568 lines
71 KiB
XML
568 lines
71 KiB
XML
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<mods:title id="6978A81A6ADB56F6DA58E538CCE35689">A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula</mods:title>
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<mods:title id="1E1DF59290E211F7501EA290E1DDDB6B">Euscorpius</mods:title>
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Genus
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Ehrenberg, 1828
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<emphasis id="B951EABB2056194AFCE8892BDE88FB9E" box="[831,968,1025,1049]" italics="true" pageId="4" pageNumber="3">Androctonus</emphasis>
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(
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) Ehrenberg
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in
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<bibRefCitation id="EFB44B582056194AFB1F892BD963FBBF" author="Hemprich & Ehrenberg" firstAuthor="Hemprich" journalOrPublisher="Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus" pageId="4" pageNumber="3" refId="ref60936" refString="HEMPRICH, F. W. & C. G. EHRENBERG. 1828. Zoologica II. Arachnoidea. In: Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg. Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus." title="Zoologica II. Arachnoidea" type="book chapter" volumeTitle="Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg" year="1828">Hemprich & Ehrenberg, 1828</bibRefCitation>
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</treatmentCitation>
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, pl.
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, fig. 5;
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<treatmentCitation id="0A8410B82056194AFB7B890AD916FBD1" author="HEMPRICH, F. W. & C. G. EHRENBERG" page="353" pageId="4" pageNumber="3" year="1829">
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<bibRefCitation id="EFB44B582056194AFB7B890AD916FBD1" author="Hemprich & Ehrenberg" firstAuthor="Hemprich" journalOrPublisher="Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin" page="353" pageId="4" pageNumber="3" pagination="348 - 362" part="1" refId="ref61033" refString="HEMPRICH, F. W. & C. G. EHRENBERG. 1829. Vorlaufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Skorpione und deren geographischen Verbreitung. Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin, 1 (6): 348 - 362." title="Vorlaufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Skorpione und deren geographischen Verbreitung" type="journal article" year="1829">Hemprich & Ehrenberg, 1829: 353</bibRefCitation>
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(in part).
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</paragraph>
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<paragraph id="8B9A36A92056194AFCE88956D9CCFB35" blockId="4.[831,1441,1025,1693]" pageId="4" pageNumber="3">
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<emphasis id="B951EABB2056194AFCE88956DE88FB13" box="[831,968,1148,1172]" italics="true" pageId="4" pageNumber="3">Androctonus</emphasis>
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(
|
||
<emphasis id="B951EABB2056194AFC0E8956D95EFB14" box="[985,1054,1148,1171]" italics="true" pageId="4" pageNumber="3">Liurus</emphasis>
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)
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||
</taxonomicName>
|
||
:
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<bibRefCitation id="EFB44B582056194AFBE08956D83AFB13" author="Hemprich & Ehrenberg" bookContentInfo="12 pp." box="[1079,1402,1148,1172]" firstAuthor="Hemprich" journalOrPublisher="Berolini ex Officina Academica, Venditur a Mittlero" pageId="4" pageNumber="3" refId="ref61080" refString="HEMPRICH, F. W. & C. G. EHRENBERG. 1831. Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici. In: Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162. Berolini ex Officina Academica, Venditur a Mittlero, 12 pp. (Latin text, pages unnumbered)." title="Animalia articulata, Arachnoidea. Scorpiones Africani et Asiatici" type="book" volumeTitle="Symbolae Physicae. Animalia Evertebrata Exclusis Insectis Percensuit Dr. C. G. Ehrenberg. Series Prima cum Tabularum Decade Prima. Continet Animalia Africana et Asiatica 162" year="1831">Hemprich & Ehrenberg, 1831</bibRefCitation>
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</treatmentCitation>
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(in part);
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||
<treatmentCitation id="0A8410B82056194AFC7889B0D9CCFB35" author="SUNDEVALL, C. J." box="[943,1164,1178,1202]" page="33" pageId="4" pageNumber="3" year="1833">
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<bibRefCitation id="EFB44B582056194AFC7889B0D9C8FB35" author="Sundevall" box="[943,1160,1178,1202]" firstAuthor="Sundevall" page="33" pageId="4" pageNumber="3" refId="ref64400" refString="SUNDEVALL, C. J. 1833. Conspectus Arachnidum. C. F. Berling, Londini Gothorum, 39 pp." type="book" year="1833">Sundevall, 1833: 33</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8B9A36A92056194AFCE88990DEA1F9E7" blockId="4.[831,1441,1025,1693]" pageId="4" pageNumber="3">
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<taxonomicName id="4C254D2A2056194AFCE88990D9DCFB56" ID-CoL="5C2F" authority=": Vachon, 1949: 83 - 88" authorityName=": Vachon" authorityPageNumber="83 - 88" authorityYear="1949" box="[831,1180,1209,1233]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B951EABB2056194AFCE88990DED0FB56" box="[831,912,1210,1233]" italics="true" pageId="4" pageNumber="3">Leiurus</emphasis>
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:
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<treatmentCitation id="0A8410B82056194AFC728993D9DCFB56" author="VACHON, M." box="[933,1180,1209,1233]" page="83-88" pageId="4" pageNumber="3" year="1949">
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<bibRefCitation id="EFB44B582056194AFC728993D9DCFB56" author="Vachon" box="[933,1180,1209,1233]" firstAuthor="Vachon" page="83 - 88" pageId="4" pageNumber="3" pagination="66 - 100" refId="ref64580" refString="VACHON, M. 1949. Etudes sur les Scorpions (suite). Chapitre III (suite). Description des scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 27 (1): 66 - 100, figs. 242 - 292." type="journal article" year="1949">Vachon, 1949: 83–88</bibRefCitation>
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</treatmentCitation>
|
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</taxonomicName>
|
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;
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<treatmentCitation id="0A8410B82056194AFB798993DE93FB77" pageId="4" pageNumber="3">
|
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<bibRefCitation id="EFB44B582056194AFB798993DEDAFB77" author="Vachon" firstAuthor="Vachon" page="199 - 200" pageId="4" pageNumber="3" pagination="152 - 216" refId="ref64629" refString="VACHON, M. 1950 a. Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28 (2): 152 - 216." type="journal article" year="1950" yearSuffix="a">Vachon, 1950a: 199– 200</bibRefCitation>
|
||
, 202
|
||
</treatmentCitation>
|
||
, figs. 550–552;
|
||
<treatmentCitation id="0A8410B82056194AFB5D89F2D82CFB77" author="VACHON, M." box="[1162,1388,1240,1264]" page="393" pageId="4" pageNumber="3" year="1950">
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<bibRefCitation id="EFB44B582056194AFB5D89F2D82CFB77" author="Vachon" box="[1162,1388,1240,1264]" firstAuthor="Vachon" page="393" pageId="4" pageNumber="3" pagination="382 - 413" refId="ref64669" refString="VACHON, M. 1950 b. Etudes sur les Scorpions. (suite). Determination des Scorpions du Nord-ouest de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28 (3): 382 - 413." type="journal article" year="1950" yearSuffix="b">Vachon, 1950b: 393</bibRefCitation>
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||
</treatmentCitation>
|
||
, fig. 589;
|
||
<treatmentCitation id="0A8410B82056194AFC7D89DCD9F7FA89" author="VACHON, M." box="[938,1207,1270,1294]" page="203-208" pageId="4" pageNumber="3" year="1952">
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<bibRefCitation id="EFB44B582056194AFC7D89DCD9F7FA89" author="Vachon" box="[938,1207,1270,1294]" firstAuthor="Vachon" page="203 - 208" pageId="4" pageNumber="3" refId="ref64708" refString="VACHON, M. 1952. Etude sur les Scorpions. Institut Pasteur d'Algerie, Alger, 482 pp." type="book" year="1952">Vachon, 1952: 203–208</bibRefCitation>
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</treatmentCitation>
|
||
, 372–374, 399, figs. 550–552;
|
||
<treatmentCitation id="0A8410B82056194AFC37883FD9EDFAAA" author="BUCHERL, W." box="[992,1197,1301,1325]" page="57" pageId="4" pageNumber="3" year="1964">
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<bibRefCitation id="EFB44B582056194AFC37883FD9EDFAAA" author="Bucherl" box="[992,1197,1301,1325]" firstAuthor="Bucherl" journalOrPublisher="Memorias do Instituto de Butantan" page="57" pageId="4" pageNumber="3" pagination="55 - 66" part="31" refId="ref59810" refString="BUCHERL, W. 1964. Distribucao geografica dos Aracnoides peconhentos temiveis. Memorias do Instituto de Butantan, 31: 55 - 66." title="Distribucao geografica dos Aracnoides peconhentos temiveis" type="journal article" year="1964">Bücherl, 1964: 57</bibRefCitation>
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</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFB68883FD8DCFAAA" author="STAHNKE, H. L." box="[1215,1436,1301,1325]" page="130" pageId="4" pageNumber="3" year="1972">
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<bibRefCitation id="EFB44B582056194AFB68883FD8DCFAAA" author="Stahnke" box="[1215,1436,1301,1325]" firstAuthor="Stahnke" page="130" pageId="4" pageNumber="3" pagination="121 - 133" refId="ref64369" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." type="journal article" year="1972">Stahnke, 1972: 130</bibRefCitation>
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</treatmentCitation>
|
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;
|
||
<treatmentCitation id="0A8410B82056194AFCB8881ED99DFACB" author="LAMORAL, B. H. & S. C. REYNDERS" box="[879,1245,1332,1356]" page="509" pageId="4" pageNumber="3" year="1975">
|
||
<bibRefCitation id="EFB44B582056194AFCB8881ED99DFACB" author="Lamoral & Reynders" box="[879,1245,1332,1356]" firstAuthor="Lamoral" journalOrPublisher="Annals of the Natal Museum" page="509" pageId="4" pageNumber="3" pagination="489 - 576" part="22" refId="ref62059" refString="LAMORAL, B. H. & S. C. REYNDERS. 1975. A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973. Annals of the Natal Museum, 22 (2): 489 - 576." title="A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973" type="journal article" year="1975">Lamoral & Reynders, 1975: 509</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFB3A881ED96BFAED" author="LEVY, G. & P. AMITAI" page="14, 46-47" pageId="4" pageNumber="3" year="1980">
|
||
<bibRefCitation id="EFB44B582056194AFB3A881EDE93FAED" author="Levy & Amitai" bookContentInfo="130 pp" firstAuthor="Levy" journalOrPublisher="The Israel Academy of Sciences and Humanities. Jerusalem" page="14" pageId="4" pageNumber="3" refId="ref62185" refString="LEVY, G. & P. AMITAI. 1980. Scorpiones. Fauna Palaestina. Arachnida I. The Israel Academy of Sciences and Humanities. Jerusalem, 130 pp .." title="Scorpiones. Fauna Palaestina. Arachnida I" type="book" year="1980">Levy & Amitai, 1980: 14</bibRefCitation>
|
||
, 46–47;
|
||
</treatmentCitation>
|
||
<treatmentCitation id="0A8410B82056194AFBE18878D85FFAED" author="FRANCKE, O. F." box="[1078,1311,1362,1386]" page="9, 15" pageId="4" pageNumber="3" year="1985">
|
||
<bibRefCitation id="EFB44B582056194AFBE18878D9B2FAED" author="Francke" box="[1078,1266,1362,1386]" firstAuthor="Francke" journalOrPublisher="Occasional Papers of the Museum, Texas Tech University" page="9" pageId="4" pageNumber="3" pagination="1 - 32" part="98" refId="ref60564" refString="FRANCKE, O. F. 1985. Conspectus genericus scorpionorum 1758 - 1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University, 98: 1 - 32." title="Conspectus genericus scorpionorum 1758 - 1982 (Arachnida: Scorpiones)" type="journal article" year="1985">Francke, 1985: 9</bibRefCitation>
|
||
, 15
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFAF98878D920FA0E" author="VACHON, M. & R. KINZELBACH." page="92" pageId="4" pageNumber="3" year="1987">
|
||
<bibRefCitation id="EFB44B582056194AFAF98878D920FA0E" author="Vachon & Kinzelbach" editor="KRUPP, F. & W. SCHNEIDER & R. KINZELBACH" firstAuthor="Vachon" journalOrPublisher="Beihefte zum Tubinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften)" page="92" pageId="4" pageNumber="3" pagination="91 - 103" part="28" refId="ref65029" refString="VACHON, M. & R. KINZELBACH. 1987. On the taxonomy and distribution of the scorpions of the Middle East. In KRUPP, F., W. SCHNEIDER & R. KINZELBACH (eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tubinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften), 28: 91 - 103." title="On the taxonomy and distribution of the scorpions of the Middle East" type="book" volumeTitle="Proceedings of the Sym-" year="1987">Vachon & Kinzelbach, 1987: 92</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFBB8885BD800FA0E" author="SISSOM, W. D." box="[1135,1344,1393,1417]" page="101" pageId="4" pageNumber="3" year="1990">
|
||
<bibRefCitation id="EFB44B582056194AFBB8885BD800FA0E" author="Sissom" box="[1135,1344,1393,1417]" editor="POLIS, G. A." firstAuthor="Sissom" journalOrPublisher="The Biology of Scorpions. Stanford University Press, Stanford, CA" page="101" pageId="4" pageNumber="3" pagination="64 - 160" refId="ref64015" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in: POLIS, G. A. (ed.) The Biology of Scorpions. Stanford University Press, Stanford, CA." title="Systematics, biogeography and paleontology" type="book chapter" year="1990">Sissom, 1990: 101</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFA87885BD960FA2F" author="NENILIN, A. B. & V. FET." page="17" pageId="4" pageNumber="3" year="1992">
|
||
<bibRefCitation id="EFB44B582056194AFA87885BD960FA2F" author="Nenilin & Fet" firstAuthor="Nenilin" journalOrPublisher="Arthropoda Selecta" page="17" pageId="4" pageNumber="3" pagination="3 - 31" part="1" refId="ref62944" refString="NENILIN, A. B. & V. FET. 1992. Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones). Arthropoda Selecta, 1 (2): 3 - 31 (in Russian)." title="Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones)" type="journal article" year="1992">Nenilin & Fet, 1992: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFBF888BAD847FA2F" author="KOVARIK, F." box="[1071,1287,1424,1448]" page="112" pageId="4" pageNumber="3" year="1998">
|
||
<bibRefCitation id="EFB44B582056194AFBF888BAD847FA2F" author="Kovarik" bookContentInfo="175 pp." box="[1071,1287,1424,1448]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar, Jihlava" page="112" pageId="4" pageNumber="3" refId="ref61762" refString="KOVARIK, F. 1998. Stiri. Madagaskar, Jihlava, 175 pp." title="Stiri" type="book" year="1998">Kovařík, 1998: 112</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFAC388BADEA2FA41" author="FET, V. & G. LOWE" page="155" pageId="4" pageNumber="3" year="2000">
|
||
<bibRefCitation id="EFB44B582056194AFAC388BADEA2FA41" author="Fet & Lowe" editor="FET, V. & W. D. SISSOM & G. LOWE & M. E. BRAUNWALDER" firstAuthor="Fet" journalOrPublisher="The New York Entomological Society" page="155" pageId="4" pageNumber="3" pagination="54 - 286" refId="ref60347" refString="FET, V. & G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE & M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." title="Family Buthidae" type="book chapter" volumeTitle="Catalog of the Scorpions of the World (1758 - 1998)" year="2000">Fet & Lowe, 2000: 155</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFC258884D809FA41" author="SOLEGLAD, M. E. & V. F. FET." box="[1010,1353,1454,1478]" page="26-27" pageId="4" pageNumber="3" year="2003">
|
||
<bibRefCitation id="EFB44B582056194AFC258884D809FA41" author="Soleglad & Fet" box="[1010,1353,1454,1478]" firstAuthor="Soleglad" journalOrPublisher="Euscorpius" page="26 - 27" pageId="4" pageNumber="3" pagination="1 - 34" part="5" refId="ref64217" refString="SOLEGLAD, M. E. & V. F. FET. 2003 a. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1 - 34." title="The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni)" type="journal article" year="2003" yearSuffix="a">Soleglad & Fet, 2003a: 26–27</bibRefCitation>
|
||
</treatmentCitation>
|
||
, tab. 2, fig. 44;
|
||
<treatmentCitation id="0A8410B82056194AFC1388E7D9A3FA62" author="SOLEGLAD, M. E. & V. F. FET." box="[964,1251,1485,1509]" page="88" pageId="4" pageNumber="3" year="2003">
|
||
<bibRefCitation id="EFB44B582056194AFC1388E7D9A3FA62" author="Soleglad & Fet" box="[964,1251,1485,1509]" firstAuthor="Soleglad" journalOrPublisher="Euscorpius" page="88" pageId="4" pageNumber="3" pagination="1 - 175" part="11" refId="ref64254" refString="SOLEGLAD, M. E. & V. F. FET. 2003 b. High-Level Systematics and Phylogeny of the Extant Scorpions (Scorpiones: Orthosterni). Euscorpius, 11: 1 - 175." title="High-Level Systematics and Phylogeny of the Extant Scorpions (Scorpiones: Orthosterni)" type="journal article" year="2003" yearSuffix="b">Soleglad & Fet, 2003b: 88</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 91, tab. 9;
|
||
<treatmentCitation id="0A8410B82056194AFAB588E4D96CF983" author="FET, V. & M. E. SOLEGLAD & G. LOWE" page="10-12" pageId="4" pageNumber="3" year="2005">
|
||
<bibRefCitation id="EFB44B582056194AFAB588E4D96CF983" author="Fet" etAl="et al." firstAuthor="Fet" journalOrPublisher="Euscorpius" page="10 - 12" pageId="4" pageNumber="3" pagination="1 - 40" part="23" refId="ref60407" refString="FET, V., M. E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1 - 40." title="A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida)" type="journal article" year="2005">Fet et al., 2005: 10–12</bibRefCitation>
|
||
</treatmentCitation>
|
||
, tab. 1, fig. 23, 29;
|
||
<treatmentCitation id="0A8410B82056194AFAF488C6D91CF9A5" author="PRENDINI, L. & W. WHEELER" page="481" pageId="4" pageNumber="3" year="2005">
|
||
<bibRefCitation id="EFB44B582056194AFAF488C6D91CF9A5" author="Prendini & Wheeler" firstAuthor="Prendini" journalOrPublisher="Cladistics" page="481" pageId="4" pageNumber="3" pagination="446 - 494" part="21" refId="ref63593" refString="PRENDINI, L. & W. WHEELER. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics, 21: 446 - 494." title="Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing" type="journal article" year="2005">Prendini & Wheeler, 2005: 481</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFBA58B20D8E0F9A5" author="HENDRIXSON, B. E." box="[1138,1440,1546,1570]" page="81-82" pageId="4" pageNumber="3" year="2006">
|
||
<bibRefCitation id="EFB44B582056194AFBA58B20D8DBF9A5" author="Hendrixson" box="[1138,1435,1546,1570]" firstAuthor="Hendrixson" journalOrPublisher="Fauna of Arabia" page="81 - 82" pageId="4" pageNumber="3" pagination="33 - 120" part="21" refId="ref61157" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." title="Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae)" type="journal article" year="2006">Hendrixson, 2006: 81–82</bibRefCitation>
|
||
;
|
||
</treatmentCitation>
|
||
<treatmentCitation id="0A8410B82056194AFCB88B03D963F9C6" author="DUPRE, G." box="[879,1059,1577,1601]" page="7" pageId="4" pageNumber="3" year="2007">
|
||
<bibRefCitation id="EFB44B582056194AFCB88B03D95EF9C6" author="Dupre" box="[879,1054,1577,1601]" firstAuthor="Dupre" journalOrPublisher="Euscorpius" page="7" pageId="4" pageNumber="3" pagination="1 - 31" part="50" refId="ref60020" refString="DUPRE, G. 2007. Conspectus genericus scorpionorum 1758 - 2006 (Arachnida: Scorpiones). Euscorpius, 50: 1 - 31." title="Conspectus genericus scorpionorum 1758 - 2006 (Arachnida: Scorpiones)" type="journal article" year="2007">Dupré, 2007: 7</bibRefCitation>
|
||
;
|
||
</treatmentCitation>
|
||
<treatmentCitation id="0A8410B82056194AFBE78B03D842F9C6" author="KOVARIK, F." box="[1072,1282,1577,1601]" page="24" pageId="4" pageNumber="3" year="2009">
|
||
<bibRefCitation id="EFB44B582056194AFBE78B03D842F9C6" author="Kovarik" bookContentInfo="170 pp." box="[1072,1282,1577,1601]" firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="24" pageId="4" pageNumber="3" refId="ref61808" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." title="Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species" type="book" year="2009">Kovařík, 2009: 24</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0A8410B82056194AFAC48B03DEA1F9E7" author="EL-HENNAWY, H. K." page="122" pageId="4" pageNumber="3" year="2009">
|
||
<bibRefCitation id="EFB44B582056194AFAC48B03DE9DF9E7" author="El-Hennawy" firstAuthor="El-Hennawy" journalOrPublisher="Serket" page="122" pageId="4" pageNumber="3" pagination="119 - 128" part="11" refId="ref60203" refString="EL-HENNAWY, H. K. 2009. Scorpions of Saudi Arabia. Serket, 11 (3 / 4): 119 - 128." title="Scorpions of Saudi Arabia" type="journal article" year="2009">El-Hennawy, 2009: 122</bibRefCitation>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F65222056194AFCE88B4DDEB7F91A" pageId="4" pageNumber="3" type="type_taxon">
|
||
<paragraph id="8B9A36A92056194AFCE88B4DDEB7F91A" blockId="4.[831,1441,1025,1693]" pageId="4" pageNumber="3">
|
||
<typeStatus id="549E880B2056194AFCE88B4DDEC2F9F9" box="[831,898,1639,1662]" pageId="4" pageNumber="3">TYPE</typeStatus>
|
||
SPECIES.
|
||
<taxonomicName id="4C254D2A2056194AFC288B4CDEB4F91A" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Leiurus">
|
||
<emphasis id="B951EABB2056194AFC288B4CD9C7F9F9" box="[1023,1159,1638,1662]" italics="true" pageId="4" pageNumber="3">Androctonus</emphasis>
|
||
(
|
||
<emphasis id="B951EABB2056194AFB4F8B4DD9A9F9F9" box="[1176,1257,1639,1662]" italics="true" pageId="4" pageNumber="3">Leiurus</emphasis>
|
||
)
|
||
<emphasis id="B951EABB2056194AFB2D8B4DD8E0F9F9" box="[1274,1440,1639,1662]" italics="true" pageId="4" pageNumber="3">quinquestriatus</emphasis>
|
||
Ehrenberg, 1828
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F65222056194AFCE88BEFDED3F8F0" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8B9A36A92056194AFCE88BEFDED3F8F0" blockId="4.[831,1441,1733,1911]" pageId="4" pageNumber="3">
|
||
ETYMOLOGY. Two subgenera of
|
||
<taxonomicName id="4C254D2A2056194AFB008BEFD81FF95A" authorityName="Ehrenberg" authorityYear="1828" box="[1239,1375,1733,1757]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABB2056194AFB008BEFD81FF95A" box="[1239,1375,1733,1757]" italics="true" pageId="4" pageNumber="3">Androctonus</emphasis>
|
||
</taxonomicName>
|
||
were created by Ehrenberg:
|
||
<taxonomicName id="4C254D2A2056194AFBB58BCFD9F4F97B" authorityName=": Vachon" authorityYear="1949" box="[1122,1204,1765,1788]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABB2056194AFBB58BCFD9F4F97B" box="[1122,1204,1765,1788]" italics="true" pageId="4" pageNumber="3">Leiurus</emphasis>
|
||
</taxonomicName>
|
||
(= ‘slender tail’), including species with more elongated metasomal segments, and
|
||
<taxonomicName id="4C254D2A2056194AFC098A08D90AF8BE" box="[990,1098,1826,1849]" class="Actinopterygii" family="Acanthuridae" genus="Prionurus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B951EABB2056194AFC098A08D90AF8BE" box="[990,1098,1826,1849]" italics="true" pageId="4" pageNumber="3">Prionurus</emphasis>
|
||
</taxonomicName>
|
||
(= ‘saw tail’) including species with heavier metasomal segments bearing serrated carinae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F65222057194BFF178DECD87FFCC4" pageId="5" pageNumber="4" type="diagnosis">
|
||
<paragraph id="8B9A36A92057194BFF178DECDC34FB75" blockId="5.[192,802,198,1266]" pageId="5" pageNumber="4">
|
||
DIAGNOSIS. Medium to large buthids (
|
||
<bibRefCitation id="EFB44B582057194BFDA98DECDE55FF59" author="Sissom" box="[638,789,198,222]" editor="POLIS, G. A." firstAuthor="Sissom" journalOrPublisher="The Biology of Scorpions. Stanford University Press, Stanford, CA" pageId="5" pageNumber="4" pagination="64 - 160" refId="ref64015" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in: POLIS, G. A. (ed.) The Biology of Scorpions. Stanford University Press, Stanford, CA." title="Systematics, biogeography and paleontology" type="book chapter" year="1990">Sissom, 1990</bibRefCitation>
|
||
), adults
|
||
<quantity id="4CDD9B4C2057194BFED18DCEDCC9FF7B" box="[262,393,228,252]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" metricValueMax="11.5" metricValueMin="5.1" pageId="5" pageNumber="4" unit="mm" value="83.0" valueMax="115.0" valueMin="51.0">51–115 mm</quantity>
|
||
; carapace and tergites with granulated carinae; carapace with distinct anterior, superciliary, central median, central lateral, posterior median and posterior lateral carinae; central lateral and posterior median carinae fused into a lyre configuration; tergites I–II, VII with 5 carinae, III–VI with 3 carinae; metasoma elongate, metasoma I–III with 10 carinae, median lateral carinae complete on I, reduced on II–III; metasoma IV with 8 carinae; metasoma V with 7 carinae, dorsolateral carinae weak, ventrolateral carinae strong with serrate or lobate dentition; telson with bulbous vesicle lacking subaculear spine or tubercle; pectines with fulcra; chelicerae with characteristic buthid dentition (
|
||
<bibRefCitation id="EFB44B582057194BFD118F7EDDB9FD0C" author="Vachon" firstAuthor="Vachon" pageId="5" pageNumber="4" pagination="161 - 166" refId="ref64728" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
|
||
), two denticles on ventral aspect of fixed finger; pedipalps orthobothriotaxic,
|
||
<typeStatus id="549E880B2057194BFDD08FBFDF75FD2A" box="[519,565,661,685]" pageId="5" pageNumber="4">type</typeStatus>
|
||
A
|
||
<emphasis id="B951EABB2057194BFD8E8FBEDF28FD2B" bold="true" box="[601,616,660,684]" pageId="5" pageNumber="4"></emphasis>
|
||
(
|
||
<bibRefCitation id="EFB44B582057194BFD578FBEDE5DFD2B" author="Vachon" box="[640,797,660,684]" firstAuthor="Vachon" pageId="5" pageNumber="4" pagination="857 - 958" refId="ref64809" refString="VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB44B582057194BFF178F98DDB9FD4D" author="Vachon" box="[192,249,690,714]" firstAuthor="Vachon" pageId="5" pageNumber="4" pagination="1597 - 1599" refId="ref64871" refString="VACHON, M. 1975. Sur l'utilisation de la tricho- bothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser. D Sciences Naturelles, 281 (21): 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
|
||
); pedipalp femur with trichobothrium
|
||
<emphasis id="B951EABB2057194BFD498F98DFEBFD4D" box="[670,683,690,714]" italics="true" pageId="5" pageNumber="4">d</emphasis>
|
||
<subScript id="17A134EC2057194BFD7C8F96DFF4FD4D" attach="left" box="[683,692,700,714]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
on dorsal surface, pedipalp patella with
|
||
<emphasis id="B951EABB2057194BFDD28FFBDF52FD6E" box="[517,530,721,745]" italics="true" pageId="5" pageNumber="4">d</emphasis>
|
||
<subScript id="17A134EC2057194BFDC58FF1DF5BFD6E" attach="left" box="[530,539,731,745]" fontSize="6" pageId="5" pageNumber="4">3</subScript>
|
||
internal to dorsomedian carina; chela smooth with carinae reduced or obsolete, dentate margins of fingers armed with linear subrows of primary denticles; subrows flanked by internal and external accessory denticles; movable finger with two enlarged subdistal internal denticles; males without scalloping at base of pedipalp fingers; tergites without macrosetae; tibial spurs present on legs III–IV; basitarsi I–III with regular series of macrosetae on retrosuperior, retroinferior and inferior margins; ventral surfaces of telotarsi with paired rows of macrosetae; prolateral and retrolateral tarsal spurs present on all legs.
|
||
<emphasis id="B951EABB2057194BFD0D8908DC7FFBDE" italics="true" pageId="5" pageNumber="4">Sexual dimorphism</emphasis>
|
||
: compared to females, males have a narrower mesosoma, more robust carination on tergites and sternites III–V, more slender pedipalps and metasoma, longer pectines with larger teeth, and weaker dentition or granulation on ventromedian carinae of metasoma II–III.
|
||
</paragraph>
|
||
<paragraph id="8B9A36A92057194BFF17883FD87FFCC4" blockId="5.[192,802,1301,1907]" lastBlockId="5.[831,1440,198,835]" pageId="5" pageNumber="4">
|
||
SPECIES COMPOSITION. The broad geographic range and morphological variation of
|
||
<taxonomicName id="4C254D2A2057194BFDC4881EDF24FACC" authorityName=": Vachon" authorityYear="1949" box="[531,612,1332,1355]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABB2057194BFDC4881EDF24FACC" box="[531,612,1332,1355]" italics="true" pageId="5" pageNumber="4">Leiurus</emphasis>
|
||
</taxonomicName>
|
||
populations have caused difficulty for taxonomists. For a long time, the genus was assumed to be represented by a single polytypic species spread over a vast area, although different authors disagreed about the definition and validity of various subspecies. More recently, several distinctive species were described from widely disparate localities:
|
||
<taxonomicName id="4C254D2A2057194BFEE28B21DF9CF9A5" authority="Lourenco et al., 2002" authorityName="Lourenco" authorityYear="2002" box="[309,732,1546,1570]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="jordanensis">
|
||
<emphasis id="B951EABB2057194BFEE28B21DC04F9A5" box="[309,324,1547,1570]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFE8F8B20DC95F9A5" box="[344,469,1546,1570]" italics="true" pageId="5" pageNumber="4">jordanensis</emphasis>
|
||
<bibRefCitation id="EFB44B582057194BFE358B21DF9CF9A5" author="Lourenco" box="[482,732,1546,1570]" etAl="et al." firstAuthor="Lourenco" pageId="5" pageNumber="4" pagination="635 - 642" refId="ref62388" refString="LOURENCO, W. R., D. MODRY & Z. AMR. 2002. Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan. Revue suisse de Zoologie, 109 (3): 635 - 642." type="journal article" year="2002">Lourenço et al., 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
, from sandstone cliffs isolated by dunes in
|
||
<collectingCountry id="F33276392057194BFD9C8B03DFD2F9C6" box="[587,658,1577,1601]" name="Jordan" pageId="5" pageNumber="4">Jordan</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="F33276392057194BFF178B6DDC11F9D8" box="[192,337,1607,1631]" name="Saudi Arabia" pageId="5" pageNumber="4">Saudi Arabia</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C254D2A2057194BFE8A8B62DFA2F9D8" authority="Lourenco et al., 2006" authorityName="Lourenco" authorityYear="2006" box="[349,738,1607,1631]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="savanicola">
|
||
<emphasis id="B951EABB2057194BFE8A8B62DC2CF9D8" box="[349,364,1608,1631]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFEAC8B6DDCAFF9D8" box="[379,495,1607,1631]" italics="true" pageId="5" pageNumber="4">savanicola</emphasis>
|
||
<bibRefCitation id="EFB44B582057194BFE208B62DFA2F9D8" author="Lourenco" box="[503,738,1607,1631]" etAl="et al." firstAuthor="Lourenco" pageId="5" pageNumber="4" pagination="97 - 101" refId="ref62489" refString="LOURENCO, W. R., J-X. QI & J. L. CLOUDSLEY- THOMPSON. 2006. The African species of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with the description of a new species. Boletin Sociedad Entomologica Aragonesa, 39: 97 - 101." type="journal article" year="2006">Lourenço et al., 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Sahel/ savannah in
|
||
<collectingCountry id="F33276392057194BFE588B4CDF42F9F9" box="[399,514,1638,1662]" name="Cameroon" pageId="5" pageNumber="4">Cameroon</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C254D2A2057194BFDDA8B4DDE5DF9F9" authority="Kovarik, 2007" authorityName="Kovarik" authorityYear="2007" box="[525,797,1638,1662]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="nasheri">
|
||
<emphasis id="B951EABB2057194BFDDA8B4DDF5CF9F9" box="[525,540,1639,1662]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFDFD8B4CDF3AF9F9" box="[554,634,1638,1662]" italics="true" pageId="5" pageNumber="4">nasheri</emphasis>
|
||
<bibRefCitation id="EFB44B582057194BFD568B4CDE5DF9F9" author="Kovarik" box="[641,797,1638,1662]" firstAuthor="Kovarik" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="5" pageNumber="4" pagination="137 - 141" part="71" refId="ref61777" refString="KOVARIK, F. 2007. Leiurus nasheri sp. nov. from Yemen (Scorpiones, Buthidae). Acta Societatis Zoologicae Bohemicae, 71: 137 - 141." title="Leiurus nasheri sp. nov. from Yemen (Scorpiones, Buthidae)" type="journal article" year="2007">Kovařík, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the humid Tihamah coastal plain in
|
||
<collectingCountry id="F33276392057194BFD458BAFDF9FF91B" box="[658,735,1669,1692]" name="Yemen" pageId="5" pageNumber="4">Yemen</collectingCountry>
|
||
(=
|
||
<taxonomicName id="4C254D2A2057194BFCDB8BACDF00F93C" authority="(Ehrenberg, 1829)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="brachycentrus">
|
||
<emphasis id="B951EABB2057194BFCDB8BACDE5BF91A" box="[780,795,1670,1693]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFF178B89DC19F93C" box="[192,345,1699,1723]" italics="true" pageId="5" pageNumber="4">brachycentrus</emphasis>
|
||
(Ehrenberg, 1829)
|
||
</taxonomicName>
|
||
stat. n.); and
|
||
<taxonomicName id="4C254D2A2057194BFCDB8B8EDF2CF95D" authority="Yagmur et al., 2009" authorityName="Yagmur" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="abdullahbayrami">
|
||
<emphasis id="B951EABB2057194BFCDB8B8EDE5BF93C" box="[780,795,1700,1723]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFF178BE8DC38F95D" box="[192,376,1730,1754]" italics="true" pageId="5" pageNumber="4">abdullahbayrami</emphasis>
|
||
<bibRefCitation id="EFB44B582057194BFE528BE8DF2CF95D" author="Yagmur" box="[389,620,1730,1754]" etAl="et al." firstAuthor="Yagmur" pageId="5" pageNumber="4" pagination="1 - 20" refId="ref65464" refString="YAGMUR, E. A., H. KOC & K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." type="journal article" year="2009">Yağmur et al., 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, from semi- arid rocky steppe in
|
||
<collectingCountry id="F33276392057194BFEBB8BCBDCF9F97E" box="[364,441,1761,1785]" name="Turkey" pageId="5" pageNumber="4">Turkey</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="F33276392057194BFD868BCBDFCDF97E" box="[593,653,1761,1785]" name="Syria" pageId="5" pageNumber="4">Syria</collectingCountry>
|
||
. Here we add four new species from the Arabian Peninsula:
|
||
<taxonomicName id="4C254D2A2057194BFCDB8A2ADC5DF8B1" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFCDB8A2ADE5BF890" box="[780,795,1792,1815]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFF178A34DC5DF8B1" box="[192,285,1822,1846]" italics="true" pageId="5" pageNumber="4">haenggii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C02057194BFEF28A35DC1FF8B1" box="[293,351,1823,1846]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
, from mountain ranges along the Red Sea coast (Hijaz and
|
||
<collectingRegion id="49E1F84B2057194BFEA28A17DCE4F8D2" box="[373,420,1853,1877]" country="Saudi Arabia" name="'Asir" pageId="5" pageNumber="4">Asir</collectingRegion>
|
||
) in
|
||
<collectingCountry id="F33276392057194BFE198A17DF1DF8D2" box="[462,605,1853,1877]" name="Saudi Arabia" pageId="5" pageNumber="4">Saudi Arabia</collectingCountry>
|
||
, the
|
||
<collectingRegion id="49E1F84B2057194BFD588A17DE45F8D2" box="[655,773,1853,1877]" country="Yemen" name="Hadramawt" pageId="5" pageNumber="4">Hadramaut</collectingRegion>
|
||
in
|
||
<collectingCountry id="F33276392057194BFF178A76DC50F8F4" box="[192,272,1884,1907]" name="Yemen" pageId="5" pageNumber="4">Yemen</collectingCountry>
|
||
, and
|
||
<collectingRegion id="49E1F84B2057194BFE9E8A71DCD4F8F4" box="[329,404,1883,1907]" country="Oman" name="Zufar" pageId="5" pageNumber="4">Dhofar</collectingRegion>
|
||
mountains of
|
||
<collectingCountry id="F33276392057194BFDE68A71DF35F8F4" box="[561,629,1883,1907]" name="Oman" pageId="5" pageNumber="4">Oman</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C254D2A2057194BFD568A76DFBBF8F4" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[641,763,1883,1907]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFD568A76DFD0F8F4" box="[641,656,1884,1907]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFD498A71DFBBF8F4" box="[670,763,1883,1907]" italics="true" pageId="5" pageNumber="4">arabicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C02057194BFCD48A76DE14FF5A" box="[771,852,198,1907]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
, from arid plains and wadis of the central Najd plateau, extending to the east coast of
|
||
<collectingCountry id="F33276392057194BFB3D8DCED83DFF7B" box="[1258,1405,228,252]" name="Saudi Arabia" pageId="5" pageNumber="4">Saudi Arabia</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C254D2A2057194BFA5C8DCFDE87FE9C" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="macroctenus" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFA5C8DCFD8DAFF7B" box="[1419,1434,229,252]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFCE88C2EDE87FE9C" box="[831,967,260,283]" italics="true" pageId="5" pageNumber="4">macroctenus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C02057194BFC068C2ED94EFE9C" box="[977,1038,260,283]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
, from coastal fog desert of Jiddat al Harasis in
|
||
<collectingCountry id="F33276392057194BFC658C0BDEB7FEBE" box="[946,1015,289,313]" name="Oman" pageId="5" pageNumber="4">Oman</collectingCountry>
|
||
; and
|
||
<taxonomicName id="4C254D2A2057194BFBE68C08D9DAFEBE" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[1073,1178,289,313]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFBE68C08D900FEBE" box="[1073,1088,290,313]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFB998C0BD9DAFEBE" box="[1102,1178,289,313]" italics="true" pageId="5" pageNumber="4">heberti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C02057194BFB768C08D99BFEBE" box="[1185,1243,290,313]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
from rocky wadis of the Jabal Samhan mountains in
|
||
<collectingRegion id="49E1F84B2057194BFB028C6AD863FEDF" box="[1237,1315,320,344]" country="Oman" name="Zufar" pageId="5" pageNumber="4">Dhofar</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33276392057194BFAE38C6AD839FEDF" box="[1332,1401,320,344]" name="Oman" pageId="5" pageNumber="4">Oman</collectingCountry>
|
||
. In addition, we propose revised diagnoses for:
|
||
<taxonomicName id="4C254D2A2057194BFAF78C4ADEA2FE11" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hebraeus">
|
||
<emphasis id="B951EABB2057194BFAF78C4AD86FFEF0" box="[1312,1327,352,375]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFAE98C75D8E0FEF0" box="[1342,1440,351,375]" italics="true" pageId="5" pageNumber="4">hebraeus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB44B582057194BFC918C54DE9BFE11" author="Birula" box="[838,987,382,406]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="5" pageNumber="4" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
stat. n. from semi-arid/ arid steppe habitats in
|
||
<collectingCountry id="F33276392057194BFC178CB6DEBEFE33" box="[960,1022,412,436]" name="Israel" pageId="5" pageNumber="4">Israel</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33276392057194BFBD88CB6D916FE33" box="[1039,1110,412,436]" name="Jordan" pageId="5" pageNumber="4">Jordan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33276392057194BFB408CB6D992FE33" box="[1175,1234,412,436]" name="Syria" pageId="5" pageNumber="4">Syria</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C254D2A2057194BFB328CB7D8E0FE33" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[1253,1440,412,436]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="brachycentrus">
|
||
<emphasis id="B951EABB2057194BFB328CB7D9B4FE33" box="[1253,1268,413,436]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFAD08CB6D8E0FE33" box="[1287,1440,412,436]" italics="true" pageId="5" pageNumber="4">brachycentrus</emphasis>
|
||
</taxonomicName>
|
||
stat. n. from the Tihamah plain along the Red Sea coast of western
|
||
<collectingCountry id="F33276392057194BFC6C8CF0D948FE76" box="[955,1032,474,497]" name="Yemen" pageId="5" pageNumber="4">Yemen</collectingCountry>
|
||
and southwestern
|
||
<collectingCountry id="F33276392057194BFB0F8CF3D82BFE76" box="[1240,1387,473,497]" name="Saudi Arabia" pageId="5" pageNumber="4">Saudi Arabia</collectingCountry>
|
||
; and for the
|
||
<typeStatus id="549E880B2057194BFC4F8CD3DE86FD96" box="[920,966,505,529]" pageId="5" pageNumber="4">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C254D2A2057194BFBF98CD3D9B6FD97" baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[1070,1270,505,528]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="quinquestriatus">
|
||
<emphasis id="B951EABB2057194BFBF98CD3D97DFD97" box="[1070,1085,505,528]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFB878CD3D9B6FD97" box="[1104,1270,505,528]" italics="true" pageId="5" pageNumber="4">quinquestriatus</emphasis>
|
||
</taxonomicName>
|
||
from the Nile Valley and surrounding deserts in
|
||
<collectingCountry id="F33276392057194BFB048F32D853FDB7" box="[1235,1299,536,560]" name="Egypt" pageId="5" pageNumber="4">Egypt</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33276392057194BFA818F3DD8DBFDA8" box="[1366,1435,535,559]" name="Sudan" pageId="5" pageNumber="4">Sudan</collectingCountry>
|
||
, bringing the total number of included species to ten. Our findings show that, like many other scorpion genera,
|
||
<taxonomicName id="4C254D2A2057194BFCE88F5EDED0FD0C" authorityName=": Vachon" authorityYear="1949" box="[831,912,628,651]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABB2057194BFCE88F5EDED0FD0C" box="[831,912,628,651]" italics="true" pageId="5" pageNumber="4">Leiurus</emphasis>
|
||
</taxonomicName>
|
||
is comprised of an assemblage of allopatric or parapatric species spread across different regions separated by physiographic barriers, each adapted to local environments and substrates. Additional species diversity may emerge when other local populations are analyzed in more detail, for example those in southern Sinai, and in more central parts of North Africa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F65222057194BFC9A8E45D8D1FC00" box="[845,1425,879,903]" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="8B9A36A92057194BFC9A8E45D8D1FC00" blockId="5.[845,1425,879,903]" box="[845,1425,879,903]" pageId="5" pageNumber="4">
|
||
<heading id="D0D281C52057194BFC9A8E45D8D1FC00" box="[845,1425,879,903]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="4" reason="4">
|
||
<emphasis id="B951EABB2057194BFC9A8E45D8D1FC00" bold="true" box="[845,1425,879,903]" pageId="5" pageNumber="4">
|
||
Key to adult
|
||
<taxonomicName id="4C254D2A2057194BFC368E45D9CBFC00" box="[993,1163,879,903]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B951EABB2057194BFC368E45D977FC00" bold="true" box="[993,1079,879,903]" italics="true" pageId="5" pageNumber="4">Leiurus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
examined in this study
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F652220571948FCE88E85DE5DFC2E" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="key">
|
||
<paragraph id="8B9A36A92057194BFCE88E85D8DDFBA4" blockId="5.[831,1440,943,1059]" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C2057194BFCE88E85D8DDFBA4" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C2057194BFCE88E85D8DDFBA4" pageId="5" pageNumber="4">1 Medial intercarinal surfaces of tergites II–III smooth or sparsely, lightly shagreened .....……………...... 2 Medial intercarinal surfaces of tergites II–III heavily or densely, finely shagreened .......……….............. 4</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A92057194BFCE88915D8DBFB77" blockId="5.[831,1440,1087,1264]" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C2057194BFCE88915D8DBFB77" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C2057194BFCE88915D8DBFB77" pageId="5" pageNumber="4">
|
||
<specimenCount id="9D23FD202057194BFCE88915DEF0FBD0" box="[831,944,1087,1111]" pageId="5" pageNumber="4" type="male">2 Males</specimenCount>
|
||
with metasoma III L/W> 2.10, metasoma IV L/W> 2.50; metasoma III ventromedian carinae with> 35 denticles .........................
|
||
<taxonomicName id="4C254D2A2057194BFB228957D81EFB13" ID-CoL="6P795" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[1269,1374,1148,1172]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="heberti" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFB228957D844FB13" box="[1269,1284,1149,1172]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFAC58956D81EFB13" box="[1298,1374,1148,1172]" italics="true" pageId="5" pageNumber="4">heberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB2057194BFAB38957D8DFFB12" bold="true" box="[1380,1439,1149,1173]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A26257C02057194BFAB38957D8DFFB12" box="[1380,1439,1149,1173]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Males with metasoma III L/W <2.10, metasoma IV L/W <2.50; metasoma III ventromedian carinae with <35 denticles ...............…………….............. 3
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A92057194BFCE88827D8DCFA39" blockId="5.[831,1441,1293,1470]" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C2057194BFCE88827D8DCFA39" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C2057194BFCE88827D8DCFA39" pageId="5" pageNumber="4">
|
||
<specimenCount id="9D23FD202057194BFCE88827DE88FAA2" box="[831,968,1293,1317]" pageId="5" pageNumber="4" type="female">3 Females</specimenCount>
|
||
with pedipalp patella L/W> 3.20,
|
||
<emphasis id="B951EABB2057194BFA9F8827D818FAA3" box="[1352,1368,1293,1316]" italics="true" pageId="5" pageNumber="4">F</emphasis>
|
||
<subScript id="17A134EC2057194BFA8F883DD81FFAA2" attach="left" box="[1368,1375,1303,1317]" fontSize="6" pageId="5" pageNumber="4">s</subScript>
|
||
> 23; female sternites III–IV with weak to moderate median carinae .........……............
|
||
<taxonomicName id="4C254D2A2057194BFB378861D819FAE5" ID-CoL="6P6X4" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[1248,1369,1354,1378]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFB378861D9AFFAE5" box="[1248,1263,1355,1378]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFB2B8860D819FAE5" box="[1276,1369,1354,1378]" italics="true" pageId="5" pageNumber="4">arabicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB2057194BFAB78860D8DCFAE5" bold="true" box="[1376,1436,1354,1378]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A26257C02057194BFAB78860D8DCFAE5" box="[1376,1436,1354,1378]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Females with pedipalp patella L/W <3.20,
|
||
<emphasis id="B951EABB2057194BFA9F8843D818FA07" box="[1352,1368,1385,1408]" italics="true" pageId="5" pageNumber="4">F</emphasis>
|
||
<subScript id="17A134EC2057194BFA8F8859D81FFA06" attach="left" box="[1368,1375,1395,1409]" fontSize="6" pageId="5" pageNumber="4">s</subScript>
|
||
<23; female sternites III–IV with weak to obsolete median carinae ...…..…...............
|
||
<taxonomicName id="4C254D2A2057194BFB37888DD819FA39" ID-CoL="6P78T" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[1248,1369,1446,1470]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="haenggii" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFB37888DD9AFFA39" box="[1248,1263,1447,1470]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFB2B888CD819FA39" box="[1276,1369,1446,1470]" italics="true" pageId="5" pageNumber="4">haenggii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB2057194BFAB7888CD8DCFA39" bold="true" box="[1376,1436,1446,1470]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A26257C02057194BFAB7888CD8DCFA39" box="[1376,1436,1446,1470]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A92057194BFCE888F0D8DBF9C9" blockId="5.[831,1440,1498,1614]" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C2057194BFCE888F0D8DBF9C9" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C2057194BFCE888F0D8DBF9C9" pageId="5" pageNumber="4">
|
||
<specimenCount id="9D23FD202057194BFCE888F0DEE6FA75" box="[831,934,1498,1522]" pageId="5" pageNumber="4" type="male">4 Male</specimenCount>
|
||
pectines with mid-pectine sensillar margin L/ metasoma I W> 0.20 .....…...…….........................
|
||
<specimenCount id="9D23FD202057194BFA5888D3DEE6F9A8" pageId="5" pageNumber="4" type="male">5 Male</specimenCount>
|
||
pectines with mid-pectine sensillar margin L/ metasoma I W <0.20 ......………........................... 6
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A92057194BFCE88B40D8DEF97A" blockId="5.[831,1440,1642,1789]" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C2057194BFCE88B40D8DEF97A" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C2057194BFCE88B40D8DEF97A" pageId="5" pageNumber="4">
|
||
<specimenCount id="9D23FD202057194BFCE88B40DEFDF905" box="[831,957,1642,1666]" pageId="5" pageNumber="4" type="female">5 Female</specimenCount>
|
||
pectines with mid-pectine sensillar margin L/ metasoma I W> 0.15 …...
|
||
<taxonomicName id="4C254D2A2057194BFB618BA0D81BF926" ID-CoL="6P6WR" authority="Lowe & Yağmur & Kovařík, 2014" authorityName="Lowe & Yağmur & Kovařík" authorityYear="2014" box="[1206,1371,1673,1697]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="macroctenus" status="sp. nov.">
|
||
<emphasis id="B951EABB2057194BFB618BA0D985F926" box="[1206,1221,1674,1697]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFB048BA0D81BF926" box="[1235,1371,1674,1697]" italics="true" pageId="5" pageNumber="4">macroctenus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB2057194BFAB58BA3D8DDF926" bold="true" box="[1378,1437,1673,1697]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A26257C02057194BFAB58BA3D8DDF926" box="[1378,1437,1673,1697]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Female pectines with mid-pectine sensillar margin L/ metasoma I W <0.15 ........……........................... .............
|
||
<taxonomicName id="4C254D2A2057194BFC1B8BCCD80BF97A" ID-CoL="6P794" authority="(Ehrenberg, 1829)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1829" box="[972,1355,1765,1789]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="brachycentrus">
|
||
<emphasis id="B951EABB2057194BFC1B8BCCDE9BF97A" box="[972,987,1766,1789]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFC3F8BCFD9C1F97A" box="[1000,1153,1765,1789]" italics="true" pageId="5" pageNumber="4">brachycentrus</emphasis>
|
||
(Ehrenberg, 1829)
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB2057194BFA858BCFD8DEF97A" bold="true" box="[1362,1438,1765,1789]" pageId="5" pageNumber="4">stat. n.</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A920571948FCE88A33DE5BFF7B" blockId="5.[831,1440,1817,1902]" lastBlockId="6.[240,801,198,253]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
<keyLead id="30D4959C20571948FCE88A33DE5BFF7B" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
<keyStep id="30D12E0C20571948FCE88A33DE5BFF7B" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
6 Metasoma III ventromedian carinae with> 30 denticles; metasoma I–IV uniformly fuscous ……......... …....
|
||
<taxonomicName id="4C254D2A2057194BFC7C8A7DD8DAF8E9" ID-CoL="6P78Q" authority="Lourenco, Modry et Amr, 2002 Metasoma III" authorityName="Lourenco, Modry et Amr" authorityYear="2002" box="[939,1434,1878,1902]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="jordanensis">
|
||
<emphasis id="B951EABB2057194BFC7C8A7DDEFAF8E9" box="[939,954,1879,1902]" italics="true" pageId="5" pageNumber="4">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB2057194BFC108A7CD903F8E9" box="[967,1091,1878,1902]" italics="true" pageId="5" pageNumber="4">jordanensis</emphasis>
|
||
<bibRefCitation id="EFB44B582057194BFB9D8A7DD8DAF8E9" author="Lourenco, Modry et Amr" box="[1098,1434,1878,1902]" firstAuthor="Lourenco" journalOrPublisher="Revue suisse de Zoologie" pageId="5" pageNumber="4" pagination="635 - 642" part="109" refId="ref62388" refString="LOURENCO, W. R., D. MODRY & Z. AMR. 2002. Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan. Revue suisse de Zoologie, 109 (3): 635 - 642." title="Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from the south of Jordan" type="journal article" year="2002">Lourenço, Modrý et Amr, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
Metasoma III ventromedian carinae with <30 denticles; metasoma I–IV yellow to yellow-orange…. 7
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A920541948FF178C3CDE5CFE0E" blockId="6.[192,802,277,393]" pageId="6" pageNumber="5">
|
||
<keyLead id="30D4959C20541948FF178C3CDE5CFE0E" pageId="6" pageNumber="5">
|
||
<keyStep id="30D12E0C20541948FF178C3CDE5CFE0E" pageId="6" pageNumber="5">
|
||
7 Sternite VII with medial intercarinal surface smooth .....
|
||
<taxonomicName id="4C254D2A20541948FECF8C1FDE5DFECB" ID-CoL="6P797" authority="Yagmur, Koc et Kunt, 2009" authorityName="Yagmur, Koc et Kunt" authorityYear="2009" box="[280,797,308,332]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="abdullahbayrami">
|
||
<emphasis id="B951EABB20541948FECF8C1FDC67FECB" box="[280,295,309,332]" italics="true" pageId="6" pageNumber="5">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB20541948FEE28C1EDCACFECB" box="[309,492,308,332]" italics="true" pageId="6" pageNumber="5">abdullahbayrami</emphasis>
|
||
<bibRefCitation id="EFB44B5820541948FE248C1EDE5DFECB" author="Yagmur, Koc et Kunt" box="[499,797,308,332]" firstAuthor="Yagmur" journalOrPublisher="Euscorpius" pageId="6" pageNumber="5" pagination="1 - 20" part="85" refId="ref65464" refString="YAGMUR, E. A., H. KOC & K. B. KUNT. 2009. Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey. Euscorpius, 85: 1 - 20." title="Des- cription of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southeastern Turkey" type="journal article" year="2009">Yağmur, Koç et Kunt, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
Sternite VII with medial intercarinal surface shagreened .........…..…................................................ 8
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9A36A920541948FF178C8CDE5DFC2E" blockId="6.[192,802,422,937]" pageId="6" pageNumber="5">
|
||
<keyLead id="30D4959C20541948FF178C8CDE5DFC2E" pageId="6" pageNumber="5">
|
||
<keyStep id="30D12E0C20541948FF178C8CDE5DFC2E" pageId="6" pageNumber="5">
|
||
8 Carapace with anteromedian intercarinal surface smooth or sparsely shagreened; posterior median furrow of carapace shallow or flat, lateral granule arcs reduced or absent; females with pectine basal piece heavily shagreened, median carinae of sternites IV–V moderate, usually granular; males with pedipalp patella L/W> 3.1, metasomal segment L/W: II> 1.55, III> 1.7, IV> 2.05 .....…….......... …………...…
|
||
<taxonomicName id="4C254D2A20541948FE478FB6DE5DFD34" ID-CoL="6P78R" authority="(Ehrenberg, 1828)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[400,797,667,691]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="quinquestriatus">
|
||
<emphasis id="B951EABB20541948FE478FB6DCDFFD34" box="[400,415,668,691]" italics="true" pageId="6" pageNumber="5">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB20541948FE7B8FB6DF12FD34" box="[428,594,668,691]" italics="true" pageId="6" pageNumber="5">quinquestriatus</emphasis>
|
||
(Ehrenberg, 1828)
|
||
</taxonomicName>
|
||
Carapace with anteromedian intercarinal surface shagreened; posterior median furrow of carapace moderately deep, flanked by lateral granule arcs; females with pectine basal piece smooth or lightly shagreened, median carinae of sternites IV–V weak, smooth or obsolete; males with pedipalp patella L/W <3.1, metasomal segment L/W: II <1.55, III <1.7, IV <2.05 ...
|
||
<taxonomicName id="4C254D2A20541948FE798EBBDF8AFC2F" ID-CoL="6P6WQ" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[430,714,912,936]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="hebraeus">
|
||
<emphasis id="B951EABB20541948FE798EBBDCFDFC2F" box="[430,445,913,936]" italics="true" pageId="6" pageNumber="5">L</emphasis>
|
||
.
|
||
<emphasis id="B951EABB20541948FE1D8EBADF6CFC2F" box="[458,556,912,936]" italics="true" pageId="6" pageNumber="5">hebraeus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB44B5820541948FDED8EBADF83FC2F" author="Birula" box="[570,707,912,936]" firstAuthor="Birula" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien" pageId="6" pageNumber="5" pagination="121 - 152" part="117" refId="ref59140" refString="BIRULA, A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, 117 2 (1): 121 - 152." title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Agyp- tischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae" type="journal article" year="1908">Birula, 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B951EABB20541948FD068EBBDE5DFC2E" bold="true" box="[721,797,913,937]" pageId="6" pageNumber="5">stat. n.</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |