195 lines
21 KiB
XML
195 lines
21 KiB
XML
<document id="8F43EEE82B0629A78BFC683B44171105" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter & Bogorodsky, Sergey V." docDate="2012" docId="038C87863F7AFFC4FF3662A64A8461FC" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Norfolkia brachylepis Schultz 1960" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="58" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="4863C9215227182B35371C3E0A1EADC1">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<mods:namePart id="80B38E99A015C6977E75116BA7B82B02">Holleman, Wouter</mods:namePart>
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<mods:namePart id="7C5CAAD1B74E207AED5E6359D8CD23F0">Bogorodsky, Sergey V.</mods:namePart>
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<mods:typeOfResource id="82613D5E522FC2768B95784211E60ED9">text</mods:typeOfResource>
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<mods:title id="EC978B0A59EF1FF26042E4B52BB71619">Zootaxa</mods:title>
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<mods:part id="F19F57A6FFB7AF91838A6C2DB63BCC06">
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<mods:date id="A4B70D8E7FC95AEFA539E56197E651BB">2012</mods:date>
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<mods:detail id="BDE5D359627853A43CD342A0EDB9C9F3" type="volume">
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<mods:number id="3855E77478F22CE1270477D00FECA539">3152</mods:number>
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<treatment id="038C87863F7AFFC4FF3662A64A8461FC" ID-DOI="http://doi.org/10.5281/zenodo.5626542" ID-GBIF-Taxon="119427996" ID-Zenodo-Dep="5626542" LSID="urn:lsid:plazi:treatment:038C87863F7AFFC4FF3662A64A8461FC" httpUri="http://treatment.plazi.org/id/038C87863F7AFFC4FF3662A64A8461FC" lastPageNumber="58" pageId="22" pageNumber="58">
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<subSubSection id="C33F651B3F7AFFC4FF3662A649166784" pageId="22" pageNumber="58" type="nomenclature">
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<paragraph id="8B9A36903F7AFFC4FF3662A64A7467E7" blockId="22.[151,606,151,210]" box="[151,606,151,177]" pageId="22" pageNumber="58">
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<heading id="D0D281FC3F7AFFC4FF3662A64A7467E7" bold="true" box="[151,606,151,177]" fontSize="11" level="1" pageId="22" pageNumber="58" reason="1">
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<taxonomicName id="4C254D133F7AFFC4FF3662A64A7467E7" ID-CoL="47QBD" authority="Schultz, 1960" authorityName="Schultz" authorityYear="1960" box="[151,606,151,177]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="brachylepis">
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<emphasis id="B951EA823F7AFFC4FF3662A64A7467E7" bold="true" box="[151,606,151,177]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF3662A649B467E7" bold="true" box="[151,414,151,177]" italics="true" pageId="22" pageNumber="58">Norfolkia brachylepis</emphasis>
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(Schultz, 1960)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9A36903F7AFFC4FF36628B49166784" blockId="22.[151,606,151,210]" box="[151,316,186,210]" pageId="22" pageNumber="58">
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<figureCitation id="131E2A153F7AFFC4FF36628B48F36784" box="[151,217,186,210]" captionStart="FIGURE 8" captionStartId="19.[151,250,1883,1906]" captionTargetBox="[179,1414,193,1852]" captionTargetId="figure@19.[179,1415,193,1853]" captionTargetPageId="19" captionText="FIGURE 8. (A) Enneapterygius obscurus, Marsa Alam; (B) Enneapterygius pallidus, Farasan Is.; (C) Enneapterygius pusillus, M, Sharm el Sheik; (D) Enneapterygius pusillus, M, Oman; (E) Enneapterygius ventermaculus, Farasan Is.; (E) Helcogramma obtusirostris, M, Sharm el Sheil; (F) Helcogramma obtusirostris, F, Sharm el Sheik; (G) Helcogramma steinitzi, F, Marsa Alam; (H) Norfolkia brachylepis, Bali. Photos by R. Field (B, E), J. E. Randall (D); G. R. Allen (H); S. V. Bogorodsky (the others)." httpUri="https://zenodo.org/record/279581/files/figure.png" pageId="22" pageNumber="58">
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Fig.
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<date id="FF9B10503F7AFFC4FF6B628B48F36784" box="[202,217,186,210]" pageId="22" pageNumber="58">8</date>
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</figureCitation>
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I; table 1
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F7AFFC4FF3663324CCF6600" pageId="22" pageNumber="58" type="reference_group">
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<paragraph id="8B9A36903F7AFFC4FF3663324AEA666E" blockId="22.[151,1253,258,342]" pageId="22" pageNumber="58">
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<treatmentCitationGroup id="AB3511BE3F7AFFC4FF3663324AEA666E" pageId="22" pageNumber="58">
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<taxonomicName id="4C254D133F7AFFC4FF36633249CE664E" ID-CoL="58ZKW" authority="Schultz" authorityName="Schultz" box="[151,484,258,280]" class="Actinopterygii" family="Tripterygiidae" genus="Tripterygion" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="brachylepis">
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<emphasis id="B951EA823F7AFFC4FF36633249B8664E" box="[151,402,258,280]" italics="true" pageId="22" pageNumber="58">Tripterygion brachylepis</emphasis>
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Schultz
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</taxonomicName>
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in Schultz
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<emphasis id="B951EA823F7AFFC4FDF663324AA0664E" box="[599,650,258,280]" italics="true" pageId="22" pageNumber="58">et al.</emphasis>
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, 1960: 291 (
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<typeStatus id="549E88323F7AFFC4FCA863324B1C664E" box="[777,822,259,280]" pageId="22" pageNumber="58">type</typeStatus>
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locality Bikini Atoll,
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<collectingCountry id="F33276003F7AFFC4FBB463334CE8664E" box="[1045,1218,258,280]" name="Marshall Islands" pageId="22" pageNumber="58">Marshall Islands</collectingCountry>
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).
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<taxonomicName id="4C254D133F7AFFC4FF36631049D56661" ID-CoL="47QBL" authority="Clark, 1980: 95" authorityName="Clark" authorityPageNumber="95" authorityYear="1980" box="[151,511,289,311]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="springeri">
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<emphasis id="B951EA823F7AFFC4FF36631049716661" box="[151,347,289,311]" italics="true" pageId="22" pageNumber="58">Norfolkia springeri</emphasis>
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<treatmentCitation id="0A8410813F7AFFC4FEC0631049D56661" author="Clark" box="[353,511,289,311]" page="95" pageId="22" pageNumber="58" year="1980">
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<bibRefCitation id="EFB44B613F7AFFC4FEC0631049F76661" author="Clark" box="[353,477,289,311]" pageId="22" pageNumber="58" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
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: 95
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</treatmentCitation>
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</taxonomicName>
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(Aqaba, Red Sea).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9A36903F7AFFC4FF3663714CCF6600" blockId="22.[151,1253,258,342]" box="[151,1253,320,342]" pageId="22" pageNumber="58">
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<treatmentCitationGroup id="AB3511BE3F7AFFC4FF3663714CCF6600" box="[151,1253,320,342]" pageId="22" pageNumber="58">
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<taxonomicName id="4C254D133F7AFFC4FF3663714A716603" ID-CoL="47QBD" authority="Holleman, 1991: 233" authorityName="Holleman" authorityPageNumber="233" authorityYear="1991" box="[151,603,320,342]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="brachylepis">
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<emphasis id="B951EA823F7AFFC4FF366371495E6600" box="[151,372,320,342]" italics="true" pageId="22" pageNumber="58">Norfolkia brachylepis</emphasis>
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:
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<treatmentCitation id="0A8410813F7AFFC4FE2363714A716603" author="Holleman" box="[386,603,320,342]" page="233" pageId="22" pageNumber="58" year="1991">
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<bibRefCitation id="EFB44B613F7AFFC4FE2363714A026603" author="Holleman" box="[386,552,320,342]" pageId="22" pageNumber="58" refString="Holleman, W. (1991) A revision of the tripterygiid fish genus Norfolkia Fowler, 1953 (Perciformes: Blennioidei). Annals of the Cape Provincial Museums (Natural History), 18 (11), 227 - 243." type="journal article" year="1991">Holleman, 1991</bibRefCitation>
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: 233
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0A8410813F7AFFC4FDC463714B366600" author="Fricke" box="[613,796,320,342]" page="465" pageId="22" pageNumber="58" year="1994">
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<bibRefCitation id="EFB44B613F7AFFC4FDC463714AC36603" author="Fricke" box="[613,745,320,342]" pageId="22" pageNumber="58" refString="Fricke, R. (1994) Tripterygiid Fishes of Australia, New Zealand and the Southwest Pacific Ocean. Koeltz Scientific Books, Koenigstein: ix + 585 pp." type="book" year="1994">Fricke, 1994</bibRefCitation>
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: 465
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</treatmentCitation>
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; 1997: 504;
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<treatmentCitation id="0A8410813F7AFFC4FC3863714CCB6600" author="Golani" box="[921,1249,320,342]" page="45" pageId="22" pageNumber="58" year="2010">
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<bibRefCitation id="EFB44B613F7AFFC4FC3863714C916603" author="Golani" box="[921,1211,320,342]" pageId="22" pageNumber="58" refString="Golani, D. & Bogorodsky, S. V. (2010) The Fishes of the Red Sea-Reappraisal and Updated Checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani & Bogorodsky, 2010</bibRefCitation>
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: 45
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F7AFFC4FF3663B34C376248" pageId="22" pageNumber="58" type="diagnosis">
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<paragraph id="8B9A36903F7AFFC4FF3663B34A326524" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF3663B3493B66CD" bold="true" box="[151,273,386,411]" pageId="22" pageNumber="58">Diagnosis.</emphasis>
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Dorsal fins IV + XIV–XV + 10–11, usually IV + XV + 11; anal fin II, 20–21, usually 21, the spines less than half the length of the rays; pectoral fin 16, upper 2–3 and lower 7 rays simple, remainder divided once; pelvicfin rays united by membrane for less than one-quarter of length of shorter ray. LL 21–24 (usually 22) pored scales ending below junction of second and third dorsal fins, and 14–19 (usually 16) notched scales; total lateral scales 32–35, usually 34. Dentary pores 5–6 + 1 + 5–6. Orbital and nasal cirri large palmate, at least equal to pupil diameter; cirri at tips of first dorsal-fin spines. Head 2.9–3.6 (3.3) in SL, eye 3.0–3.9 (3.3), maxilla 2.3–3.0 (2.6) and snout 3.3–4.9 (3.8) in head length.
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</paragraph>
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<paragraph id="8B9A36903F7AFFC4FF64604E4A806438" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF64604E494965C1" bold="true" box="[197,355,639,663]" pageId="22" pageNumber="58">Fresh colour.</emphasis>
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(from photograph of Rick Winterbottom of specimen from Kwazulu-Natal,
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<collectingCountry id="F33276003F7AFFC4FB6F604F4D4C65C0" box="[1230,1382,638,662]" name="South Africa" pageId="22" pageNumber="58">South Africa</collectingCountry>
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). No sexual dichromatism present other than that males may be more darkly coloured than females. Body pale yellow, scales outlined in yellow-ochre, with 5–6 dark brown blotches between dorsum and midline; head brownish-yellow with darker brown bar below eye and pale area behind, iris yellow; first dorsal fin brown and yellow, second translucent with patches of light red and brown, third dorsal fin with irregular bars of brown-red and cream; caudal fin with irregular brown and white barring, anal fin with yellow and black-brown bars, pelvic fins pale yellow and pectoral fins with irregular brown and yellow bars.
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</paragraph>
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<paragraph id="8B9A36903F7AFFC4FF64614B4D536310" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF64614B498564C5" bold="true" box="[197,431,890,915]" pageId="22" pageNumber="58">Colour underwater</emphasis>
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(from photograph of G.R. Allen of fresh specimen from
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<collectingCountry id="F33276003F7AFFC4FBE4614B4C9264C4" box="[1093,1208,890,914]" name="Indonesia" pageId="22" pageNumber="58">Indonesia</collectingCountry>
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): body grayish with golden-brown and gray-white blotches between dorsum and midline, blotched extending onto second and third dorsal fins; head grey with brown bar below eye, followed by cream bar and light brown preopercle, eye rim alternating brown and grey; first dorsal fin ‘dirty’ white, darker distally, second dorsal fin with white and red-brown marks, the white extending as bands across the fin; third dorsal fin the same; caudal fin with alternating, irregular white and brown bands; anal fin distinctly banded black and cream; pectoral fins irregularly banded with white.
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</paragraph>
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<paragraph id="8B9A36903F7AFFC4FF6466634C376248" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF64666349F1633D" bold="true" box="[197,475,1106,1131]" pageId="22" pageNumber="58">Colour in preservative.</emphasis>
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Body pale gray with five, irregular dark bars along dorsum, which fade towards LL; head gray with a tapering dark bar below the eye and another from the posterior margin of the orbit curving down onto the preopercle. First dorsal fin distally dark gray, second and third dorsal fins with clusters of melanophores basally, continuations of body bars, and which extend as irregular areas of spots across fin to margin; caudal fin with several, narrow dark gray bars; anal fin with faint, diagonal bands of melanophores. Mature males are much darker, with body and median fins dark gray and without any suggestion of bars.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F7AFFC4FF64671A49CC6230" pageId="22" pageNumber="58" type="key">
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<paragraph id="8B9A36903F7AFFC4FF64671A49CC6230" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF64671A49496215" bold="true" box="[197,355,1322,1347]" pageId="22" pageNumber="58">Key features.</emphasis>
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Scaled head, palmate orbital and nasal cirri, four spines in the first dorsal fin and distinctive yellow and black barred anal fin.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F7AFFC4FF64674349836284" pageId="22" pageNumber="58" type="distribution">
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<paragraph id="8B9A36903F7AFFC4FF64674349836284" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF646743497462DD" bold="true" box="[197,350,1394,1419]" pageId="22" pageNumber="58">Distribution.</emphasis>
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This is an apparently wide-spread species, currently known from the Red Sea, to
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<collectingCountry id="F33276003F7AFFC4FAA567434DB362DC" box="[1284,1433,1394,1418]" name="South Africa" pageId="22" pageNumber="58">South Africa</collectingCountry>
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, from
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<collectingCountry id="F33276003F7AFFC4FF7767A7496362F8" box="[214,329,1430,1454]" name="Sri Lanka" pageId="22" pageNumber="58">Sri Lanka</collectingCountry>
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, the east and west coast of
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<collectingCountry id="F33276003F7AFFC4FD2467A74AC562F8" box="[645,751,1430,1454]" name="Australia" pageId="22" pageNumber="58">Australia</collectingCountry>
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,
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<collectingCountry id="F33276003F7AFFC4FD5D67A74B4662F8" box="[764,876,1430,1454]" name="Indonesia" pageId="22" pageNumber="58">Indonesia</collectingCountry>
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, southern
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<collectingCountry id="F33276003F7AFFC4FC4367A64C0862F8" box="[994,1058,1431,1454]" name="Japan" pageId="22" pageNumber="58">Japan</collectingCountry>
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and east to the
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<collectingCountry id="F33276003F7AFFC4FB7867A74DBD62F8" box="[1241,1431,1430,1454]" name="Marshall Islands" pageId="22" pageNumber="58">Marshall Islands</collectingCountry>
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,
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<collectingCountry id="F33276003F7AFFC4FF36678A496E6284" box="[151,324,1466,1490]" name="New Caledonia" pageId="22" pageNumber="58">New Caledonia</collectingCountry>
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and
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<collectingCountry id="F33276003F7AFFC4FEDA678B498C6284" box="[379,422,1466,1490]" name="Fiji" pageId="22" pageNumber="58">Fiji</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F651B3F7AFFC4FF6467EE4A8461FC" pageId="22" pageNumber="58" type="discussion">
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<paragraph id="8B9A36903F7AFFC4FF6467EE4A8461FC" blockId="22.[151,1437,386,1706]" pageId="22" pageNumber="58">
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<emphasis id="B951EA823F7AFFC4FF6467EE491362A1" bold="true" box="[197,313,1503,1527]" pageId="22" pageNumber="58">Remarks.</emphasis>
|
||
<bibRefCitation id="EFB44B613F7AFFC4FEE367EF4A2F62A0" author="Holleman" box="[322,517,1502,1526]" pageId="22" pageNumber="58" refString="Holleman, W. (1991) A revision of the tripterygiid fish genus Norfolkia Fowler, 1953 (Perciformes: Blennioidei). Annals of the Cape Provincial Museums (Natural History), 18 (11), 227 - 243." type="journal article" year="1991">Holleman (1991)</bibRefCitation>
|
||
revised the genus, recognising three species,
|
||
<taxonomicName id="4C254D133F7AFFC4FBB467D14C9162A0" box="[1045,1211,1502,1527]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="brachylepis">
|
||
<emphasis id="B951EA823F7AFFC4FBB467D14C9162A0" box="[1045,1211,1502,1527]" italics="true" pageId="22" pageNumber="58">N. brachylepis</emphasis>
|
||
</taxonomicName>
|
||
, with its very wide distribution,
|
||
<taxonomicName id="4C254D133F7AFFC4FE84643549E2614D" box="[293,456,1540,1563]" family="Tripterygiidae" genus="Gillias" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="squamiceps">
|
||
<emphasis id="B951EA823F7AFFC4FE84643549E2614D" box="[293,456,1540,1563]" italics="true" pageId="22" pageNumber="58">N. squamiceps</emphasis>
|
||
</taxonomicName>
|
||
(southern Great Barrier Reef) and
|
||
<taxonomicName id="4C254D133F7AFFC4FCED64354BEF614D" box="[844,965,1539,1563]" class="Actinopterygii" family="Tripterygiidae" genus="Norfolkia" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="58" phylum="Chordata" rank="species" species="thomasi">
|
||
<emphasis id="B951EA823F7AFFC4FCED64354BEF614D" box="[844,965,1539,1563]" italics="true" pageId="22" pageNumber="58">N. thomasi</emphasis>
|
||
</taxonomicName>
|
||
, from the Great Barrier Reef and southern Pacific to Rapa
|
||
<collectingCountry id="F33276003F7AFFC4FEEA641749B96168" box="[331,403,1574,1598]" name="Iceland" pageId="22" pageNumber="58">Island</collectingCountry>
|
||
. Photographs by J. E. Randall of specimens from
|
||
<collectingCountry id="F33276003F7AFFC4FC6964164C226168" box="[968,1032,1575,1598]" name="Japan" pageId="22" pageNumber="58">Japan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33276003F7AFFC4FBE164174C876168" box="[1088,1197,1574,1598]" name="Indonesia" pageId="22" pageNumber="58">Indonesia</collectingCountry>
|
||
do not show the yellow and black anal fin. The colour described above is of a specimen from southern
|
||
<collectingCountry id="F33276003F7AFFC4FB96647B4CEA6134" box="[1079,1216,1610,1634]" name="Madagascar" pageId="22" pageNumber="58">Madagascar</collectingCountry>
|
||
.
|
||
<collectingCountry id="F33276003F7AFFC4FB6B647A4CC36135" box="[1226,1257,1611,1635]" name="American Samoa" pageId="22" pageNumber="58">As</collectingCountry>
|
||
the species was recorded by
|
||
<bibRefCitation id="EFB44B613F7AFFC4FE82645F49E661D0" author="Randall" box="[291,460,1646,1670]" pageId="22" pageNumber="58" refString="Randall, J. E. (1995) A review of the triplefin fishes (Blennioidei: Tripterygiidae) of Oman, with the description of two new species. Revue francaise d'Aquariologie, 22, 27 - 34." type="journal article" year="1995">Randall (1995)</bibRefCitation>
|
||
from the Gulf of
|
||
<collectingCountry id="F33276003F7AFFC4FD34645F4AF361D0" box="[661,729,1646,1670]" name="Oman" pageId="22" pageNumber="58">Oman</collectingCountry>
|
||
it is assumed here that the colour of the
|
||
<collectingCountry id="F33276003F7AFFC4FB02645F4D0061D0" box="[1187,1322,1646,1670]" name="Madagascar" pageId="22" pageNumber="58">Madagascar</collectingCountry>
|
||
specimen is the same as for individuals from the Red Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |