EABB20151909FF178B13DC76F9D6" bold="true" box="[192,310,1593,1617]" pageId="71" pageNumber="70">Figure 54:</emphasis>
<taxonomicName id="4C254D2A20151909FEEA8B16DC90F9D6" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[317,464,1595,1617]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="71" pageNumber="70" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB20151909FEEA8B16DC90F9D6" box="[317,464,1595,1617]" italics="true" pageId="71" pageNumber="70">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C020151909FE018B16DF4BF9D6" box="[470,523,1596,1617]" pageId="71" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
, paratype female. Right pedipalp.
<emphasis id="B951EABB20151909FC808B11DE2EF9D7" bold="true" box="[855,878,1595,1616]" pageId="71" pageNumber="70">A.</emphasis>
Femur, dorsal aspect.
<emphasis id="B951EABB20151909FB9F8B11D91EF9D7" bold="true" box="[1096,1118,1595,1616]" pageId="71" pageNumber="70">B.</emphasis>
Patella, dorsal aspect.
<emphasis id="B951EABB20151909FAEC8B11D813F9D7" bold="true" box="[1339,1363,1595,1616]" pageId="71" pageNumber="70">C.</emphasis>
Patella, external aspect.
<emphasis id="B951EABB20151909FE8C8B7EDC33F9EE" bold="true" box="[347,371,1620,1641]" pageId="71" pageNumber="70">D.</emphasis>
Chela, ventral aspect.
<emphasis id="B951EABB20151909FD9A8B7EDF24F9EE" bold="true" box="[589,612,1620,1641]" pageId="71" pageNumber="70">E.</emphasis>
Chela, dorsal aspect.
<emphasis id="B951EABB20151909FCE18B7EDE0BF9EE" bold="true" box="[822,843,1620,1641]" pageId="71" pageNumber="70">F.</emphasis>
Chela, external aspect. Jabal Samhan, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A920151909FF278BF1DFC7F8B7" blockId="71.[192,802,1693,1902]" pageId="71" pageNumber="70">
<emphasis id="B951EABB20151909FF278BF1DC02F974" bold="true" box="[240,322,1755,1779]" pageId="71" pageNumber="70">Telson.</emphasis>
Vesicle smooth, bulbous; ventral surface with scattered fine microsetae and several short macrosetae; aculeus slightly shorter than vesicle.
</paragraph>
<paragraph id="8B9A36A920151907FF278A1DDC76F8B0" blockId="71.[192,802,1693,1902]" lastBlockId="73.[192,802,1024,1909]" lastPageId="73" lastPageNumber="72" pageId="71" pageNumber="70">
<emphasis id="B951EABB20151909FF278A1DDC1CF8C8" bold="true" box="[240,348,1847,1871]" pageId="71" pageNumber="70">Pedipalp.</emphasis>
<emphasis id="B951EABB20151909FEB28A1DDCECF8C9" box="[357,428,1847,1870]" italics="true" pageId="71" pageNumber="70">Femur</emphasis>
: slender, L/
<geoCoordinate id="EE11506E20151909FDE38A1CDF3BF8C9" box="[564,635,1846,1870]" degrees="4.3" direction="west" orientation="longitude" pageId="71" pageNumber="70" precision="5555" value="-4.3">W 4.3</geoCoordinate>
; dorsoexternal, dorsointernal and ventrointernal carinae strong with regular large conical granules; internal carina strong, with irregular large dentate granules; external carina obsolete, a smooth ridge with isolated large dentate granules; dorsal and internal surfaces finely, sparsely shagreened, ventral and external surfaces nearly smooth; linear group of 1517 short macrosetae on distal 1/3 of segment along lower external surface, nearly level with or distal to trichobothrium
<emphasis id="B951EABB201B1907FE50892BDCD3FB9F" box="[391,403,1025,1048]" italics="true" pageId="73" pageNumber="72">e</emphasis>
<subScript id="17A134EC201B1907FE448920DCDCFB9F" attach="left" box="[403,412,1034,1048]" fontSize="6" pageId="73" pageNumber="72">2</subScript>
, part of total of 2021 distal external accessory macrosetae;
<emphasis id="B951EABB201B1907FE388934DF79FBB1" box="[495,569,1054,1078]" italics="true" pageId="73" pageNumber="72">patella</emphasis>
: slender, L/
<geoCoordinate id="EE11506E201B1907FD128934DE5DFBB1" box="[709,797,1054,1078]" degrees="4.48" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-4.48">W 4.48</geoCoordinate>
; dorsointernal carina moderate, with medium to fine granules; dorsomedian carina weak, finely granular; dorsoexternal and external carinae weak, smooth; ventroexternal and ventromedian carinae weak, with fine granules; ventrointernal carina obsolete, marked by noncontiguous coarse conical granules; internal carina moderate, with widely spaced large conical granules separated by fine granulation; both internal and ventrointernal carinae with proximal patellar spur; all patellar surfaces smooth;
<emphasis id="B951EABB201B1907FEF2887BDC1FFAEE" box="[293,351,1361,1385]" italics="true" pageId="73" pageNumber="72">chela</emphasis>
: slender, L/
<geoCoordinate id="EE11506E201B1907FE27887BDF0AFAEE" box="[496,586,1361,1385]" degrees="8.24" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-8.24">W 8.24</geoCoordinate>
, with tenuous upwardly curved fingers, movable finger L/ manus ventral L 2.34; dorsal marginal and ventroexternal carinae faint, smooth, other carinae obsolete; all chela surfaces smooth; manus and fixed finger with sparse short macrosetae; movable finger with numerous short macrosetae on ventral aspect, culminating in dense subapical brush; 12 primary denticle subrows on fixed and movable fingers (left fixed finger regenerated, with 8 subrows); all denticle subrows flanked by internal and external accessory denticles, except proximal subrow of fixed finger without internal accessory denticle;
<emphasis id="B951EABB201B1907FD758B89DC4BF95B" italics="true" pageId="73" pageNumber="72">Trichobothriotaxy</emphasis>
: orthobothriotaxic, type A (
<bibRefCitation id="EFB44B58201B1907FD848BEEDFB1F95B" author="Vachon" box="[595,753,1732,1756]" firstAuthor="Vachon" pageId="73" pageNumber="72" pagination="857 - 958" refId="ref64809" refString="VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
),
<emphasis id="B951EABB201B1907FCD08BEEDE62F95B" box="[775,802,1732,1756]" italics="true" pageId="73" pageNumber="72">db</emphasis>
on fixed finger distal to
<emphasis id="B951EABB201B1907FE028BC9DCB3F97D" box="[469,499,1763,1786]" italics="true" pageId="73" pageNumber="72">est</emphasis>
(
<emphasis id="B951EABB201B1907FDD18BC8DF61F97D" box="[518,545,1762,1786]" italics="true" pageId="73" pageNumber="72">db</emphasis>
missing on right fixed finger,
<emphasis id="B951EABB201B1907FEC58A2BDC65F89F" box="[274,293,1793,1816]" italics="true" pageId="73" pageNumber="72">et</emphasis>
missing on left fixed finger, both present in
<typeStatus id="549E880B201B1907FF178A35DC6BF8B0" box="[192,299,1823,1847]" pageId="73" pageNumber="72" type="paratype">paratypes</typeStatus>
).
</paragraph>
<caption id="DF5A6621201A1906FF178BD4D9C1F8A9" ID-DOI="http://doi.org/10.5281/zenodo.6505739" ID-Zenodo-Dep="6505739" httpUri="https://zenodo.org/record/6505739/files/figure.png" pageId="72" pageNumber="71" startId="72.[192,267,1790,1814]" targetBox="[286,1333,215,1760]" targetPageId="72">
<paragraph id="8B9A36A9201A1906FF178BD4D9C1F8A9" blockId="72.[192,1441,1790,1838]" pageId="72" pageNumber="71">
<emphasis id="B951EABB201A1906FF178BD4DC79F891" bold="true" box="[192,313,1790,1814]" pageId="72" pageNumber="71">Figure 55:</emphasis>
<taxonomicName id="4C254D2A201A1906FE948A2ADC9AF892" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[323,474,1791,1813]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="72" pageNumber="71" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB201A1906FE948A2ADC9AF892" box="[323,474,1791,1813]" italics="true" pageId="72" pageNumber="71">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201A1906FE348A2ADF5BF892" box="[483,539,1792,1813]" pageId="72" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
Metasoma.
<emphasis id="B951EABB201A1906FD408BD5DFEFF893" bold="true" box="[663,687,1791,1812]" pageId="72" pageNumber="71">A.</emphasis>
Male, lateral aspect.
<emphasis id="B951EABB201A1906FC5E8BD5DEE0F893" bold="true" box="[905,928,1791,1812]" pageId="72" pageNumber="71">B.</emphasis>
Male, ventral aspect.
<emphasis id="B951EABB201A1906FB568BD5D9D9F893" bold="true" box="[1153,1177,1791,1812]" pageId="72" pageNumber="71">C.</emphasis>
Female, lateral aspect.
<emphasis id="B951EABB201A1906FA5E8BD5D8E1F893" bold="true" box="[1417,1441,1791,1812]" pageId="72" pageNumber="71">D.</emphasis>
Female, ventral aspect. Male holotype, Wadi Andur, Oman. Female paratype, Jabal Samhan, Oman.
</paragraph>
</caption>
<caption id="DF5A6621201B1907FF178EA9DC42FC4B" ID-DOI="http://doi.org/10.5281/zenodo.6505741" ID-Zenodo-Dep="6505741" httpUri="https://zenodo.org/record/6505741/files/figure.png" pageId="73" pageNumber="72" startId="73.[192,267,899,923]" targetBox="[191,1440,191,869]" targetPageId="73">
<paragraph id="8B9A36A9201B1907FF178EA9DC42FC4B" blockId="73.[192,1441,899,972]" pageId="73" pageNumber="72">
<emphasis id="B951EABB201B1907FF178EA9DC77FC1C" bold="true" box="[192,311,899,923]" pageId="73" pageNumber="72">Figure 56:</emphasis>
<taxonomicName id="4C254D2A201B1907FEE88EACDC94FC1C" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[319,468,901,923]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="73" pageNumber="72" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB201B1907FEE88EACDC94FC1C" box="[319,468,901,923]" italics="true" pageId="73" pageNumber="72">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C0201B1907FE0B8EAFDF53FC1D" box="[476,531,901,922]" pageId="73" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Tarsi.
<emphasis id="B951EABB201B1907FD218EAFDE66FC1D" bold="true" box="[758,806,901,922]" pageId="73" pageNumber="72">A-D.</emphasis>
Left telotarsus and distal basitarsus, ventral aspect.
<emphasis id="B951EABB201B1907FAFF8EAFD800FC1D" bold="true" box="[1320,1344,901,922]" pageId="73" pageNumber="72">A.</emphasis>
Leg I.
<emphasis id="B951EABB201B1907FA5D8EAFD8E1FC1D" bold="true" box="[1418,1441,901,922]" pageId="73" pageNumber="72">B.</emphasis>
Leg II.
<emphasis id="B951EABB201B1907FEDE8EB4DC61FC34" bold="true" box="[265,289,926,947]" pageId="73" pageNumber="72">C.</emphasis>
Leg III.
<emphasis id="B951EABB201B1907FEAE8EB4DCD1FC34" bold="true" box="[377,401,926,947]" pageId="73" pageNumber="72">D.</emphasis>
Leg IV.
<emphasis id="B951EABB201B1907FE3D8EB4DF41FC34" bold="true" box="[490,513,926,947]" pageId="73" pageNumber="72">E.</emphasis>
Right basitarsus III retrolateral aspect. Upper scale bar:
<emphasis id="B951EABB201B1907FBFE8EB4D917FC34" bold="true" box="[1065,1111,926,947]" pageId="73" pageNumber="72">AD</emphasis>
, lower scale bar:
<emphasis id="B951EABB201B1907FAD28EB4D855FC34" bold="true" box="[1285,1301,926,947]" pageId="73" pageNumber="72">E</emphasis>
. Wadi Andur, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A9201B1907FF278A14D81EFAAB" blockId="73.[192,802,1024,1909]" lastBlockId="73.[831,1441,1024,1907]" pageId="73" pageNumber="72">
<emphasis id="B951EABB201B1907FF278A14DC6CF8D1" bold="true" box="[240,300,1854,1878]" pageId="73" pageNumber="72">Legs.</emphasis>
Long, slender, femur III L/ carapace L 1.27, patella III L/D 4.54; inferior carinae strongly denticulate on femur IIV and patella IIII, very weakly denticulate, almost smooth on patella IV; tibia IIIIV with long spurs; retrolateral pedal spurs simple, non-setose; prolateral pedal spurs basally bifurcate, bearing 23 macrosetae; basitarsi IIII with weakly developed bristlecombs, basitarsus III setal counts (left/right): retrosuperior 9/8, retroinferior (including basal accessory seta) 12/13, inferior 14/12; ventral surface of telotarsi with sparse paired rows of stout, finely tapered macrosetae. Right leg I of
<typeStatus id="549E880B201B1907FBC1883ED933FAAB" box="[1046,1139,1300,1324]" pageId="73" pageNumber="72" type="holotype">holotype</typeStatus>
malformed, vestigial.
</paragraph>
<paragraph id="8B9A36A9201B1907FCB88819DE35F91B" blockId="73.[831,1441,1024,1907]" pageId="73" pageNumber="72">
<emphasis id="B951EABB201B1907FCB88819D87DFACC" bold="true" box="[879,1341,1331,1355]" pageId="73" pageNumber="72">
Measurements of
<typeStatus id="549E880B201B1907FB918819D9E8FACC" box="[1094,1192,1331,1355]" pageId="73" pageNumber="72" type="holotype">holotype</typeStatus>
male (mm).
</emphasis>
Total L 83.00; metasoma + telson L 54.00; carapace L 8.79,
<geoCoordinate id="EE11506E201B1907FA518878DE31FA0F" degrees="9.34" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-9.34">W 9.34</geoCoordinate>
, carapace preocular L 4.00; metasomal segments (L/ D /W) I 7.00/ 4.30/ 4.96, II 8.36/ 3.93/ 4.37, III 8.87 / 3.67 / 3.86, IV 9.95 / 3.31 / 3.38, V 10.57 / 3.08 / 3.27; telson L 8.75; vesicle L 4.94, D 3.27,
<geoCoordinate id="EE11506E201B1907FB3488E7D877FA63" box="[1251,1335,1484,1508]" degrees="3.48" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-3.48">W 3.48</geoCoordinate>
; pedipalp chela L 18.21, manus ventral L 5.69, manus
<geoCoordinate id="EE11506E201B1907FA9288C1D8DBF984" box="[1349,1435,1515,1539]" degrees="2.21" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-2.21">W 2.21</geoCoordinate>
, manus D 2.57, fixed finger L 11.36, movable finger L 13.29; pedipalp femur L 9.98,
<geoCoordinate id="EE11506E201B1907FB468B03D9A6F9C7" box="[1169,1254,1576,1600]" degrees="2.32" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-2.32">W 2.32</geoCoordinate>
, patella L 11.12,
<geoCoordinate id="EE11506E201B1907FCE88B6DDED1F9D8" box="[831,913,1607,1631]" degrees="2.48" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-2.48">W 2.48</geoCoordinate>
; pectine L 12.17, mid-pectine sensillar margin L 0.98; leg III femur L 11.18; leg III patella L 9.37, D 2.06.
</paragraph>
<paragraph id="8B9A36A9201B1904FCB88B89DFE1F9CF" blockId="73.[831,1441,1024,1907]" lastBlockId="74.[192,802,1093,1914]" lastPageId="74" lastPageNumber="73" pageId="73" pageNumber="72">
<emphasis id="B951EABB201B1907FCB88B89D841F93C" bold="true" box="[879,1281,1699,1723]" pageId="73" pageNumber="72">
<typeStatus id="549E880B201B1907FCB88B89DE97F93C" box="[879,983,1699,1723]" pageId="73" pageNumber="72" type="paratype">Paratype</typeStatus>
female (Jabal Samhan).
</emphasis>
Smaller than
<typeStatus id="549E880B201B1907FCE88BEBDEDCF95E" box="[831,924,1729,1753]" pageId="73" pageNumber="72" type="holotype">holotype</typeStatus>
male, presumably sub-adult; differs as follows: metasoma, pedipalps and legs not conspicuously elongated: metasoma II L/
<geoCoordinate id="EE11506E201B1907FBE48BD5D9C7F890" box="[1075,1159,1791,1815]" degrees="1.74" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-1.74">W 1.74</geoCoordinate>
, metasoma III L/
<geoCoordinate id="EE11506E201B1907FA9F8BD5D8DCF890" box="[1352,1436,1791,1815]" degrees="1.92" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-1.92">W 1.92</geoCoordinate>
, metasoma IV L/
<geoCoordinate id="EE11506E201B1907FBD08A37D926F8B2" box="[1031,1126,1821,1845]" degrees="2.30" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-2.3">W 2.30</geoCoordinate>
, metasoma V L/
<geoCoordinate id="EE11506E201B1907FAEA8A37D8DBF8B2" box="[1341,1435,1821,1845]" degrees="2.47" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-2.47">W 2.47</geoCoordinate>
, pedipalp femur L/
<geoCoordinate id="EE11506E201B1907FBDD8A16D921F8D3" box="[1034,1121,1852,1876]" degrees="3.25" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-3.25">W 3.25</geoCoordinate>
, pedipalp patella L/
<geoCoordinate id="EE11506E201B1907FA928A16D8DBF8D3" box="[1349,1435,1852,1876]" degrees="3.09" direction="west" orientation="longitude" pageId="73" pageNumber="72" precision="555" value="-3.09">W 3.09</geoCoordinate>
, pedipalp chela L/ carapace L 1.72, pedipalp chela L/ manus
<geoCoordinate id="EE11506E20181904FEC5896CDC28FBDA" box="[274,360,1093,1117]" degrees="6.12" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-6.12">W 6.12</geoCoordinate>
, leg III patella L/D 3.23; wider mesosoma; pectines shorter, tips extending to base of trochanter IV, teeth smaller with shorter sensillar margin,
<quantity id="4CDD9B4C20181904FF38898BDC0EFB3E" box="[239,334,1185,1209]" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="8.636" metricValueMin="8.382" pageId="74" pageNumber="73" unit="in" value="33.5" valueMax="34.0" valueMin="33.0">3334 in</quantity>
number; basal pectine teeth not overlapped when anterior pectinal margins aligned with posterior margins of coxae IV; median carinae obsolete on sternites IIIV, weak on VI; lateral carinae moderate, smooth on sternites IVV, moderate, finely granular on VI; sternite VII with finer median and lateral carinae; sternites with smooth intercarinal surfaces; coarser granules and crenulation on ventromedian and ventrolateral carinae on metasoma IIIII, with more enlarged denticles on posterior half of segments; metasoma IIIII with 2025 granules on ventromedian carinae; metasoma V with larger denticles on ventrolateral carinae, posterior denticles enlarged, triangular; pedipalp femur with 27 distal external accessory macrosetae.
</paragraph>
<caption id="DF5A662120181904FF178E9ADE50FB96" ID-DOI="http://doi.org/10.5281/zenodo.6505743" ID-Zenodo-Dep="6505743" httpUri="https://zenodo.org/record/6505743/files/figure.png" pageId="74" pageNumber="73" startId="74.[192,267,944,968]" targetBox="[223,766,203,893]" targetPageId="74">
<paragraph id="8B9A36A920181904FF178E9ADE50FB96" blockId="74.[192,802,944,1041]" pageId="74" pageNumber="73">
<emphasis id="B951EABB20181904FF178E9ADC07FC4F" bold="true" box="[192,327,944,968]" pageId="74" pageNumber="73">Figure 57:</emphasis>
<taxonomicName id="4C254D2A20181904FEB78E98DF46FC40" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[352,518,945,967]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
<emphasis id="B951EABB20181904FEB78E98DF46FC40" box="[352,518,945,967]" italics="true" pageId="74" pageNumber="73">Leiurus heberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C020181904FDC98E98DF25FC40" box="[542,613,946,967]" pageId="74" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Trichobothrial map of pedipalp.
<emphasis id="B951EABB20181904FDDD8EE0DF61FC58" bold="true" box="[522,545,970,991]" pageId="74" pageNumber="73">A.</emphasis>
Femur, dorsal aspect.
<emphasis id="B951EABB20181904FCDB8EE0DE62FC58" bold="true" box="[780,802,970,991]" pageId="74" pageNumber="73">B.</emphasis>
Patella, dorsal aspect.
<emphasis id="B951EABB20181904FE718EC9DCFDFC7F" bold="true" box="[422,445,995,1016]" pageId="74" pageNumber="73">C.</emphasis>
Patella, external aspect.
<emphasis id="B951EABB20181904FD168EC9DF98FC7F" bold="true" box="[705,728,995,1016]" pageId="74" pageNumber="73">D.</emphasis>
Chela, external aspect.
<emphasis id="B951EABB20181904FE8C8ED6DC32FB96" bold="true" box="[347,370,1020,1041]" pageId="74" pageNumber="73">E.</emphasis>
Chela, ventral aspect. Wadi Andur, Oman.
</paragraph>
</caption>
<paragraph id="8B9A36A920181904FF278B65D814FEF0" blockId="74.[192,802,1093,1914]" lastBlockId="74.[831,1441,198,743]" pageId="74" pageNumber="73">
<emphasis id="B951EABB20181904FF278B65DF89F9E0" bold="true" box="[240,713,1615,1639]" pageId="74" pageNumber="73">Measurements of sub-adult female (mm).</emphasis>
Total L 73.00; metasoma + telson L 48.00; carapace L 8.43,
<geoCoordinate id="EE11506E20181904FCDF8B44DDB1F923" degrees="9.55" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-9.55">W 9.55</geoCoordinate>
, carapace preocular L 4.10; metasomal segments (L/ D/ W) I 5.76/ 4.11/ 4.77, II 7.01/ 3.87/ 4.03, III 7.33/ 3.86/ 3.82, IV 8.15/ 3.54/ 3.54, V 9.28/ 3.12/ 3.76; telson L 8.70; vesicle L 4.38, D 3.12,
<geoCoordinate id="EE11506E20181904FDC68BC3DF23F887" box="[529,611,1768,1792]" degrees="3.30" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-3.3">W 3.30</geoCoordinate>
; pedipalp chela L 14.50, manus ventral L 4.31, manus
<geoCoordinate id="EE11506E20181904FD8C8A2DDFEFF898" box="[603,687,1799,1823]" degrees="2.37" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-2.37">W 2.37</geoCoordinate>
, manus D 2.72, fixed finger L 9.08, movable finger L 10.40; pedipalp femur L 7.96,
<geoCoordinate id="EE11506E20181904FE148A6EDF56F8DB" box="[451,534,1860,1884]" degrees="2.45" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-2.45">W 2.45</geoCoordinate>
, patella L 9.04,
<geoCoordinate id="EE11506E20181904FD1D8A6EDE5EF8DB" box="[714,798,1860,1884]" degrees="2.93" direction="west" orientation="longitude" pageId="74" pageNumber="73" precision="555" value="-2.93">W 2.93</geoCoordinate>
; pectine L 8.86; mid-pectine sensillar margin L 0.54; leg III femur L 8.88; leg III patella L 7.11, D 2.20.
<emphasis id="B951EABB20181904FCE88DCFDEE2FF7B" box="[831,930,229,252]" italics="true" pageId="74" pageNumber="73">Meristics</emphasis>
: pedipalp movable fingers with 12/ 12 (left/ right) subrows of primary denticles, fixed fingers with 11/ 8 subrows (anomalous fusion of rows on right finger); basitarsus III seta counts (left/ right): retrosuperior 15/11, retroinferior 14/13, inferior 13/14.
</paragraph>
<paragraph id="8B9A36A920181904FCB88C54D963FD60" blockId="74.[831,1441,198,743]" pageId="74" pageNumber="73">
<emphasis id="B951EABB20181904FCB88C54DEA3FE11" bold="true" box="[879,995,382,406]" pageId="74" pageNumber="73">Variation.</emphasis>
Comparison of the
<typeStatus id="549E880B20181904FB348C54D800FE11" box="[1251,1344,382,406]" pageId="74" pageNumber="73" type="holotype">holotype</typeStatus>
to two smaller
<typeStatus id="549E880B20181904FC4E8CB7DEB5FE32" box="[921,1013,413,437]" pageId="74" pageNumber="73" type="paratype">paratype</typeStatus>
males showed an increasing slenderness of pedipalps, legs and metasoma with increasing size (
<tableCitation id="C6A7031220181904FCAE8CF3DE84FE76" box="[889,964,473,497]" captionStart="Table 2" captionStartId="75.[192,257,682,706]" captionTargetBox="[319,1283,200,646]" captionText="Table 2: Variation in male morphometric ratios with increasing size in Leiurus heberti sp. n." httpUri="http://table.plazi.org/id/DF5A662120191905FF178F80D909FD46" pageId="74" pageNumber="73" tableUuid="DF5A662120191905FF178F80D909FD46">Tab. 2</tableCitation>
). The smaller of the
<specimenCount id="9D23FD2020181904FB788CF0D8C4FE76" box="[1199,1412,473,498]" pageId="74" pageNumber="73" type="male" typeStatus="paratype">two paratype males</specimenCount>
is more similar to
<taxonomicName id="4C254D2A20181904FC3A8CD3D928FD97" authority="Lowe &amp; Yağmur &amp; Kovařík, 2014" authorityName="Lowe &amp; Yağmur &amp; Kovařík" authorityYear="2014" box="[1005,1128,504,528]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="74" pageNumber="73" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
<emphasis id="B951EABB20181904FC3A8CD3DEBCFD97" box="[1005,1020,505,528]" italics="true" pageId="74" pageNumber="73">L</emphasis>
.
<emphasis id="B951EABB20181904FBDC8CD2D928FD97" box="[1035,1128,504,528]" italics="true" pageId="74" pageNumber="73">haenggii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C020181904FBA68CD3D9EBFD97" box="[1137,1195,505,528]" pageId="74" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
in morphometrics and the presence of 4 enlarged denticles on ventrolateral carinae of metasoma V. Median carinae on sternites III V are weak or obsolete on the smaller
<typeStatus id="549E880B20181904FB298F7FD819FDEA" box="[1278,1369,597,621]" pageId="74" pageNumber="73" type="paratype">paratype</typeStatus>
male. The larger
<typeStatus id="549E880B20181904FC6F8F5ED953FD0B" box="[952,1043,628,652]" pageId="74" pageNumber="73" type="paratype">paratype</typeStatus>
male, like the
<typeStatus id="549E880B20181904FB6D8F59D85AFD0C" box="[1210,1306,627,651]" pageId="74" pageNumber="73" type="holotype">holotype</typeStatus>
, has weakly developed bristle combs on basitarsi IIII, with seta counts (left/right): retrosuperior 9/10, retroinferior 12/12, inferior 13/13.
</paragraph>
</subSubSection>
<subSubSection id="C33F652220181904FCE88E2CD997FCBA" pageId="74" pageNumber="73" type="distribution">
<paragraph id="8B9A36A920181904FCE88E2CD997FCBA" blockId="74.[831,1440,774,829]" pageId="74" pageNumber="73">
DISTRIBUTION. Known only from the Jabal Samhan mountains of
<collectingRegion id="49E1F84B20181904FC068E0FD9C4FCBA" box="[977,1156,805,829]" country="Oman" name="Zufar" pageId="74" pageNumber="73">Dhofar Province</collectingRegion>
,
<collectingCountry id="F332763920181904FB588E0FD993FCBA" box="[1167,1235,805,829]" name="Oman" pageId="74" pageNumber="73">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F652220181904FCE88E76D8DEFB63" pageId="74" pageNumber="73" type="biology_ecology">
<paragraph id="8B9A36A920181904FCE88E76D8DEFB63" blockId="74.[831,1440,860,1252]" pageId="74" pageNumber="73">
ECOLOGY. The male types were collected by ultraviolet detection among rocks and boulders along the edge of a lightly vegetated wadi on the northern side of the Jabal Samhan mountains. All were very agile and aggressive when captured. The only known female
<typeStatus id="549E880B20181904FCE88EDCDEDAFB89" box="[831,922,1014,1038]" pageId="74" pageNumber="73" type="paratype">paratype</typeStatus>
was collected near the base of the Jabal Samhan escarpment facing the southern coastal plain. Both sites are at lower elevations (ca.
<quantity id="4CDD9B4C20181904FB368919D866FBCC" box="[1249,1318,1075,1099]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="74" pageNumber="73" unit="m" value="400.0">400 m</quantity>
a.s.l.), and it is not known if this species occurs at higher elevations in these mountains whose peaks reach over
<quantity id="4CDD9B4C20181904FA96895AD8DFFB0F" box="[1345,1439,1136,1160]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="74" pageNumber="73" unit="m" value="2000.0">2,000 m</quantity>
a.s.l. The compressed body, elongated legs, pedipalps and metasoma, and weakly developed basitarsal bristle combs are consistent with a lithophilous ecomorphotype.
</paragraph>
</subSubSection>
</treatment>
</document>