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<mods:title id="B6D47BF2164BBAE9FAAEE88F453B2599">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<mods:namePart id="2A5D99BC32A381A53C997A035A4FE80E">Holleman, Wouter</mods:namePart>
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<mods:namePart id="0A13410EE2CF9D120B7B518B90C257EC">Bogorodsky, Sergey V.</mods:namePart>
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<treatment id="038C87863F68FFD7FF3666D34CDE64F2" ID-DOI="http://doi.org/10.5281/zenodo.5626514" ID-GBIF-Taxon="119428004" ID-Zenodo-Dep="5626514" LSID="urn:lsid:plazi:treatment:038C87863F68FFD7FF3666D34CDE64F2" httpUri="http://treatment.plazi.org/id/038C87863F68FFD7FF3666D34CDE64F2" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40">
<subSubSection id="C33F651B3F68FFD6FF3666D34988624B" pageId="4" pageNumber="40" type="nomenclature">
<paragraph id="8B9A36903F68FFD6FF3666D34AAF63AA" blockId="4.[151,645,1250,1309]" box="[151,645,1250,1276]" pageId="4" pageNumber="40">
<heading id="D0D281FC3F68FFD6FF3666D34AAF63AA" bold="true" box="[151,645,1250,1276]" fontSize="11" level="1" pageId="4" pageNumber="40" reason="1">
<taxonomicName id="4C254D133F68FFD6FF3666D34AAF63AA" ID-CoL="39VYK" authority="Klausewitz, 1960" authorityName="Klausewitz" authorityYear="1960" box="[151,645,1250,1276]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="40" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F68FFD6FF3666D34AAF63AA" bold="true" box="[151,645,1250,1276]" pageId="4" pageNumber="40">
<emphasis id="B951EA823F68FFD6FF3666D349BD63AA" bold="true" box="[151,407,1250,1276]" italics="true" pageId="4" pageNumber="40">Enneapterygius abeli</emphasis>
(
<bibRefCitation id="EFB44B613F68FFD6FE0766D34A5763AA" author="Klausewitz" box="[422,637,1250,1276]" pageId="4" pageNumber="40" refString="Klausewitz, W. (1960) Fische aus dem Roten Meer. III. Tripterygion abeli n. sp. (Pisces, Blennioidea, Clinidae). Senckenbergiana Biologica, 41 (1 - 2), 11 - 13." type="journal article" year="1960">Klausewitz, 1960</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9A36903F68FFD6FF3667344988624B" blockId="4.[151,645,1250,1309]" box="[151,418,1285,1309]" pageId="4" pageNumber="40">
<figureCitation id="131E2A153F68FFD6FF36673448C9624B" box="[151,227,1285,1309]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="4" pageNumber="40">Figs. 2</figureCitation>
A, 5AC; table 1
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F68FFD6FF3667794A6C62EE" pageId="4" pageNumber="40" type="reference_group">
<paragraph id="8B9A36903F68FFD6FF3667794A72622D" blockId="4.[151,1437,1350,1464]" pageId="4" pageNumber="40">
<treatmentCitationGroup id="AB3511BE3F68FFD6FF3667794A72622D" pageId="4" pageNumber="40">
<taxonomicName id="4C254D133F68FFD6FF3667794A07620A" ID-CoL="58ZKQ" authority="Klausewitz, 1960: 11" authorityName="Klausewitz" authorityPageNumber="11" authorityYear="1960" box="[151,557,1350,1373]" class="Actinopterygii" family="Tripterygiidae" genus="Tripterygion" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="40" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F68FFD6FF3667794965620B" box="[151,335,1351,1373]" italics="true" pageId="4" pageNumber="40">Tripterygion abeli</emphasis>
<treatmentCitation id="0A8410813F68FFD6FEF567774A07620A" author="Klausewitz" box="[340,557,1350,1372]" page="11" pageId="4" pageNumber="40" year="1960">
<bibRefCitation id="EFB44B613F68FFD6FEF567774A23620A" author="Klausewitz" box="[340,521,1350,1372]" pageId="4" pageNumber="40" refString="Klausewitz, W. (1960) Fische aus dem Roten Meer. III. Tripterygion abeli n. sp. (Pisces, Blennioidea, Clinidae). Senckenbergiana Biologica, 41 (1 - 2), 11 - 13." type="journal article" year="1960">Klausewitz, 1960</bibRefCitation>
: 11
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</taxonomicName>
,
<figureCitation id="131E2A153F68FFD6FD9967774ABD620A" box="[568,663,1350,1372]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1157,1180]" captionStartId-1="6.[151,250,1651,1674]" captionTargetBox-0="[164,1424,187,1146]" captionTargetBox-1="[184,1390,372,1603]" captionTargetId-0="figure@4.[164,1424,187,1146]" captionTargetId-1="figure@6.[151,1436,349,1608]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 1. Map of the Red Sea showing localities mentioned in the text." captionText-1="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri-0="https://zenodo.org/record/279574/files/figure.png" httpUri-1="https://zenodo.org/record/279575/files/figure.png" pageId="4" pageNumber="40">Figs. 12</figureCitation>
(Al-Ghardaqa,
<collectingCountry id="F33276003F68FFD6FC9767764B5F620A" box="[822,885,1351,1372]" name="Egypt" pageId="4" pageNumber="40">Egypt</collectingCountry>
, Red Sea).
<taxonomicName id="4C254D133F68FFD6FF3667544A79622D" ID-CoL="6LGHB" authority="Lal Mohan, 1971: 222" authorityName="Lal Mohan" authorityPageNumber="222" authorityYear="1971" box="[151,595,1381,1403]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="40" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F68FFD6FF366754494B622D" box="[151,353,1381,1403]" italics="true" pageId="4" pageNumber="40">Helcogramma abeli</emphasis>
:
<treatmentCitation id="0A8410813F68FFD6FECF67544A79622D" author="Lal" box="[366,595,1381,1403]" page="222" pageId="4" pageNumber="40" year="1971">
<bibRefCitation id="EFB44B613F68FFD6FECF67544A0A622D" author="Lal" box="[366,544,1381,1403]" pageId="4" pageNumber="40" refString="Lal Mohan, R. S. (1971) Helcogramma shinglensis, a new species of tripterygiid fish from the Gulf of Mannar with a key to the fishes of the family Tripterygiidae of the eastern and central Indian Ocean. Senckenbergiana Biologica 52 (3 / 5), 219 - 223." type="journal article" year="1971">Lal Mohan, 1971</bibRefCitation>
: 222
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</taxonomicName>
.
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</paragraph>
<paragraph id="8B9A36903F68FFD6FF3667B44A6C62EE" blockId="4.[151,1437,1350,1464]" pageId="4" pageNumber="40">
<treatmentCitationGroup id="AB3511BE3F68FFD6FF3667B44A6C62EE" pageId="4" pageNumber="40">
<taxonomicName id="4C254D133F68FFD6FF3667B44A3062CF" ID-CoL="39VYK" authority="Clark, 1980: 97" authorityName="Clark" authorityPageNumber="97" authorityYear="1980" box="[151,538,1412,1434]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="40" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F68FFD6FF3667B4494462CC" box="[151,366,1412,1434]" italics="true" pageId="4" pageNumber="40">Enneapterygius abeli</emphasis>
:
<treatmentCitation id="0A8410813F68FFD6FEDA67B54A3062CF" author="Clark" box="[379,538,1412,1434]" page="97" pageId="4" pageNumber="40" year="1980">
<bibRefCitation id="EFB44B613F68FFD6FEDA67B549DF62CF" author="Clark" box="[379,501,1412,1434]" pageId="4" pageNumber="40" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
: 97
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="131E2A153F68FFD6FD8567B54A4062CF" box="[548,618,1412,1434]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="4" pageNumber="40">Figs. 2</figureCitation>
a, 2b, 4c, &amp; 10; Holleman, 1986: 756, Fig. 236.2; 2005: 45,
<figureCitation id="131E2A153F68FFD6FB6F67B54D2162CF" box="[1230,1291,1412,1434]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="4" pageNumber="40">Fig. 2</figureCitation>
;
<treatmentCitation id="0A8410813F68FFD6FAB467B548DA62EE" author="Fricke" page="468" pageId="4" pageNumber="40" year="1999">
<bibRefCitation id="EFB44B613F68FFD6FAB467B54DB262CF" author="Fricke" box="[1301,1432,1412,1434]" pageId="4" pageNumber="40" refString="Fricke, R. (1999) Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). Koeltz Scientific Books, Koenigstein: viii + 759 pp." type="book" year="1999">Fricke, 1999</bibRefCitation>
: 468
</treatmentCitation>
;
<treatmentCitation id="0A8410813F68FFD6FF5B67934A6862EE" author="Golani" box="[250,578,1442,1464]" page="44" pageId="4" pageNumber="40" year="2010">
<bibRefCitation id="EFB44B613F68FFD6FF5B67934A3662EE" author="Golani" box="[250,540,1442,1464]" pageId="4" pageNumber="40" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The Fishes of the Red Sea-Reappraisal and Updated Checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky, 2010</bibRefCitation>
: 44
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F68FFD7FF3667EF48CB66A2" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40" type="diagnosis">
<paragraph id="8B9A36903F68FFD6FF3667EF4CC661D0" blockId="4.[151,1436,1502,2031]" pageId="4" pageNumber="40">
<emphasis id="B951EA823F68FFD6FF3667EF493B62A1" bold="true" box="[151,273,1502,1527]" pageId="4" pageNumber="40">Diagnosis.</emphasis>
Dorsal fins III + XIXIII + 910 (usually III + XII + 10); anal fin I, 1718 (usually 18 rays); pectoralfin 15: usually 3, 5, 7. LL 1214 pored scales and 2023 (usually 22) notched scales; longitudinal scale series 31; dentary pores 3 + 1 + 3 (
<figureCitation id="131E2A153F68FFD6FE0A641749C56168" box="[427,495,1574,1598]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="4" pageNumber="40">Fig. 2</figureCitation>
A). Head
<quantity id="4CDD9B753F68FFD6FDFD64174AE46168" box="[604,718,1574,1598]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="9.652" metricValueMin="8.128" pageId="4" pageNumber="40" unit="in" value="3.5" valueMax="3.8" valueMin="3.2">3.23.8 in</quantity>
SL; eye
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head length. Nape partially scaled, abdomen without scales, single row of scales at base of caudal fin; supratemporal sensory canal crescent-shaped; supraorbital cirrus small and pointed; first dorsal fin about equal in height to second in males, lower in females.
</paragraph>
<paragraph id="8B9A36903F68FFD6FF6464A249B760F0" blockId="4.[151,1436,1502,2031]" pageId="4" pageNumber="40">
<emphasis id="B951EA823F68FFD6FF6464A2494B61FD" bold="true" box="[197,353,1683,1707]" pageId="4" pageNumber="40">Fresh colour.</emphasis>
<emphasis id="B951EA823F68FFD6FEC964A2498761FD" box="[360,429,1683,1707]" italics="true" pageId="4" pageNumber="40">Males</emphasis>
with semi-translucent, deep yellow body, scale margins orange-red; head, chest, and base of pectoral and pelvic fins of males with numerous melanophores (entire head becomes black during courtship); eye with alternating brown and greenish bands; first dorsal-fin membranes white, suffused with yellow and with some melanophores along margin; second dorsal-fin membranes translucent with spines yellow and orange, some small melanophores in patches along margin; third dorsal fin similar with patches of small melanophores about half-way in from margin; caudal fin pale yellow and orange on rays; anal fin lemon yellow with orange spots at base of alternate rays; pectoral-fin rays deep yellow, membranes pale yellow; pelvic fins yellow with some black proximally and at base.
</paragraph>
<paragraph id="8B9A36903F68FFD7FF6465824B3967A1" blockId="4.[151,1436,1502,2031]" lastBlockId="5.[151,1437,151,932]" lastPageId="5" lastPageNumber="41" pageId="4" pageNumber="40">
<emphasis id="B951EA823F68FFD6FF646582490E609D" box="[197,292,1971,1995]" italics="true" pageId="4" pageNumber="40">Females</emphasis>
: body translucent with scale margins orange-brown, forming irregular bars, darkest at caudal-fin base; head and nape brownish, snout and lower half of head white, dark brown stripe from upper lip to anterior margin of eye, vertical orange mark below eye, orange marks on opercle and pectoral-fin base; first dorsal fin greenish with two orange bars on first spine; second and third dorsal fins with oblique banding, colour on rays only, anal and pectoral-fin rays pale yellow, all fin membranes transparent.
</paragraph>
<paragraph id="8B9A36903F69FFD7FF64633248CB66A2" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B951EA823F69FFD7FF646332499E664A" bold="true" box="[197,436,259,284]" pageId="5" pageNumber="41">Colour underwater.</emphasis>
<emphasis id="B951EA823F69FFD7FE1F63354A28664A" box="[446,514,260,284]" italics="true" pageId="5" pageNumber="41">Males</emphasis>
semi-translucent greenish yellow, scale margins orange-brown, head with scattered melanophores to black, eye with orange-brown and green radii, all fins transparent, spines and rays of dorsal and caudal fins translucent, rays of anal and pectoral fins pale yellow, in moderate to faintly pigmented males a brown stripe from corner of eye to upper lip;
<emphasis id="B951EA823F69FFD7FD3D63414ADB66DE" box="[668,753,368,392]" italics="true" pageId="5" pageNumber="41">females</emphasis>
paler than males, top of head greenish and without melanophores, brown stripe from corner of eye to upper lip, head below eye white with cluster of orange spots below eye. Both males and females with vertebral column coloured in alternating brown and white, the white often quite bright.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F69FFD7FF646031493A6536" pageId="5" pageNumber="41" type="key">
<paragraph id="8B9A36903F69FFD7FF646031493A6536" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B951EA823F69FFD7FF646031494F654E" bold="true" box="[197,357,511,536]" pageId="5" pageNumber="41">Key features.</emphasis>
Low first dorsal fin, brown and green radii around eye, males with yellow body and melanophores on head to base of pectoral fins; females with yellow-green body, brown stripe on nose and orange bar below eye.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F69FFD7FF64605A4A026461" pageId="5" pageNumber="41" type="distribution">
<paragraph id="8B9A36903F69FFD7FF64605A4A4D659A" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B951EA823F69FFD7FF64605A497465D2" bold="true" box="[197,350,619,644]" pageId="5" pageNumber="41">Distribution.</emphasis>
Known from the Red Sea, where it is common in the north, the east coast of Africa to as far south as KwaZulu-Natal, the
<collectingCountry id="F33276003F69FFD7FE3F60BE4A3365F1" box="[414,537,655,679]" name="Seychelles" pageId="5" pageNumber="41">Seychelles</collectingCountry>
, southern
<collectingCountry id="F33276003F69FFD7FD2C60BE4B3C65F1" box="[653,790,655,679]" name="Madagascar" pageId="5" pageNumber="41">Madagascar</collectingCountry>
,
<collectingCountry id="F33276003F69FFD7FC8060BE4BAA65F1" box="[801,896,655,679]" name="Réunion" pageId="5" pageNumber="41">Réunion</collectingCountry>
and
<collectingCountry id="F33276003F69FFD7FC1960BE4C0265F1" box="[952,1064,655,679]" name="Mauritius" pageId="5" pageNumber="41">Mauritius</collectingCountry>
, St Brandon Shoals and the Chagos Archipelago; common in some areas.
</paragraph>
<paragraph id="8B9A36903F69FFD7FF6460E64A026461" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B951EA823F69FFD7FF6460E6490265A6" bold="true" box="[197,296,727,752]" pageId="5" pageNumber="41">Habitat.</emphasis>
Shallow, sheltered waters, on hard corals.
<taxonomicName id="4C254D133F69FFD7FCAD60E84B4965A6" box="[780,867,728,752]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F69FFD7FCAD60E84B4965A6" box="[780,867,728,752]" italics="true" pageId="5" pageNumber="41">E. abeli</emphasis>
</taxonomicName>
are found on the upper part of coral reefs, usually along the reef margin in depths of about
<quantity id="4CDD9B753F69FFD7FDD060CD4A8A6442" box="[625,672,764,788]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="41" unit="m" value="1.0">1 m</quantity>
, often on corals covered by low algae. Found in bays or sheltered lagoons.
<taxonomicName id="4C254D133F69FFD7FF5D61104979646E" box="[252,339,800,824]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="41" phylum="Chordata" rank="species" species="abeli">
<emphasis id="B951EA823F69FFD7FF5D61104979646E" box="[252,339,800,824]" italics="true" pageId="5" pageNumber="41">E. abeli</emphasis>
</taxonomicName>
attains
<quantity id="4CDD9B753F69FFD7FE0B612E49D3646E" box="[426,505,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="41" unit="mm" value="25.0">25 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F69FFD7FF6461724CDE64F2" pageId="5" pageNumber="41" type="materials_examined">
<paragraph id="8B9A36903F69FFD7FF6461724CDE64F2" blockId="5.[151,1437,151,932]" pageId="5" pageNumber="41">
<emphasis id="B951EA823F69FFD7FF6461724985640A" bold="true" box="[197,431,835,860]" pageId="5" pageNumber="41">Material examined.</emphasis>
Additional to that listed in
<bibRefCitation id="EFB44B613F69FFD7FD4461754BBD640A" author="Holleman" box="[741,919,836,860]" pageId="5" pageNumber="41" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman 2005</bibRefCitation>
:
<collectionCode id="ED34AE553F69FFD7FC0361754BDC640A" LSID="urn:lsid:biocol.org:col:34984" box="[930,1014,836,860]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="5" pageNumber="41">SAIAB</collectionCode>
88711 (5:
<quantity id="4CDD9B753F69FFD7FBD161754CC0640A" box="[1136,1258,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="5" pageNumber="41" unit="mm" value="18.0" valueMax="21.0" valueMin="15.0">1521 mm</quantity>
SL), Mangrove Bay, El Quseir (
<geoCoordinate id="EE1150573F69FFD7FEF56156499D6429" box="[340,439,871,895]" direction="north" orientation="latitude" pageId="5" pageNumber="41" precision="555" value="25.52">25.52°N</geoCoordinate>
,
<geoCoordinate id="EE1150573F69FFD7FE6461564A0E6429" box="[453,548,871,895]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="555" value="34.24">34.24°E</geoCoordinate>
), collected S. Bogorodsky, 8 &amp;
<date id="FF9B10503F69FFD7FC3261564C186429" box="[915,1074,871,896]" pageId="5" pageNumber="41" value="2008-06-10">10 June 2008</date>
;
<collectionCode id="ED34AE553F69FFD7FBE361564CBC6429" LSID="urn:lsid:biocol.org:col:34984" box="[1090,1174,871,895]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="5" pageNumber="41">SAIAB</collectionCode>
88721 (
<quantity id="4CDD9B753F69FFD7FB5861564D4B64D6" box="[1273,1377,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="5" pageNumber="41" unit="mm" value="18.1">18.1 mm</quantity>
SL), Abingdon Reef,
<collectingCountry id="F33276003F69FFD7FEF061BD49BD64F2" box="[337,407,908,932]" name="Sudan" pageId="5" pageNumber="41">Sudan</collectingCountry>
(
<geoCoordinate id="EE1150573F69FFD7FE0761BD4A2364F2" box="[422,521,908,932]" direction="north" orientation="latitude" pageId="5" pageNumber="41" precision="555" value="20.5">20.50°N</geoCoordinate>
,
<geoCoordinate id="EE1150573F69FFD7FDB261BD4A5864F2" box="[531,626,908,932]" direction="east" orientation="longitude" pageId="5" pageNumber="41" precision="555" value="37.25">37.25°E</geoCoordinate>
), depth
<quantity id="4CDD9B753F69FFD7FD6A61BD4B2C64F2" box="[715,774,908,932]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="41" unit="m" value="14.0">14 m</quantity>
, collected S. Bogorodsky,
<date id="FF9B10503F69FFD7FB9061BD4CDB64F2" box="[1073,1265,908,932]" pageId="5" pageNumber="41" value="2009-10-15">15 October 2009</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>