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<document id="AFA7F5C2BF112C0B64D7B0CAF8328F00" ID-CLB-Dataset="30108" ID-DOI="10.11646/zootaxa.4418.5.5" ID-GBIF-Dataset="d9133d62-096e-4ec1-873a-4a3af73c050c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1245022" ID-ZooBank="4757CFC6-6307-4A37-B210-ACC16EB666F8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525974619032" checkinUser="plazi" docAuthor="Özbek, Hasan Hüseyin &amp; Mašán, Peter" docDate="2018" docId="038C87B5FFB3B11853ECF8AAFD3D3A8D" docLanguage="en" docName="zootaxa.4418.5.5.pdf" docOrigin="Zootaxa 4418 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachylaelaps (Pachylaelaps) armiger Özbek &amp; Mašán 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="490" masterDocId="FFB5FFCDFFB6B111537BFFA5FFE23841" masterDocTitle="A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae)" masterLastPageNumber="492" masterPageNumber="481" pageNumber="486" updateTime="1698543763606" updateUser="ExternalLinkService">
<mods:mods id="96333A63B80B1D7977190F7454595526" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B4E307A302AB770C1BD1567DAF801AE1">
<mods:title id="FBA57F1AEA1D0118DBCC148A5C6DDFEA">A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae)</mods:title>
</mods:titleInfo>
<mods:name id="DAAAB5BB20844120016AF4E7947AB135" type="personal">
<mods:role id="90CE3319CEFB9117F2AAB623DBEFEA5B">
<mods:roleTerm id="6322D4F796A50ED152A459B486F66FA4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F1FAA43426259A8BBB8689F0803754BB">Özbek, Hasan Hüseyin</mods:namePart>
</mods:name>
<mods:name id="433BEA8B3C5347BD3941D9D6BB791461" type="personal">
<mods:role id="A4C1A2BA8E1B17AFAC95B06BEFEEF5A8">
<mods:roleTerm id="3D132EF40ABF9179E52B67AD4EED7B99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBC71BCB66F9CF6790CF6566ED4FC447">Mašán, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="9E4B97E0ACC2E5F31519BA4B7EB28CA4">text</mods:typeOfResource>
<mods:relatedItem id="A8ED3311CA9D1CC40807224AD34526B6" type="host">
<mods:titleInfo id="875B69F7C2BB4E61435A16E341FF0FEF">
<mods:title id="82FC7CDB2768E7957DB802F4CA4FFDBF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E0CE138C07726BAAEBE08D57419D26BA">
<mods:date id="43BB39A24A9616F3AB835E34524A46D3">2018</mods:date>
<mods:detail id="637A5130C3193559A078B8ED6AED5D92" type="volume">
<mods:number id="4384B3E8E72006D5FC6AC2F9FE006D44">4418</mods:number>
</mods:detail>
<mods:detail id="58A8DD29395E9814E03A56289B48928A" type="issue">
<mods:number id="77442188E63CC9AA65AECB147D94A3C6">5</mods:number>
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<mods:extent id="F7332ACBFF8EDC93E7283EC220B42844" unit="page">
<mods:start id="35F3A95E0E6B3EEF8A7674AC54BCDF34">481</mods:start>
<mods:end id="8298EEBB67D20021EB0A9CCC0CEA3EB5">492</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8A806E59CE1E47D63F18F34CD9D44B9B">journal article</mods:classification>
<mods:identifier id="90FA750D6E7D67987A7365F797ADEAAE" type="CLB-Dataset">30108</mods:identifier>
<mods:identifier id="172FC0F436ECA2E3C120299C5AC25482" type="DOI">10.11646/zootaxa.4418.5.5</mods:identifier>
<mods:identifier id="1D1E0651D66A95EEF3006FF40D80A52D" type="GBIF-Dataset">d9133d62-096e-4ec1-873a-4a3af73c050c</mods:identifier>
<mods:identifier id="CF6DD807765F379939073CB984AF997D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4959DC796175CF3743D0AF0607A36B6F" type="Zenodo-Dep">1245022</mods:identifier>
<mods:identifier id="0D18B20E5954DB1F5AA2F2336B946803" type="ZooBank">4757CFC6-6307-4A37-B210-ACC16EB666F8</mods:identifier>
</mods:mods>
<treatment id="038C87B5FFB3B11853ECF8AAFD3D3A8D" ID-DOI="http://doi.org/10.5281/zenodo.5966876" ID-GBIF-Taxon="143929644" ID-Zenodo-Dep="5966876" LSID="urn:lsid:plazi:treatment:038C87B5FFB3B11853ECF8AAFD3D3A8D" httpUri="http://treatment.plazi.org/id/038C87B5FFB3B11853ECF8AAFD3D3A8D" lastPageId="9" lastPageNumber="490" pageId="5" pageNumber="486">
<subSubSection id="C33F6528FFB3B11453ECF8AAFEC53F0A" pageId="5" pageNumber="486" type="nomenclature">
<paragraph id="8B9A36A3FFB3B11453ECF8AAFD5F3F6B" blockId="5.[151,701,1807,1867]" box="[151,701,1807,1834]" pageId="5" pageNumber="486">
<heading id="D0D281CFFFB3B11453ECF8AAFD5F3F6B" bold="true" box="[151,701,1807,1834]" fontSize="11" level="1" pageId="5" pageNumber="486" reason="1">
<emphasis id="B951EAB1FFB3B11453ECF8AAFD5F3F6B" bold="true" box="[151,701,1807,1834]" pageId="5" pageNumber="486">
<taxonomicName id="4C254D20FFB3B11453ECF8AAFDBA3F68" ID-CoL="8Y7L2" authority="Özbek &amp; Mašán, 2018" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[151,600,1807,1834]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="486" phylum="Arthropoda" rank="species" species="armiger" status="sp. nov." subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFB3B11453ECF8AAFED53F68" bold="true" box="[151,311,1807,1833]" italics="true" pageId="5" pageNumber="486">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFB3B114523CF8AAFE043F68" bold="true" box="[327,486,1807,1833]" italics="true" pageId="5" pageNumber="486">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFB3B1145283F8AAFDBA3F68" bold="true" box="[504,600,1807,1833]" italics="true" pageId="5" pageNumber="486">armiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257CAFFB3B1145124F8B5FD5F3F6B" box="[607,701,1808,1834]" pageId="5" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36A3FFB3B11453ECF897FEC53F0A" blockId="5.[151,701,1807,1867]" box="[151,295,1842,1867]" pageId="5" pageNumber="486">
(
<figureCitation id="131E2A26FFB3B11453E4F897FEFD3F0A" box="[159,287,1842,1867]" captionStart-0="FIGURES 15–24" captionStart-1="FIGURES 25–26" captionStart-2="FIGURES 27–33" captionStart-3="FIGURES 34–39" captionStartId-0="4.[151,263,1909,1932]" captionStartId-1="6.[151,263,1515,1538]" captionStartId-2="7.[151,263,1244,1267]" captionStartId-3="8.[151,263,1932,1955]" captionTargetBox-0="[178,1406,195,1885]" captionTargetBox-1="[162,1427,350,1485]" captionTargetBox-2="[158,1430,309,1215]" captionTargetBox-3="[151,1436,373,1911]" captionTargetId-0="figure@4.[178,1406,195,1885]" captionTargetId-1="figure@6.[151,1436,337,1494]" captionTargetId-2="figure@7.[151,1436,301,1223]" captionTargetId-3="figure@8.[151,1436,373,1911]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURES 15–24. Pachylaelaps spp. 15‒23. Pachylaelaps (Longipachys) anatolicus. 15. Chelicera, male, variant fOrms; 16. Chelicera, female; 17. Terminal parts Of palps (tarsi and tibiae), male; 18. Leg II (tarsus, tibia and genu), female; 19. Leg II (tarsus, tibia, genu and femur), male; 20. Spermathecal duct, female; 21‒22. EpistOme, female, variant fOrms; 23. EpistOme, male. 24. Pachylaelaps (Pachylaelaps) armiger sp. nov., chelicera, female.NOt scaled." captionText-1="FIGURES 25–26. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 25. DOrsal idiOsOma; 26. Ventral idiOsOma. Scale bar:200 µm." captionText-2="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." captionText-3="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri-0="https://zenodo.org/record/1245028/files/figure.png" httpUri-1="https://zenodo.org/record/1245030/files/figure.png" httpUri-2="https://zenodo.org/record/1245032/files/figure.png" httpUri-3="https://zenodo.org/record/1245034/files/figure.png" pageId="5" pageNumber="486">Figs 24‒38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F6528FFB3B11753ECF8DFFD99395D" lastPageId="6" lastPageNumber="487" pageId="5" pageNumber="486" type="materials_examined">
<paragraph id="8B9A36A3FFB3B11753ECF8DFFD99395D" blockId="5.[151,1437,1914,2011]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="487" pageId="5" pageNumber="486">
<materialsCitation id="3B4D3CFEFFB3B11453ECF8DFFC5C3FF6" ID-GBIF-Occurrence="1844420396" collectingDate="2012-05-19" collectorName="Type &amp; Turkey" country="Turkey" elevation="1430" latitude="39.716667" location="Geyikli Village" longLatPrecision="1167" longitude="39.666668" pageId="5" pageNumber="486" specimenCount="2" stateProvince="Erzincan Province" typeStatus="holotype">
<emphasis id="B951EAB1FFB3B11453ECF8DFFE0A3FD2" bold="true" box="[151,488,1914,1939]" pageId="5" pageNumber="486">
<collectorName id="26D05375FFB3B11453ECF8DFFF303FD2" box="[151,210,1914,1939]" pageId="5" pageNumber="486">
<typeStatus id="549E8801FFB3B11453ECF8DFFF303FD2" box="[151,210,1914,1939]" pageId="5" pageNumber="486">Type</typeStatus>
</collectorName>
specimens examined.
</emphasis>
<typeStatus id="549E8801FFB3B1145282F8DFFD833FD2" box="[505,609,1914,1939]" pageId="5" pageNumber="486" type="holotype">Holotype</typeStatus>
female—NE
<collectorName id="26D05375FFB3B114506AF8DFFC803FD2" box="[785,866,1914,1939]" pageId="5" pageNumber="486">
<collectingCountry id="F3327633FFB3B114506AF8DFFC803FD2" box="[785,866,1914,1939]" name="Turkey" pageId="5" pageNumber="486">Turkey</collectingCountry>
</collectorName>
,
<collectingRegion id="49E1F841FFB3B114500CF8DFFBB03FD2" box="[887,1106,1914,1939]" country="Turkey" name="Erzincan" pageId="5" pageNumber="486">Erzincan Province</collectingRegion>
,
<location id="8EFA6078FFB3B1145713F8DFFAFC3FD2" LSID="urn:lsid:plazi:treatment:038C87B5FFB3B11853ECF8AAFD3D3A8D:8EFA6078FFB3B1145713F8DFFAFC3FD2" box="[1128,1310,1914,1939]" country="Turkey" latitude="39.716667" longLatPrecision="1167" longitude="39.666668" name="Geyikli Village" pageId="5" pageNumber="486" stateProvince="Erzincan Province">Geyikli Village</location>
(
<geoCoordinate id="EE115064FFB3B1145643F8DEFA753FD2" box="[1336,1431,1915,1939]" direction="north" orientation="latitude" pageId="5" pageNumber="486" precision="925" value="39.716667">39°43'N</geoCoordinate>
,
<geoCoordinate id="EE115064FFB3B11453ECF83AFF113FF6" box="[151,243,1951,1975]" direction="east" orientation="longitude" pageId="5" pageNumber="486" precision="925" value="39.666668">39°40'E</geoCoordinate>
), altitude
<elevation id="0008D190FFB3B1145215F83AFE293FF7" box="[366,459,1951,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="5" pageNumber="486" unit="m" value="1430.0">
<quantity id="4CDD9B46FFB3B1145215F83AFE293FF7" box="[366,459,1951,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="5" pageNumber="486" unit="m" value="1430.0">1430 m</quantity>
</elevation>
, leaf litter under
<taxonomicName id="4C254D20FFB3B11451DAF83BFD3B3FF7" box="[673,729,1950,1974]" class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="486" phylum="Magnoliopsida" rank="genus">
<emphasis id="B951EAB1FFB3B11451DAF83BFD3B3FF7" box="[673,729,1950,1974]" italics="true" pageId="5" pageNumber="486">Salix</emphasis>
</taxonomicName>
sp.,
<collectingDate id="EFDFE98BFFB3B1145061F83AFC5C3FF6" box="[794,958,1951,1975]" pageId="5" pageNumber="486" value="2012-05-19">
<date id="FF9B1063FFB3B1145061F83AFC5C3FF6" box="[794,958,1951,1975]" pageId="5" pageNumber="486" value="2012-05-19">19 May 2012</date>
</collectingDate>
</materialsCitation>
. Paratypes: one female (deposited in IZ SAV)—
<materialsCitation id="3B4D3CFEFFB3B114538AF867FD983F9A" ID-GBIF-Occurrence="1844420397" box="[241,634,1986,2011]" collectingDate="2012-05-19" collectorName="Type &amp; Turkey" country="Turkey" elevation="1430" latitude="39.716667" location="Geyikli Village" longLatPrecision="1167" longitude="39.666668" pageId="5" pageNumber="486" specimenCount="1" stateProvince="Erzincan Province" typeStatus="holotype">
with the same data as in
<typeStatus id="549E8801FFB3B114516FF867FD983F9A" box="[532,634,1986,2011]" pageId="5" pageNumber="486" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="3B4D3CFEFFB3B11751FCF866FD7538F1" ID-GBIF-Occurrence="1844420420" collectingDate="2017-05-30" collectionCode="SAV" collectorName="Type &amp; Turkey" country="Turkey" elevation="1430" lastPageId="6" lastPageNumber="487" latitude="39.716667" location="Geyikli Village" longLatPrecision="1167" longitude="39.666668" pageId="5" pageNumber="486" specimenCount="1" stateProvince="Erzincan Province" typeStatus="holotype">
two females—with the same data as in
<typeStatus id="549E8801FFB3B1145729F867FB553F9A" box="[1106,1207,1986,2011]" pageId="5" pageNumber="486" type="holotype">holotype</typeStatus>
,
<collectingDate id="EFDFE98BFFB3B11457BFF866FA813F9A" box="[1220,1379,1987,2011]" pageId="5" pageNumber="486" value="2017-05-30">
<date id="FF9B1063FFB3B11457BFF866FA813F9A" box="[1220,1379,1987,2011]" pageId="5" pageNumber="486" value="2017-05-30">30 May 2017</date>
</collectingDate>
; five females (of which one deposited in IZ
<collectionCode id="ED34AE66FFB0B1175122FF3DFD6D38F1" box="[601,655,152,176]" country="Slovakia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14906" name="Institute of Botany, Slovak Academy of Sciences" pageId="6" pageNumber="487">SAV</collectionCode>
)
</materialsCitation>
—
<materialsCitation id="3B4D3CFEFFB0B11751C8FF3DFE4038B9" ID-GBIF-Occurrence="1844420503" collectingDate="2017-05-25" collectorName="E Turkey &amp; The" country="Turkey" location="Acarology Laboratory of Erzincan University" pageId="6" pageNumber="487" specimenCount="2" stateProvince="Tunceli Province" typeStatus="holotype">
<collectorName id="26D05375FFB0B11751C8FF3DFCFC38F1" box="[691,798,151,176]" pageId="6" pageNumber="487">
E
<collectingCountry id="F3327633FFB0B11751B6FF32FCFC38F1" box="[717,798,151,176]" name="Turkey" pageId="6" pageNumber="487">Turkey</collectingCountry>
</collectorName>
,
<collectingRegion id="49E1F841FFB0B1175057FF32FC1338F1" box="[812,1009,151,176]" country="Turkey" name="Tunceli" pageId="6" pageNumber="487">Tunceli Province</collectingRegion>
, Pülümür District, Uzuntarla Village, leaf litter under
<taxonomicName id="4C254D20FFB0B1175230FF19FE4B3892" box="[331,425,188,211]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="487" phylum="Tracheophyta" rank="genus">
<emphasis id="B951EAB1FFB0B1175230FF19FE4B3892" box="[331,425,188,211]" italics="true" pageId="6" pageNumber="487">Quercus</emphasis>
</taxonomicName>
sp.,
<collectingDate id="EFDFE98BFFB0B11752A5FF19FD9B3895" box="[478,633,188,213]" pageId="6" pageNumber="487" value="2017-05-25">
<date id="FF9B1063FFB0B11752A5FF19FD9B3895" box="[478,633,188,213]" pageId="6" pageNumber="487" value="2017-05-25">25 May 2017</date>
</collectingDate>
.
<collectorName id="26D05375FFB0B11751F8FF19FD4D3894" box="[643,687,188,213]" pageId="6" pageNumber="487">The</collectorName>
<typeStatus id="549E8801FFB0B11751CEFF19FCF53894" box="[693,791,188,213]" pageId="6" pageNumber="487" type="holotype">holotype</typeStatus>
and
<typeStatus id="549E8801FFB0B1175035FF18FC5B3894" box="[846,953,189,213]" pageId="6" pageNumber="487" type="paratype">paratypes</typeStatus>
are deposited at the
<location id="8EFA6078FFB0B11757D9FF19FE4038B9" LSID="urn:lsid:plazi:treatment:038C87B5FFB3B11853ECF8AAFD3D3A8D:8EFA6078FFB0B11757D9FF19FE4038B9" country="Turkey" name="Acarology Laboratory of Erzincan University" pageId="6" pageNumber="487" stateProvince="Tunceli Province">Acarology Laboratory of Erzincan University</location>
</materialsCitation>
,
<materialsCitation id="3B4D3CFEFFB0B11752C9FF7AFD9A395D" ID-GBIF-Occurrence="1844420506" collectionCode="SAV" country="Turkey" county="Institute of Zoology" location="Bratislava" municipality="Slovak Academy of Sciences" pageId="6" pageNumber="487" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F3327633FFB0B11752C9FF7AFDE438B9" box="[434,518,223,248]" name="Turkey" pageId="6" pageNumber="487">Turkey</collectingCountry>
; two
<typeStatus id="549E8801FFB0B1175137FF45FD5538B9" box="[588,695,224,248]" pageId="6" pageNumber="487" type="paratype">paratypes</typeStatus>
are deposited at the
<collectingCounty id="62FB4E2FFFB0B11750CCFF7AFB4138B9" box="[951,1187,223,248]" pageId="6" pageNumber="487">Institute of Zoology</collectingCounty>
,
<collectingMunicipality id="6BFEACD9FFB0B11757C8FF7AFF1E395C" pageId="6" pageNumber="487">Slovak Academy of Sciences</collectingMunicipality>
,
<location id="8EFA6078FFB0B117527CFEA1FE9B395C" LSID="urn:lsid:plazi:treatment:038C87B5FFB3B11853ECF8AAFD3D3A8D:8EFA6078FFB0B117527CFEA1FE9B395C" box="[263,377,260,285]" country="Turkey" county="Institute of Zoology" municipality="Slovak Academy of Sciences" name="Bratislava" pageId="6" pageNumber="487">Bratislava</location>
(indicated as IZ
<collectionCode id="ED34AE66FFB0B117514DFEA1FD8D395D" box="[566,623,260,284]" country="Slovakia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14906" name="Institute of Botany, Slovak Academy of Sciences" pageId="6" pageNumber="487">SAV</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5A662BFFB0B11753ECFA4EFEFB3E60" httpUri="https://zenodo.org/record/1245030/files/figure.png" pageId="6" pageNumber="487" startId="6.[151,263,1515,1538]" targetBox="[162,1427,350,1485]" targetPageId="6">
<paragraph id="8B9A36A3FFB0B11753ECFA4EFEFB3E60" blockId="6.[151,1436,1515,1569]" pageId="6" pageNumber="487">
<emphasis id="B951EAB1FFB0B11753ECFA4EFEB73E40" bold="true" box="[151,341,1515,1538]" pageId="6" pageNumber="487">FIGURES 25–26.</emphasis>
<taxonomicName id="4C254D20FFB0B1175226FA49FD003E43" authority="Özbek &amp; Mašán, 2018" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[349,738,1516,1538]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="487" phylum="Arthropoda" rank="species" species="armiger" status="sp. nov." subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFB0B1175226FA49FE073E40" box="[349,485,1516,1537]" italics="true" pageId="6" pageNumber="487">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFB0B117528CFA49FD9D3E40" box="[503,639,1516,1537]" italics="true" pageId="6" pageNumber="487">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFB0B11751EBFA48FD003E43" box="[656,738,1517,1538]" italics="true" pageId="6" pageNumber="487">armiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257CAFFB0B1175191FA49FCDE3E40" box="[746,828,1516,1537]" pageId="6" pageNumber="487" rank="species">
<emphasis id="B951EAB1FFB0B1175191FA49FCDE3E40" bold="true" box="[746,828,1516,1537]" pageId="6" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 25. DOrsal idiOsOma; 26. Ventral idiOsOma. Scale bar: 200 µm.
</paragraph>
</caption>
<subSubSection id="C33F6528FFB0B11753BCF9EDFC4A3F54" pageId="6" pageNumber="487" type="diagnosis">
<paragraph id="8B9A36A3FFB0B11753BCF9EDFC4A3F54" blockId="6.[151,1437,1608,2029]" pageId="6" pageNumber="487">
<emphasis id="B951EAB1FFB0B11753BCF9EDFEA33E20" bold="true" box="[199,321,1608,1633]" pageId="6" pageNumber="487">Diagnosis.</emphasis>
Dorsal setae relatively short: dorsocentral j- and J-setae with their tips not exceeding bases of following seta. Openings of slit-like glandular poroids gdZ1 and gdS4 with separate position on posterolateral margins of dorsal shield. Prestigmatic section of peritremes long, with anterior tips reaching dorsal surface between setae z1 and z2. Soft integument with 15 pairs of setae. Cheliceral digits polydentate. Trochanter III with large posterolateral projection. Sperm induction system with tubular components well developed: tubes hyaline, flattened, progressively narrowed apically and usually curved distally.
</paragraph>
</subSubSection>
<subSubSection id="C33F6528FFB0B11853BCF885FAC738B9" lastPageId="9" lastPageNumber="490" pageId="6" pageNumber="487" type="description">
<paragraph id="8B9A36A3FFB0B11753BCF885FE5F3F76" blockId="6.[151,1437,1608,2029]" box="[199,445,1824,1849]" pageId="6" pageNumber="487">
<emphasis id="B951EAB1FFB0B11753BCF885FE5F3F76" bold="true" box="[199,445,1824,1849]" pageId="6" pageNumber="487">Description. Female.</emphasis>
</paragraph>
<paragraph id="8B9A36A3FFB0B11653BCF8E1FF5E38B9" blockId="6.[151,1437,1608,2029]" lastBlockId="7.[151,1436,151,248]" lastPageId="7" lastPageNumber="488" pageId="6" pageNumber="487">
<emphasis id="B951EAB1FFB0B11753BCF8E1FE603F1D" box="[199,386,1860,1884]" italics="true" pageId="6" pageNumber="487">Dorsal idiosoma</emphasis>
(
<figureCitation id="131E2A26FFB0B11752EBF8E1FE063F1C" box="[400,484,1860,1885]" captionStart="FIGURES 25–26" captionStartId="6.[151,263,1515,1538]" captionTargetBox="[162,1427,350,1485]" captionTargetId="figure@6.[151,1436,337,1494]" captionTargetPageId="6" captionText="FIGURES 25–26. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 25. DOrsal idiOsOma; 26. Ventral idiOsOma. Scale bar:200 µm." httpUri="https://zenodo.org/record/1245030/files/figure.png" pageId="6" pageNumber="487">Fig. 25</figureCitation>
). Dorsal shield 715‒815 µm long and 405‒455 µm wide, suboval and oblong (length/ width 1.7‒1.96), delicately and evenly reticulate on whole surface, and bearing 30 pairs of smooth and needleshaped dorsal setae. Setae J5 strongly reduced in length, vestigial microsetae; setae z1 short, 20‒26 µm long; other setae relatively well developed, similar in length and uniform. Length and spacing of some selected dorsal shield setae as follows: j1 45‒53 µm, j4‒j6, and z5 50‒60 µm, other setae 60‒80 µm, length of j5 similar or shorter than distance between j5-z5, and length of J1 similar or shorter than distance between J1-J2. Two posterolateral poroid structures (gdZ1 and gdS4) with their openings slit-like, separate and placed approximately between setae Z2 and S4.
</paragraph>
<caption id="DF5A662BFFB1B11653ECFB79FA793D50" httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="7" pageNumber="488" startId="7.[151,263,1244,1267]" targetBox="[158,1430,309,1215]" targetPageId="7">
<paragraph id="8B9A36A3FFB1B11653ECFB79FA793D50" blockId="7.[151,1435,1244,1297]" pageId="7" pageNumber="488">
<emphasis id="B951EAB1FFB1B11653ECFB79FEBA3CB0" bold="true" box="[151,344,1244,1267]" pageId="7" pageNumber="488">FIGURES 27–33.</emphasis>
<taxonomicName id="4C254D20FFB1B1165219FB79FD093CB3" authority="Özbek &amp; Mašán, 2018" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[354,747,1244,1266]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="488" phylum="Arthropoda" rank="species" species="armiger" status="sp. nov." subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFB1B1165219FB79FE083CB0" box="[354,490,1244,1265]" italics="true" pageId="7" pageNumber="488">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFB1B1165285FB79FD643CB0" box="[510,646,1244,1265]" italics="true" pageId="7" pageNumber="488">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFB1B11651E2FB78FD093CB3" box="[665,747,1245,1266]" italics="true" pageId="7" pageNumber="488">armiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257CAFFB1B116518DFB78FCA93CB3" box="[758,843,1245,1266]" pageId="7" pageNumber="488" rank="species">
<emphasis id="B951EAB1FFB1B116518DFB78FCA93CB3" bold="true" box="[758,843,1245,1266]" pageId="7" pageNumber="488">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms; 29. Ventral gnathOsOma; 30. EpistOme; 31. TrOchanter III; 32. Tarsus II; 33. Ventral spur On femur II. Scale bar: 100 µm.
</paragraph>
</caption>
<paragraph id="8B9A36A3FFB1B11653BCFA9BFAF13EDB" blockId="7.[151,1437,1341,2014]" pageId="7" pageNumber="488">
<emphasis id="B951EAB1FFB1B11653BCFA9BFE673D17" box="[199,389,1342,1366]" italics="true" pageId="7" pageNumber="488">Ventral idiosoma</emphasis>
(
<figureCitation id="131E2A26FFB1B11652EFFA98FE053D17" box="[404,487,1341,1366]" captionStart="FIGURES 25–26" captionStartId="6.[151,263,1515,1538]" captionTargetBox="[162,1427,350,1485]" captionTargetId="figure@6.[151,1436,337,1494]" captionTargetPageId="6" captionText="FIGURES 25–26. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 25. DOrsal idiOsOma; 26. Ventral idiOsOma. Scale bar:200 µm." httpUri="https://zenodo.org/record/1245030/files/figure.png" pageId="7" pageNumber="488">Fig. 26</figureCitation>
). Sternal shield 277‒290 µm long, proportionally 1.02‒1.15 longer than genitiventral shield, with slightly concave anterior margin and two small projections close to bases of sternal setae st1. Genitiventral shield slightly longer than wide or subequal in length and width (length 235‒278 µm, width 230‒277 µm, length/width 1‒1.07). Anal shield subtriangular, 111‒128 µm long and 138‒172 µm wide (length/width 0.65‒0.80); anus with three circum-anal setae situated close to posterior margin of shield. Peritremes normally developed, relatively long, with anterior end reaching dorsal surface between setae z1 and z2. Peritrematal shields with weak longitudinal sculptural lines, other ventral shields distinctly and evenly reticulate and usually delicately punctate on surface. Metapodal platelets minuscule, free and well separate from peritrematal shields. Lateral soft integument with seven pairs of marginal setae (r6, R1, R3‒R7). Ventral soft integument with eight pairs of opisthogastric setae (JV2‒JV5, ZV2‒ZV5). Ventrally placed setae similar to those on dorsal idiosoma.
</paragraph>
<paragraph id="8B9A36A3FFB1B11653BCF903FF0E3F6B" blockId="7.[151,1437,1341,2014]" pageId="7" pageNumber="488">
<emphasis id="B951EAB1FFB1B11653BCF903FE3C3EFF" box="[199,478,1702,1726]" italics="true" pageId="7" pageNumber="488">Spermathecal apparatus</emphasis>
(
<figureCitation id="131E2A26FFB1B116528BF900FDA83EFF" box="[496,586,1701,1726]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="7" pageNumber="488">Figs 28</figureCitation>
,
<figureCitation id="131E2A26FFB1B1165121F903FD453EFF" box="[602,679,1702,1726]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="7" pageNumber="488">37, 38</figureCitation>
). Tubes of sperm induction system relatively well developed but weakly sclerotised, hyaline, flattened, with broader base and narrowed distal part; basal section widely abutting to inner anterolateral margin of coxa IV; distal part usually curved and irregularly formed, sometimes straightened inward.
</paragraph>
<paragraph id="8B9A36A3FFB1B11953BCF893FDD43901" blockId="7.[151,1437,1341,2014]" lastBlockId="8.[151,1436,151,320]" lastPageId="8" lastPageNumber="489" pageId="7" pageNumber="488">
<emphasis id="B951EAB1FFB1B11653BCF893FE373F0F" box="[199,469,1846,1870]" italics="true" pageId="7" pageNumber="488">Gnathosomal structures</emphasis>
(
<figureCitation id="131E2A26FFB1B116529DF890FDDC3F0F" box="[486,574,1845,1870]" captionStart="FIGURES 15–24" captionStartId="4.[151,263,1909,1932]" captionTargetBox="[178,1406,195,1885]" captionTargetId="figure@4.[178,1406,195,1885]" captionTargetPageId="4" captionText="FIGURES 15–24. Pachylaelaps spp. 15‒23. Pachylaelaps (Longipachys) anatolicus. 15. Chelicera, male, variant fOrms; 16. Chelicera, female; 17. Terminal parts Of palps (tarsi and tibiae), male; 18. Leg II (tarsus, tibia and genu), female; 19. Leg II (tarsus, tibia, genu and femur), male; 20. Spermathecal duct, female; 21‒22. EpistOme, female, variant fOrms; 23. EpistOme, male. 24. Pachylaelaps (Pachylaelaps) armiger sp. nov., chelicera, female.NOt scaled." httpUri="https://zenodo.org/record/1245028/files/figure.png" pageId="7" pageNumber="488">Figs 24</figureCitation>
,
<figureCitation id="131E2A26FFB1B1165137F893FD233F0F" box="[588,705,1846,1870]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="7" pageNumber="488">27, 29, 30</figureCitation>
,
<figureCitation id="131E2A26FFB1B11651B4F893FD0D3F0F" box="[719,751,1846,1870]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="7" pageNumber="488">34</figureCitation>
). Corniculi elongated and horn-like; laciniae densely pilose, lanceolate, longer than corniculi; deutosternum slightly widened anteriorly, with six transverse rows of denticles; subcapitular setae smooth and needle-shaped: setae h1 longest, h2 similar in length to pc, and shorter than h3 (
<figureCitation id="131E2A26FFB1B1165615F8D8FF553FFB" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="7" pageNumber="488">Fig. 29</figureCitation>
). Epistome with wide, subtriangular and marginally serrate base, and central projection; projection long and narrow, smooth, with inconspicuous neck and bifurcate brush-like apex; each furca delicately serrate apically, with a row of 1‒5 minute prongs (
<figureCitation id="131E2A26FFBEB1195293FF32FDDC38F1" box="[488,574,151,176]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="8" pageNumber="489">Fig. 30</figureCitation>
). Chelicerae pincer-like, with relatively short and robust polydentate digits; fixed digit 94‒100 µm long, having bidentate terminal hook, one robust distal tooth associated with minute pilus dentilis, and 10‒15 small and sharp teeth on masticatory area; movable digit 92‒105 µm long, 42‒48% of the length of medial segment (210‒220 µm), armed with simple hook and 7‒9 sharp teeth, including the biggest one with distal position (
<figureCitation id="131E2A26FFBEB1195207FE82FE313901" box="[380,467,295,320]" captionStart="FIGURES 15–24" captionStartId="4.[151,263,1909,1932]" captionTargetBox="[178,1406,195,1885]" captionTargetId="figure@4.[178,1406,195,1885]" captionTargetPageId="4" captionText="FIGURES 15–24. Pachylaelaps spp. 15‒23. Pachylaelaps (Longipachys) anatolicus. 15. Chelicera, male, variant fOrms; 16. Chelicera, female; 17. Terminal parts Of palps (tarsi and tibiae), male; 18. Leg II (tarsus, tibia and genu), female; 19. Leg II (tarsus, tibia, genu and femur), male; 20. Spermathecal duct, female; 21‒22. EpistOme, female, variant fOrms; 23. EpistOme, male. 24. Pachylaelaps (Pachylaelaps) armiger sp. nov., chelicera, female.NOt scaled." httpUri="https://zenodo.org/record/1245028/files/figure.png" pageId="8" pageNumber="489">Figs 24</figureCitation>
,
<figureCitation id="131E2A26FFBEB11952A4FE8DFE1F3901" box="[479,509,296,320]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="8" pageNumber="489">27</figureCitation>
,
<figureCitation id="131E2A26FFBEB1195172FE8DFDCB3901" box="[521,553,296,320]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="8" pageNumber="489">34</figureCitation>
).
</paragraph>
<caption id="DF5A662BFFBEB11953ECF829FC663FA1" httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="8" pageNumber="489" startId="8.[151,263,1932,1955]" targetBox="[151,1436,373,1911]" targetPageId="8">
<paragraph id="8B9A36A3FFBEB11953ECF829FC663FA1" blockId="8.[151,1436,1932,2016]" pageId="8" pageNumber="489">
<emphasis id="B951EAB1FFBEB11953ECF829FEBA3FE0" bold="true" box="[151,344,1932,1955]" pageId="8" pageNumber="489">FIGURES 34–39.</emphasis>
<taxonomicName id="4C254D20FFBEB1195218F829FD723FE3" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[355,656,1932,1954]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB1FFBEB1195218F829FE093FE0" box="[355,491,1932,1953]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFBEB1195284F829FD653FE0" box="[511,647,1932,1953]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
)
</taxonomicName>
spp., females. 34‒38.
<taxonomicName id="4C254D20FFBEB11950FCF829FAF33FE3" authority="Özbek &amp; Mašán, 2018" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[903,1297,1932,1954]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="489" phylum="Arthropoda" rank="species" species="armiger" status="sp. nov." subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFBEB11950FCF829FBED3FE0" box="[903,1039,1932,1953]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFBEB1195758F829FB493FE0" box="[1059,1195,1932,1953]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFBEB11957C4F828FAF33FE3" box="[1215,1297,1933,1954]" italics="true" pageId="8" pageNumber="489">armiger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257CAFFBEB1195660F828FA933FE3" box="[1307,1393,1933,1954]" pageId="8" pageNumber="489" rank="species">
<emphasis id="B951EAB1FFBEB1195660F828FA933FE3" bold="true" box="[1307,1393,1933,1954]" pageId="8" pageNumber="489">sp. nov.</emphasis>
</taxonomicNameLabel>
34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39.
<taxonomicName id="4C254D20FFBEB11953ECF86FFDE43FA1" authorityName="Costa" authorityYear="1971" box="[151,518,1994,2016]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="489" phylum="Arthropoda" rank="species" species="evansi" subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFBEB11953ECF86FFEFD3F9E" box="[151,287,1994,2015]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFBEB1195255F86FFE543F9E" box="[302,438,1994,2015]" italics="true" pageId="8" pageNumber="489">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFBEB11952BEF86EFDE43FA1" box="[453,518,1995,2016]" italics="true" pageId="8" pageNumber="489">evansi</emphasis>
</taxonomicName>
, part Of ventral idiOsOma. NOt scaled.
</paragraph>
</caption>
<paragraph id="8B9A36A3FFBFB11853BCFF3CFAC738B9" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="490">
<emphasis id="B951EAB1FFBFB11853BCFF3CFF1E38F1" box="[199,252,153,176]" italics="true" pageId="9" pageNumber="490">Legs</emphasis>
(
<figureCitation id="131E2A26FFBFB1185276FF32FE7238F1" box="[269,400,151,176]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="9" pageNumber="490">Figs 31‒33</figureCitation>
,
<figureCitation id="131E2A26FFBFB11852E5FF32FE0538F1" box="[414,487,151,176]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="9" pageNumber="490">35, 36</figureCitation>
). Leg setation normal for genus (
<bibRefCitation id="EFB44B52FFBFB1185012FF3DFBE138F1" box="[873,1027,152,176]" pageId="9" pageNumber="490" refId="ref5383" refString="Masan, P. (2007) A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea). NOI Press, Bratislava, 247 pp." type="book">Mašán, 2007</bibRefCitation>
). Tarsus II with two spur-like distal setae, pl1 and pl2 (
<figureCitation id="131E2A26FFBFB1185212FF19FE223895" box="[361,448,188,213]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="9" pageNumber="490">Figs 32</figureCitation>
,
<figureCitation id="131E2A26FFBFB11852B7FF19FE093894" box="[460,491,188,213]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="9" pageNumber="490">35</figureCitation>
). Femur II with small spine-like process and associated seta on distal ventral surface (
<figureCitation id="131E2A26FFBFB11853E4FF7AFF1738B9" box="[159,245,223,248]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="9" pageNumber="490">Figs 33</figureCitation>
,
<figureCitation id="131E2A26FFBFB118527AFF7AFEC338B9" box="[257,289,223,248]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="9" pageNumber="490">35</figureCitation>
). Trochanter III with conspicuous bump-shaped projection directed backward (
<figureCitation id="131E2A26FFBFB11857EDFF7AFB0F38B9" box="[1174,1261,223,248]" captionStart="FIGURES 27–33" captionStartId="7.[151,263,1244,1267]" captionTargetBox="[158,1430,309,1215]" captionTargetId="figure@7.[151,1436,301,1223]" captionTargetPageId="7" captionText="FIGURES 27–33. Pachylaelaps (Pachylaelaps) armiger sp. nov., female. 27. Chelicera; 28. Spermathecal ducts, variant fOrms;29.Ventral gnathOsOma;30. EpistOme;31. TrOchanter III; 32.Tarsus II; 33.Ventral spur On femur II. Scale bar:100 µm." httpUri="https://zenodo.org/record/1245032/files/figure.png" pageId="9" pageNumber="490">Figs 31</figureCitation>
,
<figureCitation id="131E2A26FFBFB1185782FF7AFAFB38B9" box="[1273,1305,223,248]" captionStart="FIGURES 34–39" captionStartId="8.[151,263,1932,1955]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@8.[151,1436,373,1911]" captionTargetPageId="8" captionText="FIGURES 34–39. Pachylaelaps (Pachylaelaps) spp., females. 34‒38. Pachylaelaps (Pachylaelaps) armiger sp. nov. 34. Chelicera; 35. Leg II (femur, genu, tibia and tarsus); 36. TrOchanter III; 37‒38. Spermathecal ducts, variant fOrms. 39. Pachylaelaps (Pachylaelaps) evansi, part Of ventral idiOsOma. NOt scaled." httpUri="https://zenodo.org/record/1245034/files/figure.png" pageId="9" pageNumber="490">36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6528FFBFB11853BCFEA1FD3D3A8D" pageId="9" pageNumber="490" type="etymology">
<paragraph id="8B9A36A3FFBFB11853BCFEA1FEFB3924" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="490">
<emphasis id="B951EAB1FFBFB11853BCFEA1FEAC395C" bold="true" box="[199,334,260,285]" pageId="9" pageNumber="490">Etymology.</emphasis>
The specific name of the new species is derived from the Latin words &quot;
<emphasis id="B951EAB1FFBFB118570CFEA0FB50395D" box="[1143,1202,261,284]" italics="true" pageId="9" pageNumber="490">arma</emphasis>
&quot; (instrument of war, arm) and &quot;
<emphasis id="B951EAB1FFBFB1185274FE8CFEA03901" box="[271,322,297,320]" italics="true" pageId="9" pageNumber="490">gerō</emphasis>
&quot; (carry or bear), and it alludes to the fact that the chelicerae of these mites are heavily armed with sharp teeth.
</paragraph>
<paragraph id="8B9A36A3FFBFB11853BCFECAFD3D3A8D" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="490">
<emphasis id="B951EAB1FFBFB11853BCFECAFE7E39C9" bold="true" box="[199,412,367,392]" pageId="9" pageNumber="490">Taxonomic notes.</emphasis>
<bibRefCitation id="EFB44B52FFBFB11852D3FED5FDAB39C9" box="[424,585,368,392]" pageId="9" pageNumber="490" refId="ref5383" refString="Masan, P. (2007) A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea). NOI Press, Bratislava, 247 pp." type="book">Mašán (2007)</bibRefCitation>
divided the European members of the subgenus
<taxonomicName id="4C254D20FFBFB11857EEFED5FACE39C9" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[1173,1324,368,392]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAB1FFBFB11857EEFED5FACE39C9" box="[1173,1324,368,392]" italics="true" pageId="9" pageNumber="490">Pachylaelaps</emphasis>
</taxonomicName>
into five clusters of species. In his classification scheme,
<taxonomicName id="4C254D20FFBFB11851B4FE31FB9D39ED" authorityName="Özbek &amp; Mašán" authorityYear="2018" box="[719,1151,404,428]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="armiger" subGenus="Pachylaelaps">
<emphasis id="B951EAB1FFBFB11851B4FE31FC8439ED" box="[719,870,404,428]" italics="true" pageId="9" pageNumber="490">Pachylaelaps</emphasis>
(
<emphasis id="B951EAB1FFBFB1185001FE31FBF339ED" box="[890,1041,404,428]" italics="true" pageId="9" pageNumber="490">Pachylaelaps</emphasis>
)
<emphasis id="B951EAB1FFBFB118575EFE30FB9D39ED" box="[1061,1151,405,428]" italics="true" pageId="9" pageNumber="490">armiger</emphasis>
</taxonomicName>
could be classified as a species of the
<taxonomicName id="4C254D20FFBFB118524DFE1DFE443991" authorityName="Berlese" authorityYear="1920" box="[310,422,440,464]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="B951EAB1FFBFB118524DFE1DFE443991" box="[310,422,440,464]" italics="true" pageId="9" pageNumber="490">bellicosus</emphasis>
</taxonomicName>
group, which is representated by
<taxonomicName id="4C254D20FFBFB118505BFE1DFB2D3991" authority="Berlese, 1920" authorityName="Berlese" authorityYear="1920" box="[800,1231,439,464]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="B951EAB1FFBFB118505BFE1DFBCC3991" box="[800,1070,440,464]" italics="true" pageId="9" pageNumber="490">Pachylaelaps bellicosus</emphasis>
Berlese, 1920
</taxonomicName>
and
<taxonomicName id="4C254D20FFBFB118567EFE1DFDD539B4" authority="Evans &amp; Hyatt, 1956" authorityName="Evans &amp; Hyatt" authorityYear="1956" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="multidentatus">
<emphasis id="B951EAB1FFBFB118567EFE1DFED239B5" italics="true" pageId="9" pageNumber="490">Pachylaelaps multidentatus</emphasis>
<bibRefCitation id="EFB44B52FFBFB1185240FE78FDD539B4" box="[315,567,476,501]" pageId="9" pageNumber="490" refId="ref5109" refString="Evans, G. O. &amp; HYatt, K. H. (1956) British mites Of the genus Pachylaelaps Berlese (Gamasina - PachYlaelaptidae). Entomologist's Monthly Magazine, 92, 118 - 129." type="journal article">Evans &amp; Hyatt, 1956</bibRefCitation>
</taxonomicName>
. The
<taxonomicName id="4C254D20FFBFB1185106FE79FD0F39B5" authorityName="Berlese" authorityYear="1920" box="[637,749,476,500]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="B951EAB1FFBFB1185106FE79FD0F39B5" box="[637,749,476,500]" italics="true" pageId="9" pageNumber="490">bellicosus</emphasis>
</taxonomicName>
group is characterised by the separate position of slit-like poroid structures gdZ1 and gdS4 on the dorsal shield, multidentate cheliceral digits, weakly sclerotised and hyaline tubiform structures of sperm induction system, and males apparently absent. The new species can be easily distinguished from both above mentioned congeners especially by the presence of large bump-like projection on posterior surface of trochanter III. This conspicuous structure is easily detectable and not found in its relatives in the species group, including
<taxonomicName id="4C254D20FFBFB118529FFD35FBC13AE9" authority="Moraza &amp; Pena, 2005" authorityName="Moraza &amp; Pena" authorityYear="2005" box="[484,1059,655,680]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B951EAB1FFBFB118529FFD35FCF83AE9" box="[484,794,656,680]" italics="true" pageId="9" pageNumber="490">Pachylaelaps mandibularis</emphasis>
Moraza &amp; Peña, 2005
</taxonomicName>
previously synonymised with
<taxonomicName id="4C254D20FFBFB11856FCFD34FEE53A8D" authorityName="Berlese" authorityYear="1920" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bellicosus">
<emphasis id="B951EAB1FFBFB11856FCFD34FEE53A8D" italics="true" pageId="9" pageNumber="490">P. bellicosus</emphasis>
</taxonomicName>
by Mašán in 2007 (Moraza, pers. comm).
</paragraph>
</subSubSection>
</treatment>
</document>
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