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<document id="D69CA3C26782D25C3B049B0D8BCAC120" ID-DOI="10.11646/zootaxa.5376.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10208788" ID-ZooBank="9ECCB3F7-5481-47C2-8A5A-E9A3F38C31BA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701073812311" checkinUser="plazi" docAuthor="Angulo, Arturo, Betts, Joel T., González-Alemán, Néstor J., Castañeda, Edgar, Berghe, Eric Van Den, Elías, Diego J., Mcmahan, Caleb D. &amp; Matamoros, Wilfredo A." docDate="2023" docId="038C8788E870FF86FF792FEFFC873212" docLanguage="en" docName="zootaxa.5376.1.1.pdf" docOrigin="Zootaxa 5376 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5376.1.1/52351" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Herotilapia multispinosa" docType="treatment" docVersion="1" lastPageNumber="52" masterDocId="FFB5FFF0E843FFB5FFEE2C1FFFA1347C" masterDocTitle="Continental fishes of Nicaragua: diversity, distribution and conservation status; with an annotated and illustrated checklist of species and an identification guide to families" masterLastPageNumber="89" masterPageNumber="1" pageNumber="52" updateTime="1701194607627" updateUser="felipe">
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<mods:title id="BEDB54F05F4310876466D0926466F6F7">Continental fishes of Nicaragua: diversity, distribution and conservation status; with an annotated and illustrated checklist of species and an identification guide to families</mods:title>
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<mods:namePart id="660279532C9DA3DEB147872A83C8155A">Betts, Joel T.</mods:namePart>
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<mods:affiliation id="64F10CCDEA2DBF819AAD7467EE62D60F">Department of Fisheries and Wildlife, Michigan State University, 480 Wilson Rd # 13, East Lansing, MI 48824, USA joel. t. betts @ gmail. com; https: // orcid. org / 0000 - 0002 - 6018 - 8995</mods:affiliation>
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<mods:affiliation id="C6CA0F01FBFF60D8077CE63BF9E6604C">Universidad Americana (UAM, Costado Noroeste Camino de Oriente, Managua, Nicaragua edgar. ecologica @ gmail. com; https: // orcid. org / 0000 - 0001 - 8148 - 3195</mods:affiliation>
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<mods:affiliation id="3C820B2D36F92535AEC16307E1A6F694">Department of Environment and Development, Zamorano Biodiversity Center. Zamorano University of Tegucigalpa, P. O. Box 93, Honduras evandenberghe @ zamorano. edu; https: // orcid. org / 0000 - 0001 - 7566 - 0415</mods:affiliation>
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<mods:affiliation id="25EEDFC7A982FFCB92D0A7EF1D30B9C9">Field Museum of Natural History (FMNH). Chicago, Illinois, United States of America delias @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 4215 - 0384 cmcmahan @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2113 - 8554</mods:affiliation>
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<taxonomicName id="4C254D1DE870FF86FF792FEFFDC73076" ID-CoL="3L2V5" authority="(Gunther 1867)" baseAuthorityName="Gunther" baseAuthorityYear="1867" box="[151,614,1008,1034]" family="Cichlidae" genus="Herotilapia" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="multispinosa">
<emphasis id="B951EA8CE870FF86FF792FEFFE0E3076" box="[151,431,1008,1034]" italics="true" pageId="51" pageNumber="52">Herotilapia multispinosa</emphasis>
(Günther 1867)
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. Rainbow Cichlid; Mojarra, Mojarra Arcoiris, Cholesca, Tilapia
</paragraph>
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<paragraph id="8B9A369EE870FF86FF79280BFC873212" blockId="51.[151,1437,1008,1647]" pageId="51" pageNumber="52">
<emphasis id="B951EA8CE870FF86FF79280BFEB13052" bold="true" box="[151,272,1044,1070]" pageId="51" pageNumber="52">Vouchers:</emphasis>
UCR 0447-003 (5), 0478-001 (3), 0536-006 (3), 0546-005 (14), 0549-004 (7), 1051-007 (2), 1052-005 (0), 1054-006 (8), 1055-013 (2) and 3115-006 (1); FMNH 6001 (1), 6002 (1), 6003 (39), 6004 (11) and 52525 (3); LSUMZ 15364 (26), 15445 (1), 15454 (4), 15461 (4) and 15471 (2); other collections: CAS 28489 (140), 61376 (15), 61381 (3), 76029 (1), IBUNAM 13739 (1), MNHN 2001-1173 (1), UMMZ 180621 (1) and 213939 (1).
<emphasis id="B951EA8CE870FF86FF7928BBFE9230C2" bold="true" box="[151,307,1188,1214]" pageId="51" pageNumber="52">Distribution:</emphasis>
Central America; from eastern Honduras to northeastern Costa Rica, Atlantic and Pacific drainages;
<emphasis id="B951EA8CE870FF86FF7928D7FE86309E" bold="true" box="[151,295,1224,1250]" pageId="51" pageNumber="52">
(
<collectingCountry id="F332760EE870FF86FF7128D7FEBE309E" box="[159,287,1224,1250]" name="Nicaragua" pageId="51" pageNumber="52">Nicaragua</collectingCountry>
)
</emphasis>
WK, Es, BF, IM, To, Sa, SJ, RF, LN (Atlantic) and Ne (Pacific);
<quantity id="4CDD9B7BE870FF86FC1B28D7FBCA309E" box="[1013,1131,1224,1251]" metricMagnitude="1" metricUnit="m" metricValue="3.85" metricValueMax="7.5" metricValueMin="0.2" pageId="51" pageNumber="52" unit="m" value="38.5" valueMax="75.0" valueMin="2.0">275 masl</quantity>
; Sec, Pot.
<emphasis id="B951EA8CE870FF86FB3028D7FE27317A" bold="true" pageId="51" pageNumber="52">Occurrence and conservation status:</emphasis>
Nat; LC (2020), population trend unknown.
<emphasis id="B951EA8CE870FF86FC6E28F3FBA4317A" bold="true" box="[896,1029,1260,1286]" pageId="51" pageNumber="52">Literature:</emphasis>
<bibRefCitation id="EFB44B6FE870FF86FBE228F3FB70317A" author="Meek" box="[1036,1233,1260,1286]" firstAuthor="Meek" page="131" pageId="51" pageNumber="52" pagination="97 - 132" refId="ref84723" refString="Meek, S. E. (1907) Synopsis of the fishes of the Great Lakes of Nicaragua. Field Columbian Museum Zoological Series, 7, 97 - 132. https: // doi. org / 10.5962 / bhl. title. 3761" type="journal article" year="1907">Meek (1907: 131</bibRefCitation>
; brief description, including an identification key, and information on distribution and ecology), Astorqui (1975: 144; brief description, including illustrations and an identification key, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE870FF86FB36292BFA363132" author="Villa" box="[1240,1431,1332,1358]" firstAuthor="Villa" page="202" pageId="51" pageNumber="52" refId="ref86666" refString="Villa, J. (1982) Peces nicaraguenses de agua dulce. Editorial Union Cardoza y Cia. Ltda, Managua, 253 pp." type="book" year="1982">Villa (1982: 202</bibRefCitation>
; detailed description, including illustrations and an identification key, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE870FF86FF792963FE2531EB" author="Bussing" box="[151,388,1404,1431]" firstAuthor="Bussing" page="351" pageId="51" pageNumber="52" refId="ref81779" refString="Bussing, W. A. (1998). Freshwater fishes of Costa Rica. Editorial Universidad de Costa Rica, San Jose´, Costa Rica, 468 pp." type="book" year="1998">Bussing (1998: 351</bibRefCitation>
; brief description, including illustrations and an identification key, and information on distribution, with a map, and ecology), González-Alemán (2006: 59; brief description, including an illustration, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE870FF86FD7929DBFC1231A2" author="Schmitter-Soto" box="[663,947,1476,1502]" firstAuthor="Schmitter-Soto" page="30" pageId="51" pageNumber="52" pagination="1 - 76" refId="ref85937" refString="Schmitter-Soto, J. J. (2007) A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species. Zootaxa, 1603 (1), 1, 1 - 76. https: // doi. org / 10.11646 / zootaxa. 1603.1.1" type="journal article" year="2007">Schmitter-Soto (2007: 30</bibRefCitation>
, as
<taxonomicName id="4C254D1DE870FF86FC3229DBFAB331A2" ID-CoL="G9WL" baseAuthorityName="Gunther" baseAuthorityYear="1867" box="[988,1298,1476,1502]" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="multispinosus">
<emphasis id="B951EA8CE870FF86FC3229DBFAB331A2" box="[988,1298,1476,1502]" italics="true" pageId="51" pageNumber="52">Archocentrus multispinosus</emphasis>
</taxonomicName>
—synonym; detailed description, including illustrations and an identification key, and information on distribution, with a map, and ecology),
<bibRefCitation id="EFB44B6FE870FF86FEAE2A12FDD8325A" author="van den Berghe" box="[320,633,1548,1575]" firstAuthor="van den Berghe" page="32" pageId="51" pageNumber="52" pagination="1 - 55" refId="ref86561" refString="van den Berghe, E. (2015) Peces de la cuenca del ri´o Punta Gorda, vertiente del Caribe de Nicaragua. Revista Nicaraguense de Biodiversidad, 3, 1 - 55. [http: // www. bio-nica. info / RevNicaBiodiv / 03 - VandenBerghe-PecesPuntaGorda-final. pdf]" type="journal article" year="2015">van den Berghe (2015: 32</bibRefCitation>
; listed, including an illustration and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE870FF86FEE42A2FFE7D3236" author="Angulo, A." box="[266,476,1584,1610]" pageId="51" pageNumber="52" pagination="1 - 72" refId="ref80295" refString="Angulo, A. (2021) New records and range extensions to the Costa Rican freshwater fish fauna, with an updated checklist. Zootaxa, 5083 (1), 1 - 72, https: // doi. org / 10.11646 / zootaxa. 5083.1.1" type="journal article" year="2021">Angulo (2021: 49</bibRefCitation>
; listed, including information on distribution) and
<bibRefCitation id="EFB44B6FE870FF86FBC32A2FFA383236" author="Angulo &amp; San Gil-Leon" box="[1069,1433,1584,1610]" firstAuthor="Angulo" pageId="51" pageNumber="52" pagination="4029" refId="ref80408" refString="Angulo, A. &amp; San Gil-Leon, J. (2022) Claves para la identificacion de los peces de las aguas continentales e insulares de Costa Rica. Parte II: Cichliformes, Cichlidae (Mojarras, Guapotes, Tilapias y afines). 1740280795 UNED Research Journal, 14 (2), e 4029. https: // doi. org / 10.22458 / urj. v 14 i 2.4029" type="journal article" year="2022">Angulo &amp; San Gil-León (2022</bibRefCitation>
: e4029; identification key and information on distribution).
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