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<document id="591C402D9E85B66BB2CF0CDBC1718A70" ID-DOI="10.11646/zootaxa.5376.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10208788" ID-ZooBank="9ECCB3F7-5481-47C2-8A5A-E9A3F38C31BA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701073812311" checkinUser="plazi" docAuthor="Angulo, Arturo, Betts, Joel T., González-Alemán, Néstor J., Castañeda, Edgar, Berghe, Eric Van Den, Elías, Diego J., Mcmahan, Caleb D. &amp; Matamoros, Wilfredo A." docDate="2023" docId="038C8788E86CFF85FF792A28FC453665" docLanguage="en" docName="zootaxa.5376.1.1.pdf" docOrigin="Zootaxa 5376 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5376.1.1/52351" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Amphilophus citrinellus" docType="treatment" docVersion="1" lastPageNumber="48" masterDocId="FFB5FFF0E843FFB5FFEE2C1FFFA1347C" masterDocTitle="Continental fishes of Nicaragua: diversity, distribution and conservation status; with an annotated and illustrated checklist of species and an identification guide to families" masterLastPageNumber="89" masterPageNumber="1" pageNumber="48" updateTime="1701194607627" updateUser="felipe">
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<mods:title id="ED690F31D297BFEA9B5AAF448F9DF75D">Continental fishes of Nicaragua: diversity, distribution and conservation status; with an annotated and illustrated checklist of species and an identification guide to families</mods:title>
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<mods:namePart id="F667B7641C414CBE83139A723995A1F5">Angulo, Arturo</mods:namePart>
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<mods:namePart id="DFD5DB4A1AECB5896CF2609556B16C65">Betts, Joel T.</mods:namePart>
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<mods:affiliation id="702E1EF6E1C95FB8DD292061863853CE">Department of Fisheries and Wildlife, Michigan State University, 480 Wilson Rd # 13, East Lansing, MI 48824, USA joel. t. betts @ gmail. com; https: // orcid. org / 0000 - 0002 - 6018 - 8995</mods:affiliation>
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<mods:affiliation id="0649C52BB74148B408AB00FAEFAB2021">Universidad Americana (UAM, Costado Noroeste Camino de Oriente, Managua, Nicaragua edgar. ecologica @ gmail. com; https: // orcid. org / 0000 - 0001 - 8148 - 3195</mods:affiliation>
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<mods:affiliation id="84BC573FBE429A5742F885DF39B3A0B5">Department of Environment and Development, Zamorano Biodiversity Center. Zamorano University of Tegucigalpa, P. O. Box 93, Honduras evandenberghe @ zamorano. edu; https: // orcid. org / 0000 - 0001 - 7566 - 0415</mods:affiliation>
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<mods:namePart id="6A74D6BDD202A7435C2BB7E233ACB141">Elías, Diego J.</mods:namePart>
<mods:affiliation id="8AE9FF3D9E130CCA707C6F5874A40777">Field Museum of Natural History (FMNH). Chicago, Illinois, United States of America delias @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 4215 - 0384 cmcmahan @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2113 - 8554</mods:affiliation>
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<mods:affiliation id="4A3B8CCFCF4CF1FDE5B809FF46B37D96">Field Museum of Natural History (FMNH). Chicago, Illinois, United States of America delias @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 4215 - 0384 cmcmahan @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2113 - 8554</mods:affiliation>
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<mods:date id="531D59EC710A60A1CCAC1EDB40B9EEB8">2023</mods:date>
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<subSubSection id="C33F6515E86CFF9AFF792A28FB70322D" box="[151,1233,1591,1617]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8B9A369EE86CFF9AFF792A28FB70322D" blockId="47.[151,1437,1591,2013]" box="[151,1233,1591,1617]" pageId="47" pageNumber="48">
<heading id="D0D281F2E86CFF9AFF792A28FB70322D" box="[151,1233,1591,1617]" fontSize="11" level="2" pageId="47" pageNumber="48" reason="5">
<taxonomicName id="4C254D1DE86CFF9AFF792A28FDFA322D" ID-CoL="D3GS" authority="(Gunther 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[151,603,1591,1617]" family="Cichlidae" genus="Amphilophus" kingdom="Animalia" order="Perciformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="citrinellus">
<emphasis id="B951EA8CE86CFF9AFF792A28FE05322D" box="[151,420,1591,1617]" italics="true" pageId="47" pageNumber="48">Amphilophus citrinellus</emphasis>
(Günther 1864)
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. Midas Cichlid; Mojarra, Mojarra Midas, Cíclido Midas
</heading>
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<paragraph id="8B9A369EE86CFF85FF792A44FC453665" blockId="47.[151,1437,1591,2013]" lastBlockId="48.[151,1437,150,537]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B951EA8CE86CFF9AFF792A44FEB13209" bold="true" box="[151,272,1627,1653]" pageId="47" pageNumber="48">Vouchers:</emphasis>
UCR 0011-004 (4), 0024-015 (21), 0049-002 (3), 0050-002 (108), 0061-006 (7), 0062-004 (10), 0063- 002 (100), 0185-002 (15), 0188-001 (127), 0189-001 (20), 0190-001 (16), 0242-002 (13), 1051-004 (9), 1052- 017 (1), 3115-001 (1) and 3134-003 (1); FMNH 5948 (2), 5949 (3), 5950 (1), 5951 (1), 5952 (2), 5953 (6), 5954 (17), 5955 (8), 5956 (3), 5957 (5), 5958 (21), 5959 (11), 5960 (28), 5961 (11), 5962 (6), 5963 (13), 5964 (60), 5965 (16), 5966 (5), 5967 (9), 5968 (15), 5969 (14), 6206 (2), 6207 (7), 6208 (4), 6209 (1), 6212 (3), 6214 (2), 6215 (3), 6216 (5), 6220 (9), 33007 (3), 33010 (9), 52518 (1), 52519 (1), 52520 (1), 52521 (1), 52522 (1), 52523 (3), 52532 (1) and 95001 (25); LSUMZ 15443 (1); other collections: IBUNAM 13696 (1), 13698 (1), 13700 (1), 13701 (1) and UMMZ 245140 (1).
<emphasis id="B951EA8CE86CFF9AFDCA2B49FD1E330C" bold="true" box="[548,703,1878,1904]" pageId="47" pageNumber="48">Distribution:</emphasis>
Central America; from central Nicaragua to northern Costa Rica, Atlantic drainages;
<emphasis id="B951EA8CE86CFF9AFE972B65FDA933E8" bold="true" box="[377,520,1914,1940]" pageId="47" pageNumber="48">
(
<collectingCountry id="F332760EE86CFF9AFE6E2B65FDA133E8" box="[384,512,1914,1940]" name="Nicaragua" pageId="47" pageNumber="48">Nicaragua</collectingCountry>
)
</emphasis>
BF, IM, To, Sa, SJ, RF and LN (Atlantic);
<quantity id="4CDD9B7BE86CFF9AFBE42B65FB3333E9" box="[1034,1170,1914,1941]" metricMagnitude="1" metricUnit="m" metricValue="6.95" metricValueMax="13.5" metricValueMin="0.4" pageId="47" pageNumber="48" unit="m" value="69.5" valueMax="135.0" valueMin="4.0">4135 masl</quantity>
; Sec, Pot.
<emphasis id="B951EA8CE86CFF9AFAFF2B65FE1C33C5" bold="true" pageId="47" pageNumber="48">Occurrence and conservation status:</emphasis>
Nat; LC (2020), population trend stable.
<emphasis id="B951EA8CE86CFF9AFC782B81FBBD33C4" bold="true" box="[918,1052,1950,1976]" pageId="47" pageNumber="48">Literature:</emphasis>
<bibRefCitation id="EFB44B6FE86CFF9AFBCA2B80FB4833C5" author="Meek" box="[1060,1257,1951,1977]" firstAuthor="Meek" page="121" pageId="47" pageNumber="48" pagination="97 - 132" refId="ref84723" refString="Meek, S. E. (1907) Synopsis of the fishes of the Great Lakes of Nicaragua. Field Columbian Museum Zoological Series, 7, 97 - 132. https: // doi. org / 10.5962 / bhl. title. 3761" type="journal article" year="1907">Meek (1907: 121</bibRefCitation>
, as
<taxonomicName id="4C254D1DE86CFF9AFAF82B80FEA633A1" baseAuthorityName="Gïnther" baseAuthorityYear="1864" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="citinellum">
<emphasis id="B951EA8CE86CFF9AFAF82B80FEA633A1" italics="true" pageId="47" pageNumber="48">Cichlasoma citinellum</emphasis>
</taxonomicName>
—synonym, also as
<taxonomicName id="4C254D1DE86CFF9AFE0D2BDCFC2733A1" authority="(Meek, 1097)" baseAuthorityName="Meek" baseAuthorityYear="1097" box="[483,902,1987,2013]" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="granadense">
<emphasis id="B951EA8CE86CFF9AFE0D2BDCFD5033A1" box="[483,753,1987,2013]" italics="true" pageId="47" pageNumber="48">Cichlasoma granadense</emphasis>
(Meek, 1097)
</taxonomicName>
—misidentification; brief description, including an identification key, and information on distribution and ecology), Astorqui (1975: 134, as
<taxonomicName id="4C254D1DE873FF85FB792C88FA3734CD" baseAuthorityName="Gïnther" baseAuthorityYear="1864" box="[1175,1430,151,177]" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="citrinellus">
<emphasis id="B951EA8CE873FF85FB792C88FA3734CD" box="[1175,1430,151,177]" italics="true" pageId="48" pageNumber="49">Cichlasoma citrinellus</emphasis>
</taxonomicName>
, in part—synonym; brief description, including illustrations and an identification key, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE873FF85FED52CC0FE583485" author="Villa" box="[315,505,223,249]" firstAuthor="Villa" page="190" pageId="48" pageNumber="49" refId="ref86666" refString="Villa, J. (1982) Peces nicaraguenses de agua dulce. Editorial Union Cardoza y Cia. Ltda, Managua, 253 pp." type="book" year="1982">Villa (1982: 190</bibRefCitation>
, as
<taxonomicName id="4C254D1DE873FF85FDC72CC0FD663485" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[553,711,223,249]" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="citrinellum">
<emphasis id="B951EA8CE873FF85FDC72CC0FD663485" box="[553,711,223,249]" italics="true" pageId="48" pageNumber="49">C. citrinellum</emphasis>
</taxonomicName>
, in part—synonym; detailed description, including illustrations and an identification key, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE873FF85FC0A2D1CFB6B3561" author="Bussing" box="[996,1226,259,285]" firstAuthor="Bussing" page="312" pageId="48" pageNumber="49" refId="ref81779" refString="Bussing, W. A. (1998). Freshwater fishes of Costa Rica. Editorial Universidad de Costa Rica, San Jose´, Costa Rica, 468 pp." type="book" year="1998">Bussing (1998: 312</bibRefCitation>
; brief description, including illustrations and an identification key, and information on distribution, with a map, and ecology),
<bibRefCitation id="EFB44B6FE873FF85FAAD2D38FE213519" author="Stauffer &amp; McKaye" firstAuthor="Stauffer" page="8" pageId="48" pageNumber="49" pagination="1 - 18" refId="ref86208" refString="Stauffer, J. R. Jr. &amp; McKaye, K. R. (2002) Descriptions of three new species of cichlid fishes (Teleostei: Cichlidae) from Lake Xiloa, Nicaragua. Cuadernos de investigacion de la UCA, Universidad Centroamericana, 12, 1 - 18." type="journal article" year="2002">Stauffer &amp; McKaye (2002: 8</bibRefCitation>
; comparative morphological data and information on distribution), González-Alemán (2006: 49; brief description and information on distribution and ecology; the photo illustrating the species corresponds to a specimen of
<taxonomicName id="4C254D1DE873FF85FEB42D8CFC8E35D1" authority="(Gunther 1867))" authorityName="" baseAuthorityName="Gunther" baseAuthorityYear="1867" box="[346,815,403,429]" family="Cichlidae" genus="Cribroheros" kingdom="Animalia" order="Perciformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="B951EA8CE873FF85FEB42D8CFDCC35D1" box="[346,621,403,429]" italics="true" pageId="48" pageNumber="49">Cribroheros longimanus</emphasis>
(Günther 1867))
</taxonomicName>
,
<bibRefCitation id="EFB44B6FE873FF85FCD52D8CFBFB35D1" author="Stauffer" box="[827,1114,403,429]" etAl="et al." firstAuthor="Stauffer" page="117" pageId="48" pageNumber="49" pagination="117 - 129" refId="ref86137" refString="Stauffer, J. R. Jr., McCrary, J. K. &amp; Black, K. E. (2008) Three new species of cichlid fishes (Teleostei: Cichlidae) from Lake Apoyo, Nicaragua. Proceedings of the Biological Society of Washington, 121 (1), 117 - 129. https: // doi. org / 10.2988 / 06 - 37.1" type="journal article" year="2008">
Stauffer
<emphasis id="B951EA8CE873FF85FC732D8CFC7735D1" box="[925,982,403,429]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
(2008: 117
</bibRefCitation>
; comparative morphological data and information on distribution),
<bibRefCitation id="EFB44B6FE873FF85FDA52DA8FC0135AD" author="Klingenberg" box="[587,928,439,465]" etAl="et al." firstAuthor="Klingenberg" page="397" pageId="48" pageNumber="49" pagination="397 - 408" refId="ref83333" refString="Klingenberg, C. P., Barluenga, M. &amp; Meyer, A. (2003) Body shape variation in cichlid fishes of the Amphilophus citrinellus species complex. Biological Journal of the Linnean Society, 80 (3), 397 - 408. https: // doi. org / 10.1046 / j. 1095 - 8312.2003.00246. x" type="journal article" year="2003">
Klingenberg
<emphasis id="B951EA8CE873FF85FD312DA8FCB835AD" box="[735,793,439,465]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
(2003: 397
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; body shape variation and comparison, teeth morphology, and distribution),
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