312 lines
35 KiB
XML
312 lines
35 KiB
XML
<document id="D7A7713D6CB4D37C18A19F385708D965" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter & Bogorodsky, Sergey V." docDate="2012" docId="038C87863F7EFFC6FF36641349DF61D5" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Helcogramma obtusirostris Klunzinger 1871" docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="54" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="ACC67DF05E07736A0911EAA502E5D845">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<mods:namePart id="454A2655AC9522B6AD6E25FEACA67F0C">Holleman, Wouter</mods:namePart>
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<mods:namePart id="21D0F7008A0D86E8C9A0DF92B822119E">Bogorodsky, Sergey V.</mods:namePart>
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<mods:title id="EB16815EDE1FF242D2E938D37A6EDBAD">Zootaxa</mods:title>
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<mods:date id="B759D49C69FA7D43380CF295F1D9AC18">2012</mods:date>
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<treatment id="038C87863F7EFFC6FF36641349DF61D5" ID-DOI="http://doi.org/10.5281/zenodo.5626536" ID-GBIF-Taxon="119427994" ID-Zenodo-Dep="5626536" LSID="urn:lsid:plazi:treatment:038C87863F7EFFC6FF36641349DF61D5" httpUri="http://treatment.plazi.org/id/038C87863F7EFFC6FF36641349DF61D5" lastPageId="20" lastPageNumber="56" pageId="18" pageNumber="54">
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<subSubSection id="C33F651B3F7EFFC0FF3664134950610B" pageId="18" pageNumber="54" type="nomenclature">
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<paragraph id="8B9A36903F7EFFC0FF3664134AE5616A" blockId="18.[151,719,1570,1629]" box="[151,719,1570,1596]" pageId="18" pageNumber="54">
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<heading id="D0D281FC3F7EFFC0FF3664134AE5616A" bold="true" box="[151,719,1570,1596]" fontSize="11" level="1" pageId="18" pageNumber="54" reason="1">
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<taxonomicName id="4C254D133F7EFFC0FF3664134AE5616A" ID-CoL="3K47Q" authority="Klunzinger, 1871" authorityName="Klunzinger" authorityYear="1871" box="[151,719,1570,1596]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="obtusirostris">
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<emphasis id="B951EA823F7EFFC0FF3664134AE5616A" bold="true" box="[151,719,1570,1596]" pageId="18" pageNumber="54">
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<emphasis id="B951EA823F7EFFC0FF36641349CB616A" bold="true" box="[151,481,1570,1596]" italics="true" pageId="18" pageNumber="54">Helcogramma obtusirostris</emphasis>
|
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(
|
||
<bibRefCitation id="EFB44B613F7EFFC0FE4E64134AED616A" author="Klunzinger" box="[495,711,1570,1596]" pageId="18" pageNumber="54" refString="Klunzinger, C. B. (1871) Synopsis der Fische de Rothen Meeres. Verhandellungen der Kaiserlich-Konichlichen Zoologish-botanischen Gesellschaft in Wien, 21, 441 - 688." type="journal article" year="1871">Klunzinger, 1871</bibRefCitation>
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||
)
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B9A36903F7EFFC0FF3664744950610B" blockId="18.[151,719,1570,1629]" box="[151,378,1605,1629]" pageId="18" pageNumber="54">
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<figureCitation id="131E2A153F7EFFC0FF36647448FC610B" box="[151,214,1605,1629]" captionStart="FIGURE 8" captionStartId="19.[151,250,1883,1906]" captionTargetBox="[179,1414,193,1852]" captionTargetId="figure@19.[179,1415,193,1853]" captionTargetPageId="19" captionText="FIGURE 8. (A) Enneapterygius obscurus, Marsa Alam; (B) Enneapterygius pallidus, Farasan Is.; (C) Enneapterygius pusillus, M, Sharm el Sheik; (D) Enneapterygius pusillus, M, Oman; (E) Enneapterygius ventermaculus, Farasan Is.; (E) Helcogramma obtusirostris, M, Sharm el Sheil; (F) Helcogramma obtusirostris, F, Sharm el Sheik; (G) Helcogramma steinitzi, F, Marsa Alam; (H) Norfolkia brachylepis, Bali. Photos by R. Field (B, E), J. E. Randall (D); G. R. Allen (H); S. V. Bogorodsky (the others)." httpUri="https://zenodo.org/record/279581/files/figure.png" pageId="18" pageNumber="54">Fig. 8</figureCitation>
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F & G; table 1
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F651B3F7EFFC0FF3664B94C726000" pageId="18" pageNumber="54" type="reference_group">
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||
<paragraph id="8B9A36903F7EFFC0FF3664B94C6461CA" blockId="18.[151,1436,1670,1878]" box="[151,1102,1670,1693]" pageId="18" pageNumber="54">
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<treatmentCitationGroup id="AB3511BE3F7EFFC0FF3664B94C6461CA" box="[151,1102,1670,1693]" pageId="18" pageNumber="54">
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<taxonomicName id="4C254D133F7EFFC0FF3664B94AA061CA" authority="Klunzinger, 1871: 498" authorityName="Klunzinger" authorityPageNumber="498" authorityYear="1871" box="[151,650,1670,1693]" class="Magnoliopsida" family="Celastraceae" genus="Tripterygium" kingdom="Animalia" order="Celastrales" pageId="18" pageNumber="54" phylum="Tracheophyta" rank="species" species="obtusirostris">
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<emphasis id="B951EA823F7EFFC0FF3664B9498A61CB" box="[151,416,1671,1693]" italics="true" pageId="18" pageNumber="54">Tripterygium obtusirostris</emphasis>
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<treatmentCitation id="0A8410813F7EFFC0FE0664B74AA061CA" author="Klunzinger" box="[423,650,1670,1692]" page="498" pageId="18" pageNumber="54" year="1871">
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<bibRefCitation id="EFB44B613F7EFFC0FE0664B74A7061CA" author="Klunzinger" box="[423,602,1670,1692]" pageId="18" pageNumber="54" refString="Klunzinger, C. B. (1871) Synopsis der Fische de Rothen Meeres. Verhandellungen der Kaiserlich-Konichlichen Zoologish-botanischen Gesellschaft in Wien, 21, 441 - 688." type="journal article" year="1871">Klunzinger, 1871</bibRefCitation>
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||
: 498
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(Red Sea).
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<treatmentCitation id="0A8410813F7EFFC0FCA364B74C6061CA" author="Golani" box="[770,1098,1670,1692]" page="45" pageId="18" pageNumber="54" year="2010">
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<bibRefCitation id="EFB44B613F7EFFC0FCA364B74C0E61CA" author="Golani" box="[770,1060,1670,1692]" pageId="18" pageNumber="54" refString="Golani, D. & Bogorodsky, S. V. (2010) The Fishes of the Red Sea-Reappraisal and Updated Checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani & Bogorodsky, 2010</bibRefCitation>
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: 45
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8B9A36903F7EFFC0FF3664944AC8618F" blockId="18.[151,1436,1670,1878]" pageId="18" pageNumber="54">
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<treatmentCitationGroup id="AB3511BE3F7EFFC0FF3664944AC8618F" pageId="18" pageNumber="54">
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||
<taxonomicName id="4C254D133F7EFFC0FF3664944A7F61ED" ID-CoL="58ZLV" authority="Clark, 1980: 85" authorityName="Clark" authorityPageNumber="85" authorityYear="1980" box="[151,597,1701,1723]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="obtusirostre">
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||
<emphasis id="B951EA823F7EFFC0FF366494498C61ED" box="[151,422,1701,1723]" italics="true" pageId="18" pageNumber="54">Helcogramma obtusirostre</emphasis>
|
||
:
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||
<treatmentCitation id="0A8410813F7EFFC0FE1564944A7F61ED" author="Clark" box="[436,597,1701,1723]" page="85" pageId="18" pageNumber="54" year="1980">
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<bibRefCitation id="EFB44B613F7EFFC0FE1564944A1A61ED" author="Clark" box="[436,560,1701,1723]" pageId="18" pageNumber="54" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
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: 85
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, figs. 3e, 7 & plate 1; Holleman in:
|
||
<treatmentCitation id="0A8410813F7EFFC0FC6764944CD161ED" author="Smith" box="[966,1275,1701,1723]" page="757" pageId="18" pageNumber="54" year="1986">
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||
<bibRefCitation id="EFB44B613F7EFFC0FC6764944CE261ED" author="Smith" box="[966,1224,1701,1723]" pageId="18" pageNumber="54" refString="Smith, M. M. & Heemstra, P. C. (Eds.) (1986) Smiths' Sea Fishes. Macmillan, Johannesburg, xx + 1047 pp. 144 pls." type="book" year="1986">Smith & Heemstra, 1986</bibRefCitation>
|
||
: 757
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="0A8410813F7EFFC0FAA7649448DA618F" author="Randall" page="309" pageId="18" pageNumber="54" year="1995">
|
||
<bibRefCitation id="EFB44B613F7EFFC0FAA764944DB261ED" author="Randall" box="[1286,1432,1701,1723]" pageId="18" pageNumber="54" refString="Randall, J. E. (1995) A review of the triplefin fishes (Blennioidei: Tripterygiidae) of Oman, with the description of two new species. Revue francaise d'Aquariologie, 22, 27 - 34." type="journal article" year="1995">Randall, 1995</bibRefCitation>
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||
: 309
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||
</treatmentCitation>
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||
, fig. 872;
|
||
<treatmentCitation id="0A8410813F7EFFC0FEF464F24A0B618F" author="Holleman" box="[341,545,1731,1753]" page="68" pageId="18" pageNumber="54" year="2007">
|
||
<bibRefCitation id="EFB44B613F7EFFC0FEF464F249D6618F" author="Holleman" box="[341,508,1731,1753]" pageId="18" pageNumber="54" refString="Holleman, W. (2007) Fishes of the genus Helcogramma (Blennioidei: Tripterygiidae) in the Western Indian Ocean, including Sri Lanka, with descriptions of four new species. Smithiana Bulletin, 7, 51 - 81." type="journal article" year="2007">Holleman, 2007</bibRefCitation>
|
||
: 68
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||
</treatmentCitation>
|
||
,
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||
<figureCitation id="131E2A153F7EFFC0FD8D64F24A4D618F" box="[556,615,1731,1753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1157,1180]" captionTargetBox="[164,1424,187,1146]" captionTargetId="figure@4.[164,1424,187,1146]" captionTargetPageId="4" captionText="FIGURE 1. Map of the Red Sea showing localities mentioned in the text." httpUri="https://zenodo.org/record/279574/files/figure.png" pageId="18" pageNumber="54">Fig. 1</figureCitation>
|
||
& 10, Pl. 2.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8B9A36903F7EFFC0FF3664D34B1261AE" blockId="18.[151,1436,1670,1878]" box="[151,824,1762,1785]" pageId="18" pageNumber="54">
|
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<treatmentCitationGroup id="AB3511BE3F7EFFC0FF3664D34B1261AE" box="[151,824,1762,1785]" pageId="18" pageNumber="54">
|
||
<taxonomicName id="4C254D133F7EFFC0FF3664D34A5361AE" ID-CoL="3K47Q" authority="Hansen, 1986: 341" authorityName="Hansen" authorityPageNumber="341" authorityYear="1986" box="[151,633,1762,1784]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="obtusirostris">
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||
<emphasis id="B951EA823F7EFFC0FF3664D3498761AE" box="[151,429,1762,1784]" italics="true" pageId="18" pageNumber="54">Helcogramma obtusirostris</emphasis>
|
||
:
|
||
<treatmentCitation id="0A8410813F7EFFC0FE1B64D24A5361AE" author="Hansen" box="[442,633,1762,1784]" page="341" pageId="18" pageNumber="54" year="1986">
|
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<bibRefCitation id="EFB44B613F7EFFC0FE1B64D24A6361AE" author="Hansen" box="[442,585,1762,1784]" pageId="18" pageNumber="54" refString="Hansen, P. E. H. (1986) Revision of the tripterygiid fish genus Helcogramma, including descriptions of four new species. Bulletin of Marine Science, 38 (2), 313 - 354." type="journal article" year="1986">Hansen, 1986</bibRefCitation>
|
||
: 341
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(in part: Red Sea).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8B9A36903F7EFFC0FF3665304B866041" blockId="18.[151,1436,1670,1878]" box="[151,940,1793,1815]" pageId="18" pageNumber="54">
|
||
<treatmentCitationGroup id="AB3511BE3F7EFFC0FF3665304B866041" box="[151,940,1793,1815]" pageId="18" pageNumber="54">
|
||
<taxonomicName id="4C254D133F7EFFC0FF36653049A56041" ID-CoL="3K485" box="[151,399,1793,1815]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="trigloides">
|
||
<emphasis id="B951EA823F7EFFC0FF36653049A56041" box="[151,399,1793,1815]" italics="true" pageId="18" pageNumber="54">Helcogramma trigloides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B951EA823F7EFFC0FE3F653349EE6041" box="[414,452,1794,1815]" italics="true" pageId="18" pageNumber="54">non</emphasis>
|
||
Bleeker):
|
||
<treatmentCitation id="0A8410813F7EFFC0FD8F65304AD16041" author="Marshall" box="[558,763,1793,1815]" page="242" pageId="18" pageNumber="54" year="1952">
|
||
<bibRefCitation id="EFB44B613F7EFFC0FD8F65304AE16041" author="Marshall" box="[558,715,1793,1815]" pageId="18" pageNumber="54" refString="Marshall, N. B. (1952) The Manihine Expedition to the Gulf of Aqaba, 1948 - 49. Bulletin of the British Museum (Natural History), Zoology, 1 (8), 221 - 252." type="journal article" year="1952">
|
||
<collectingCountry id="F33276003F7EFFC0FD8F65304AA16041" box="[558,651,1793,1815]" name="Marshall Islands" pageId="18" pageNumber="54">Marshall</collectingCountry>
|
||
, 1952
|
||
</bibRefCitation>
|
||
: 242
|
||
</treatmentCitation>
|
||
(Gulf of Aqaba).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F7EFFC0FF3665104C726000" blockId="18.[151,1436,1670,1878]" pageId="18" pageNumber="54">
|
||
<treatmentCitationGroup id="AB3511BE3F7EFFC0FF3665104C726000" pageId="18" pageNumber="54">
|
||
<emphasis id="B951EA823F7EFFC0FF36651049E06060" box="[151,458,1824,1846]" italics="true" pageId="18" pageNumber="54">
|
||
non
|
||
<taxonomicName id="4C254D133F7EFFC0FF62651149E06060" ID-CoL="3K47X" authority="Lal Mohan, 1971: 219" authorityName="Lal Mohan" authorityPageNumber="219" authorityYear="1971" box="[195,458,1824,1846]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="shinglensis">Helcogramma shinglensis</taxonomicName>
|
||
</emphasis>
|
||
:
|
||
<treatmentCitation id="0A8410813F7EFFC0FE7965114A966063" author="Lal" box="[472,700,1824,1846]" page="219" pageId="18" pageNumber="54" year="1971">
|
||
<bibRefCitation id="EFB44B613F7EFFC0FE7965114AA06063" author="Lal" box="[472,650,1824,1846]" pageId="18" pageNumber="54" refString="Lal Mohan, R. S. (1971) Helcogramma shinglensis, a new species of tripterygiid fish from the Gulf of Mannar with a key to the fishes of the family Tripterygiidae of the eastern and central Indian Ocean. Senckenbergiana Biologica 52 (3 / 5), 219 - 223." type="journal article" year="1971">Lal Mohan, 1971</bibRefCitation>
|
||
: 219
|
||
</treatmentCitation>
|
||
, fig. 1 (Gulf of Mannar,
|
||
<collectingCountry id="F33276003F7EFFC0FC1765114BC56060" box="[950,1007,1824,1846]" name="India" pageId="18" pageNumber="54">India</collectingCountry>
|
||
).
|
||
<emphasis id="B951EA823F7EFFC0FF64657049D66000" box="[197,508,1856,1878]" italics="true" pageId="18" pageNumber="54">
|
||
non
|
||
<taxonomicName id="4C254D133F7EFFC0FF50657149D66000" ID-CoL="3K46K" authority="Lubbock, 1980: 294" authorityName="Lubbock" authorityPageNumber="294" authorityYear="1980" box="[241,508,1856,1878]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="54" phylum="Chordata" rank="species" species="ascensionis">Helcogramma ascensionis</taxonomicName>
|
||
</emphasis>
|
||
:
|
||
<treatmentCitation id="0A8410813F7EFFC0FDAB65714AF36000" author="Lubbock" box="[522,729,1856,1878]" page="294" pageId="18" pageNumber="54" year="1980">
|
||
<bibRefCitation id="EFB44B613F7EFFC0FDAB65714A8D6000" author="Lubbock" box="[522,679,1856,1878]" pageId="18" pageNumber="54" refString="Lubbock, R. (1980) The shore fishes of Ascension Island. Journal of Fish Biology, 17, 283 - 303." type="journal article" year="1980">Lubbock, 1980</bibRefCitation>
|
||
: 294
|
||
</treatmentCitation>
|
||
, fig. 2 (
|
||
<collectingCountry id="F33276003F7EFFC0FC8765714BBB6000" box="[806,913,1856,1878]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="18" pageNumber="54">Ascension</collectingCountry>
|
||
<collectingCountry id="F33276003F7EFFC0FC3665714BFD6000" box="[919,983,1856,1878]" name="Iceland" pageId="18" pageNumber="54">Island</collectingCountry>
|
||
, S Atlantic).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5A66183F7FFFC1FF36656A48CE60BA" httpUri="https://zenodo.org/record/279581/files/figure.png" pageId="19" pageNumber="55" targetBox="[179,1414,193,1852]" targetPageId="19">
|
||
<paragraph id="8B9A36903F7FFFC1FF36656A48CE60BA" blockId="19.[151,1437,1883,2028]" pageId="19" pageNumber="55">
|
||
<emphasis id="B951EA823F7FFFC1FF36656A49386026" bold="true" box="[151,274,1883,1906]" pageId="19" pageNumber="55">FIGURE 8.</emphasis>
|
||
(A)
|
||
<taxonomicName id="4C254D133F7FFFC1FEE0656D4AE76027" authority="Marsa Alam" authorityName="Marsa Alam" box="[321,717,1883,1905]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="obscurus">
|
||
<emphasis id="B951EA823F7FFFC1FEE0656D4A6A6027" box="[321,576,1883,1905]" italics="true" pageId="19" pageNumber="55">Enneapterygius obscurus</emphasis>
|
||
, Marsa Alam
|
||
</taxonomicName>
|
||
; (B)
|
||
<taxonomicName id="4C254D133F7FFFC1FD5F656D4C5A6027" authority="Farasan Is." authorityName="Farasan Is." box="[766,1136,1883,1905]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="pallidus">
|
||
<emphasis id="B951EA823F7FFFC1FD5F656D4BDE6027" box="[766,1012,1883,1905]" italics="true" pageId="19" pageNumber="55">Enneapterygius pallidus</emphasis>
|
||
, Farasan Is.
|
||
</taxonomicName>
|
||
; (C)
|
||
<taxonomicName id="4C254D133F7FFFC1FB03656D4DBC6027" box="[1186,1430,1883,1905]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="pusillus">
|
||
<emphasis id="B951EA823F7FFFC1FB03656D4DBC6027" box="[1186,1430,1883,1905]" italics="true" pageId="19" pageNumber="55">Enneapterygius pusillus</emphasis>
|
||
</taxonomicName>
|
||
, M, Sharm el Sheik; (D)
|
||
<taxonomicName id="4C254D133F7FFFC1FE2D654A4AAB60C6" box="[396,641,1914,1936]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="pusillus">
|
||
<emphasis id="B951EA823F7FFFC1FE2D654A4AAB60C6" box="[396,641,1914,1936]" italics="true" pageId="19" pageNumber="55">Enneapterygius pusillus</emphasis>
|
||
</taxonomicName>
|
||
, M, Oman; (E)
|
||
<taxonomicName id="4C254D133F7FFFC1FC82654A4CF360C6" authority="Farasan Is." authorityName="Farasan Is." box="[803,1241,1914,1936]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="ventermaculus">
|
||
<emphasis id="B951EA823F7FFFC1FC82654A4C7660C6" box="[803,1116,1914,1936]" italics="true" pageId="19" pageNumber="55">Enneapterygius ventermaculus</emphasis>
|
||
, Farasan Is.
|
||
</taxonomicName>
|
||
; (E)
|
||
<taxonomicName id="4C254D133F7FFFC1FAAA654B493C60F8" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="obtusirostris">
|
||
<emphasis id="B951EA823F7FFFC1FAAA654B493C60F8" italics="true" pageId="19" pageNumber="55">Helcogramma obtusirostris</emphasis>
|
||
</taxonomicName>
|
||
, M, Sharm el Sheil; (F)
|
||
<taxonomicName id="4C254D133F7FFFC1FD8365A94B1160F8" box="[546,827,1944,1966]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="obtusirostris">
|
||
<emphasis id="B951EA823F7FFFC1FD8365A94B1160F8" box="[546,827,1944,1966]" italics="true" pageId="19" pageNumber="55">Helcogramma obtusirostris</emphasis>
|
||
</taxonomicName>
|
||
, F, Sharm el Sheik; (G)
|
||
<taxonomicName id="4C254D133F7FFFC1FBE665A94D1B60F9" box="[1095,1329,1944,1967]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B951EA823F7FFFC1FBE665A94D1B60F9" box="[1095,1329,1944,1967]" italics="true" pageId="19" pageNumber="55">Helcogramma steinitzi</emphasis>
|
||
</taxonomicName>
|
||
, F, Marsa Alam; (H)
|
||
<taxonomicName id="4C254D133F7FFFC1FEAD658649C6609B" box="[268,492,1975,1997]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="55" phylum="Chordata" rank="species" species="brachylepis">
|
||
<emphasis id="B951EA823F7FFFC1FEAD658649C6609B" box="[268,492,1975,1997]" italics="true" pageId="19" pageNumber="55">Norfolkia brachylepis</emphasis>
|
||
</taxonomicName>
|
||
, Bali. Photos by R. Field (B, E), J.E. Randall (D); G.R. Allen (H); S.V. Bogorodsky (the others).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F651B3F78FFC6FF3662A6498F64AD" pageId="20" pageNumber="56" type="diagnosis">
|
||
<paragraph id="8B9A36903F78FFC6FF3662A648C366D7" blockId="20.[151,1436,151,1019]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF3662A6493B67E6" bold="true" box="[151,273,151,176]" pageId="20" pageNumber="56">Diagnosis.</emphasis>
|
||
Dorsal fins III + XII–XIII + 9–10 (rarely with XII spines or 9 rays); anal fin I, 18–19 (usually 19); pectoral fin 16: usually 2, 7, 7. LL 20–23 (usually 21–22) tubed scales, ending below junction of second and third dorsal fins; total lateral scales 36–38 (usually 37–38); transverse scales 7/5. Dentary pores 4 + 1 + 4. Head length 3.3–3.7 (3.5) in SL. Nape and belly naked; scales do not extend to bases of first dorsal and anterior of anal fins; 2 rows of scales on base of caudal fin. Pelvic–fin rays united by membrane for half shorter ray, longest ray reaching vent. First dorsal fin half height of second. Maxilla reaches vertical through centre of pupil; orbital cirrus small and simple.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F78FFC6FF6463BC4B0865EA" blockId="20.[151,1436,151,1019]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF6463BC494B66F3" bold="true" box="[197,353,397,421]" pageId="20" pageNumber="56">Fresh colour.</emphasis>
|
||
(From colour photograph by the second author and J. E. Randall.) Body of
|
||
<emphasis id="B951EA823F78FFC6FB1163BD4CDB66F2" box="[1200,1265,396,420]" italics="true" pageId="20" pageNumber="56">males</emphasis>
|
||
mottled greenish with darker interconnecting areas, with 4 narrow, pale saddle marks extending from base of dorsal fins to midline, 2 below second dorsal fin, 2 below third dorsal fin; series of paler spots along midline; dorso–anteriorly suffused with dull red; belly pink. Head dark, black below eye and to branchiostegal membrane; bright blue stripe extends from lower lip across cheek, broadly edged with black above and below. Pectoral–fin base with red spot above and below, the lower one with blue above and anterior; pelvic fins dark pink proximally; first dorsal fin dusky with some orange, darkest on the margin, and with small black spot at base of membrane connecting first and second dorsal fins; second dorsal fin dusky with dull orange band basally; third dorsal fin dusky; caudal fin dusky; anal fin dark, with an even cover of melanophores.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F78FFC6FF6460F64CB66472" blockId="20.[151,1436,151,1019]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF6460F6490E6589" box="[197,292,711,735]" italics="true" pageId="20" pageNumber="56">Females</emphasis>
|
||
dull green, the dark areas forming a zig–zag pattern down the body, on a pale green background; body pale ventrally. Head dorsally dark green, ventrally pale; median fins with light sprinkling of melanophores, first dorsal fin with small black spots as in males, basal third of dorsal and entire anal fins white.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F78FFC6FF64611E498F64AD" blockId="20.[151,1436,151,1019]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF64611E4998641E" bold="true" box="[197,434,815,840]" pageId="20" pageNumber="56">Colour underwater.</emphasis>
|
||
Body of males translucent with reddish ‘H’-bars and streak of white marks from dorsal fins onto body, belly and pelvic fins pinkish-red; head reddish above and with many melanophores below eyes, extending onto base of pectoral fin; blue-white line from lower lip, through corner of mouth to preopercle, with blue patch below; dorsal fins translucent with red and white marks, caudal fin pink, pectoral fins translucent with red and white marks. Females paler versions of males and lack the distinctive pigmentation on the head and belly, which are largely white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F78FFC6FF64661D48DF63DD" pageId="20" pageNumber="56" type="key">
|
||
<paragraph id="8B9A36903F78FFC6FF64661D48DF63DD" blockId="20.[151,1437,1067,1667]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF64661D49496312" bold="true" box="[197,355,1067,1092]" pageId="20" pageNumber="56">Key features.</emphasis>
|
||
Iridescent blue band from corner of mouth to opercle in males; males in ‘breeding dress’ nearly entirely black with 3 or 4 white ‘slashes’ below second and third dorsal fins; females whitish with irregular brownish bars.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F78FFC6FF6466A64BC663E6" box="[197,1004,1175,1200]" pageId="20" pageNumber="56" type="distribution">
|
||
<paragraph id="8B9A36903F78FFC6FF6466A64BC663E6" blockId="20.[151,1437,1067,1667]" box="[197,1004,1175,1200]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF6466A6497463E6" bold="true" box="[197,350,1175,1200]" pageId="20" pageNumber="56">Distribution.</emphasis>
|
||
<taxonomicName id="4C254D133F78FFC6FEC466A84A3E63E6" box="[357,532,1176,1200]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="obtusirostris">
|
||
<emphasis id="B951EA823F78FFC6FEC466A84A3E63E6" box="[357,532,1176,1200]" italics="true" pageId="20" pageNumber="56">H. obtusirostris</emphasis>
|
||
</taxonomicName>
|
||
occurs in the Red Sea,
|
||
<collectingCountry id="F33276003F78FFC6FCBA66A84B4363E6" box="[795,873,1177,1200]" name="Yemen" pageId="20" pageNumber="56">Yemen</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33276003F78FFC6FC0166A94BC263E6" box="[928,1000,1176,1200]" name="Oman" pageId="20" pageNumber="56">Oman</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F651B3F78FFC6FF64668D49DF61D5" pageId="20" pageNumber="56" type="discussion">
|
||
<paragraph id="8B9A36903F78FFC6FF64668D49BA6235" blockId="20.[151,1437,1067,1667]" pageId="20" pageNumber="56">
|
||
<emphasis id="B951EA823F78FFC6FF64668D49136382" bold="true" box="[197,313,1212,1236]" pageId="20" pageNumber="56">Remarks.</emphasis>
|
||
<taxonomicName id="4C254D133F78FFC6FEE1668C49DA6382" box="[320,496,1212,1236]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="obtusirostris">
|
||
<emphasis id="B951EA823F78FFC6FEE1668C49DA6382" box="[320,496,1212,1236]" italics="true" pageId="20" pageNumber="56">H. obtusirostris</emphasis>
|
||
</taxonomicName>
|
||
is one of a group of widespread Indo-west Pacific species with at least four sibling species, which all have an iridescent blue line running from the lower lip to the corner of the mouth and onto the opercle. It is easily distinguished from
|
||
<taxonomicName id="4C254D133F78FFC6FDE667344AEA624A" box="[583,704,1285,1308]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B951EA823F78FFC6FDE667344AEA624A" box="[583,704,1285,1308]" italics="true" pageId="20" pageNumber="56">H. steinitzi</emphasis>
|
||
</taxonomicName>
|
||
, which is reddish in general colour, has a large head and lacks the blue line from the corner of the mouth. Usually seen in pairs in coral and rocky areas of lagoons in depths of
|
||
<quantity id="4CDD9B753F78FFC6FAFC6719489A6235" metricMagnitude="0" metricUnit="m" metricValue="4.25" metricValueMax="8.0" metricValueMin="0.5" pageId="20" pageNumber="56" unit="m" value="4.25" valueMax="8.0" valueMin="0.5">
|
||
0.5–
|
||
<specimenCount id="9D23FD193F78FFC6FA316719489A6235" pageId="20" pageNumber="56" type="generic">8 m</specimenCount>
|
||
</quantity>
|
||
. Attains
|
||
<quantity id="4CDD9B753F78FFC6FEB2677A49486235" box="[275,354,1355,1379]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="56" unit="mm" value="38.0">38 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F78FFC6FF6467414AF562A5" blockId="20.[151,1437,1067,1667]" pageId="20" pageNumber="56">
|
||
the west coast of Indian and has been recorded from
|
||
<collectingCountry id="F33276003F78FFC6FCB567414B8962DE" box="[788,931,1392,1416]" name="South Africa" pageId="20" pageNumber="56">South Africa</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33276003F78FFC6FC7B67414C6D62DE" box="[986,1095,1392,1416]" name="Mauritius" pageId="20" pageNumber="56">Mauritius</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB44B613F78FFC6FBF467414D3A62DE" author="Holleman" box="[1109,1296,1392,1416]" pageId="20" pageNumber="56" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C254D133F78FFC6FA8267404DB162DE" box="[1315,1435,1392,1416]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="pallidus">
|
||
<emphasis id="B951EA823F78FFC6FA8267404DB162DE" box="[1315,1435,1392,1416]" italics="true" pageId="20" pageNumber="56">E. pallidus</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the Red Sea and, again, is found throughout.
|
||
<taxonomicName id="4C254D133F78FFC6FC9167A44BD962FA" box="[816,1011,1428,1452]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="ventermaculus">
|
||
<emphasis id="B951EA823F78FFC6FC9167A44BD962FA" box="[816,1011,1428,1452]" italics="true" pageId="20" pageNumber="56">E. ventermaculus</emphasis>
|
||
</taxonomicName>
|
||
occurs in tide pools and to a depth of some
|
||
<quantity id="4CDD9B753F78FFC6FF78678949386286" box="[217,274,1464,1488]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="56" unit="m" value="12.0">12 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFB44B613F78FFC6FE80678949E96286" author="Randall" box="[289,451,1464,1488]" pageId="20" pageNumber="56" refString="Randall, J. E. (1995) A review of the triplefin fishes (Blennioidei: Tripterygiidae) of Oman, with the description of two new species. Revue francaise d'Aquariologie, 22, 27 - 34." type="journal article" year="1995">Randall, 1995</bibRefCitation>
|
||
), while
|
||
<taxonomicName id="4C254D133F78FFC6FDBD67884ABC6286" box="[540,662,1464,1488]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="pallidus">
|
||
<emphasis id="B951EA823F78FFC6FDBD67884ABC6286" box="[540,662,1464,1488]" italics="true" pageId="20" pageNumber="56">E. pallidus</emphasis>
|
||
</taxonomicName>
|
||
is found on the upper part of reefs, on reef flats, often on corals covered in short algae, in fairly sheltered environments.
|
||
</paragraph>
|
||
<paragraph id="8B9A36903F78FFC6FF66643149DF61D5" blockId="20.[151,1437,1067,1667]" pageId="20" pageNumber="56">
|
||
The odd species is
|
||
<taxonomicName id="4C254D133F78FFC6FE3E643049DD614E" box="[415,503,1536,1560]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="abeli">
|
||
<emphasis id="B951EA823F78FFC6FE3E643049DD614E" box="[415,503,1536,1560]" italics="true" pageId="20" pageNumber="56">E. abeli</emphasis>
|
||
</taxonomicName>
|
||
, in which the males are markedly sexually distinct with the prominent black head, chest and pectoral-fin bases. This species too is widely distributed throughout the Red Sea and down the East African coast to KwaZulu-Natal and Mascarene Islands. The species has a sibling species,
|
||
<taxonomicName id="4C254D133F78FFC6FBC464784CE36136" box="[1125,1225,1608,1632]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="56" phylum="Chordata" rank="species" species="elaine">
|
||
<emphasis id="B951EA823F78FFC6FBC464784CE36136" box="[1125,1225,1608,1632]" italics="true" pageId="20" pageNumber="56">E. elaine</emphasis>
|
||
</taxonomicName>
|
||
, which is endemic to Rodrigues (
|
||
<bibRefCitation id="EFB44B613F78FFC6FE94645A49C361D5" author="Holleman" box="[309,489,1643,1667]" pageId="20" pageNumber="56" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |