treatments-xml/data/03/8C/87/038C87863F62FFC2FF3666E34B0E62F3.xml
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<document id="1CA8DB9CCAA8B0D91812DE564F9C55E4" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter &amp; Bogorodsky, Sergey V." docDate="2012" docId="038C87863F62FFC2FF3666E34B0E62F3" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enneapterygius qirmiz Holleman &amp; Bogorodsky, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="52" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="50" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="5BCC04E80A4F77F20E682C98CF9BC917">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<taxonomicName id="4C254D133F62FFDCFF3666E3498063BA" ID-CoL="39W2X" box="[151,426,1234,1260]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="species" species="qirmiz" status="sp. nov.">
<emphasis id="B951EA823F62FFDCFF3666E3498063BA" bold="true" box="[151,426,1234,1260]" italics="true" pageId="14" pageNumber="50">Enneapterygius qirmiz</emphasis>
</taxonomicName>
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H, 6 &amp; 7; table 1
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<taxonomicName id="4C254D133F62FFDCFF366708491C6218" ID-CoL="4BXF" box="[151,310,1337,1358]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B951EA823F62FFDCFF366708491C6218" box="[151,310,1337,1358]" italics="true" pageId="14" pageNumber="50">Enneapterygius</emphasis>
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sp.1
<treatmentCitation id="0A8410813F62FFDCFECD67094A33621B" author="Clark" box="[364,537,1336,1358]" page="101" pageId="14" pageNumber="50" year="1980">
<bibRefCitation id="EFB44B613F62FFDCFECD670949CD621B" author="Clark" box="[364,487,1336,1358]" pageId="14" pageNumber="50" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
: 101
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,
<figureCitation id="131E2A153F62FFDCFD8267094A42621B" box="[547,616,1336,1358]" captionStart="FIGURE 3" captionStartId="7.[151,250,1501,1524]" captionTargetBox="[151,1436,267,1477]" captionTargetId="figure@7.[151,1436,267,1477]" captionTargetPageId="7" captionText="FIGURE 3. Enneapterygius altipinnis, M (A) grey form; (B) green form. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/279576/files/figure.png" pageId="14" pageNumber="50">Figs. 3</figureCitation>
c &amp; 12, Gulf of Aqaba, Ras Muhammad and
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.
<taxonomicName id="4C254D133F62FFDCFF366769491C623B" ID-CoL="4BXF" box="[151,310,1368,1389]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B951EA823F62FFDCFF366769491C623B" box="[151,310,1368,1389]" italics="true" pageId="14" pageNumber="50">Enneapterygius</emphasis>
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sp.:
<treatmentCitation id="0A8410813F62FFDCFEC767674A1C623A" author="Debelius" box="[358,566,1366,1388]" page="174" pageId="14" pageNumber="50" year="1998">
<bibRefCitation id="EFB44B613F62FFDCFEC767674A29623A" author="Debelius" box="[358,515,1366,1388]" pageId="14" pageNumber="50" refString="Debelius, H. (1998) Red Sea Reef Guide. IKAN-Unterwasserarchiv, Frankfurt am Main, 321 pp." type="book" year="1998">Debelius, 1998</bibRefCitation>
: 174
</treatmentCitation>
;
<treatmentCitation id="0A8410813F62FFDCFDE167674B70623A" author="Lieske" box="[576,858,1366,1388]" page="177" pageId="14" pageNumber="50" year="2004">
<bibRefCitation id="EFB44B613F62FFDCFDE167674B0D623A" author="Lieske" box="[576,807,1366,1388]" pageId="14" pageNumber="50" refString="Lieske, E. &amp; Myers, R. (2004) Coral reef guide to the Red Sea. Harper Collins, London, 384 pp." type="book" year="2004">Lieske &amp; Myers, 2004</bibRefCitation>
: 177
</treatmentCitation>
.
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</paragraph>
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<taxonomicName id="4C254D133F62FFDCFF36674749B262DD" ID-CoL="39W2F" box="[151,408,1397,1419]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F62FFDCFF36674749B262DD" box="[151,408,1397,1419]" italics="true" pageId="14" pageNumber="50">Enneapterygius obscurus</emphasis>
</taxonomicName>
non
<bibRefCitation id="EFB44B613F62FFDCFE6B67444A6F62DD" author="Clark" box="[458,581,1397,1419]" pageId="14" pageNumber="50" refString="Clark, E. (1980) Red Sea fishes of the family Tripterygiidae, with descriptions of eight new species. Israel Journal of Zoology, 28 [for 1979], 65 - 113." type="journal article" year="1980">Clark, 1980</bibRefCitation>
:
<treatmentCitation id="0A8410813F62FFDCFDEE67444B3062DD" author="Holleman" box="[591,794,1397,1419]" page="15" pageId="14" pageNumber="50" year="2005">
<bibRefCitation id="EFB44B613F62FFDCFDEE67444ADF62DD" author="Holleman" box="[591,757,1397,1419]" pageId="14" pageNumber="50" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
: 15
</treatmentCitation>
, Fig. 13, Plate 2A.
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</paragraph>
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.
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<collectionCode id="ED34AE553F62FFDCFEB0678849636287" LSID="urn:lsid:biocol.org:col:34838" box="[273,329,1465,1489]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33093, male
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SL; Uqban, Kamaran
<collectingCountry id="F33276003F62FFDCFC9E67884BAC6287" box="[831,902,1465,1489]" name="Iceland" pageId="14" pageNumber="50">Island</collectingCountry>
, Red Sea,
<collectingCountry id="F33276003F62FFDCFC5A678B4C636287" box="[1019,1097,1466,1489]" name="Yemen" pageId="14" pageNumber="50">Yemen</collectingCountry>
(
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,
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), collected by P.C. Heemstra &amp; Uwe Zajonz,
<date id="FF9B10503F62FFDCFD2067EF4B3462A0" box="[641,798,1502,1526]" pageId="14" pageNumber="50" value="1998-04-12">12 April 1998</date>
(ex
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29594).
</paragraph>
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<typeStatus id="549E88323F62FFDCFF6464354915614A" box="[197,319,1540,1564]" pageId="14" pageNumber="50" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED34AE553F62FFDCFEF0643449B5614A" LSID="urn:lsid:biocol.org:col:34697" box="[337,415,1541,1564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="50">BPBM</collectionCode>
35721 (
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SL), Parkin Rock, Hanish Group, southern Red Sea (
<geoCoordinate id="EE1150573F62FFDCFAA264354DBD614A" box="[1283,1431,1540,1564]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="15" value="13.62">13°37'12&quot; N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFF366418490E6117" box="[151,292,1577,1601]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="15" value="42.815">42°48'54&quot; E</geoCoordinate>
), collected by J.E. Randall,
<date id="FF9B10503F62FFDCFDCB64184B2D6117" box="[618,775,1577,1602]" pageId="14" pageNumber="50" value="1993-05-18">18 May 1993</date>
;
<collectionCode id="ED34AE553F62FFDCFCB464184B676117" LSID="urn:lsid:biocol.org:col:34838" box="[789,845,1577,1601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
29357 (11.1 &amp;
<quantity id="4CDD9B753F62FFDCFBA964184C446117" box="[1032,1134,1577,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="14" pageNumber="50" unit="mm" value="15.6">15.6 mm</quantity>
SL), Uqban, east coast of Kamaran
<collectingCountry id="F33276003F62FFDCFEA5647F49616130" box="[260,331,1614,1638]" name="Iceland" pageId="14" pageNumber="50">Island</collectingCountry>
, Red Sea,
<collectingCountry id="F33276003F62FFDCFE1E647E4A276130" box="[447,525,1615,1638]" name="Yemen" pageId="14" pageNumber="50">Yemen</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFDBC647F4A806130" box="[541,682,1614,1638]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="15" value="15.522222">15°31'20&quot; N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFD14647F4B156130" box="[693,831,1614,1638]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="15" value="42.375">42°22'30&quot; E</geoCoordinate>
), collected U. Zajonz,
<date id="FF9B10503F62FFDCFB9A647F4CE66130" box="[1083,1228,1614,1638]" pageId="14" pageNumber="50" value="1998-04-01">1 April 1998</date>
;
<collectionCode id="ED34AE553F62FFDCFB76647F4D256130" LSID="urn:lsid:biocol.org:col:34838" box="[1239,1295,1614,1638]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
29487 (
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SL), same data as
<collectionCode id="ED34AE553F62FFDCFE03644549F061DA" LSID="urn:lsid:biocol.org:col:34838" box="[418,474,1652,1676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
29357;
<collectionCode id="ED34AE553F62FFDCFD9B64454A5861DA" LSID="urn:lsid:biocol.org:col:34838" box="[570,626,1652,1676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
29492 (15.5 &amp;
<quantity id="4CDD9B753F62FFDCFC8F64454BBE61DA" box="[814,916,1652,1676]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="14" pageNumber="50" unit="mm" value="16.5">16.5 mm</quantity>
SL); Kamaran
<collectingCountry id="F33276003F62FFDCFBE864454CBA61DA" box="[1097,1168,1652,1676]" name="Iceland" pageId="14" pageNumber="50">Island</collectingCountry>
,
<collectingCountry id="F33276003F62FFDCFB3C64444CC161DA" box="[1181,1259,1653,1676]" name="Yemen" pageId="14" pageNumber="50">Yemen</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFB5C64454DB261DA" box="[1277,1432,1652,1676]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="15.484616">15°29.077' N</geoCoordinate>
, 42°23'805&quot; E), collected by M. Eisinger,
<date id="FF9B10503F62FFDCFDC764A84B2D61E7" box="[614,775,1689,1713]" pageId="14" pageNumber="50" value="1998-04-10">10 April 1998</date>
;
<collectionCode id="ED34AE553F62FFDCFCB264A84B6161E7" LSID="urn:lsid:biocol.org:col:34838" box="[787,843,1689,1713]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
29594 (
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SL), collected with
<typeStatus id="549E88323F62FFDCFB5264A84D7361E7" box="[1267,1369,1689,1713]" pageId="14" pageNumber="50" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED34AE553F62FFDCFAC564A84DB661E7" LSID="urn:lsid:biocol.org:col:34838" box="[1380,1436,1689,1713]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33588 (
<quantity id="4CDD9B753F62FFDCFF4E648F497D6180" box="[239,343,1726,1750]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="14" pageNumber="50" unit="mm" value="25.3">25.3 mm</quantity>
SL), Al Lith,
<collectingCountry id="F33276003F62FFDCFDA2648F4AB26180" box="[515,664,1726,1750]" name="Saudi Arabia" pageId="14" pageNumber="50">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFD0A648F4B6D6180" box="[683,839,1726,1750]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="20.249416">20°14.965' N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFCF4648F4BC76180" box="[853,1005,1726,1750]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="1" value="39.993683">39°59.621' E</geoCoordinate>
), collected S. Bogorodsky,
<date id="FF9B10503F62FFDCFA8C648F48FB61AA" pageId="14" pageNumber="50" value="2011-03-20">20 March 2011</date>
;
<collectionCode id="ED34AE553F62FFDCFF7F64D5493C61AA" LSID="urn:lsid:biocol.org:col:34838" box="[222,278,1764,1788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33589 (19.0 mm SL), Al Wajh.
<collectingCountry id="F33276003F62FFDCFD2A64D54B3561AA" box="[651,799,1764,1788]" name="Saudi Arabia" pageId="14" pageNumber="50">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFC8E64D54BE361AA" box="[815,969,1764,1788]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="26.175867">26°10.552' N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFC7764D54C4661AA" box="[982,1132,1764,1788]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="1" value="36.374134">36°22.448' E</geoCoordinate>
), collected S. Bogorodsky,
<date id="FF9B10503F62FFDCFF36653849146077" box="[151,318,1801,1825]" pageId="14" pageNumber="50" value="2011-04-10">10 April 2011</date>
;
<collectionCode id="ED34AE553F62FFDCFEEF653849AC6077" LSID="urn:lsid:biocol.org:col:34838" box="[334,390,1801,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33590 (
<quantity id="4CDD9B753F62FFDCFE4B65384A796077" box="[490,595,1801,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="14" pageNumber="50" unit="mm" value="18.1">18.1 mm</quantity>
SL), Al Wajh,
<collectingCountry id="F33276003F62FFDCFCAF65384B8F6077" box="[782,933,1801,1825]" name="Saudi Arabia" pageId="14" pageNumber="50">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFC1965384C7E6077" box="[952,1108,1801,1825]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="26.175867">26°10.552' N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFBC565384CD66077" box="[1124,1276,1801,1825]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="1" value="36.374134">36°22.448' E</geoCoordinate>
), collected S. Bogorodsky,
<date id="FF9B10503F62FFDCFE8D651F49E66010" box="[300,460,1838,1862]" pageId="14" pageNumber="50" value="2011-04-10">10 April 2011</date>
;
<collectionCode id="ED34AE553F62FFDCFE78651F4A3B6010" LSID="urn:lsid:biocol.org:col:34838" box="[473,529,1838,1862]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33591 (
<quantity id="4CDD9B753F62FFDCFDCC651F4AF96010" box="[621,723,1838,1862]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="14" pageNumber="50" unit="mm" value="16.9">16.9 mm</quantity>
SL), Al Wajh,
<collectingCountry id="F33276003F62FFDCFCDE651F4C386010" box="[895,1042,1838,1862]" name="Saudi Arabia" pageId="14" pageNumber="50">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFB83651F4C916010" box="[1058,1211,1838,1862]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="26.175867">26°10.552' N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFB66651F4D716010" box="[1223,1371,1838,1862]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="1" value="36.374134">36°22.448' E</geoCoordinate>
), collected S. Bogorodsky,
<date id="FF9B10503F62FFDCFE3765654A1C603A" box="[406,566,1876,1900]" pageId="14" pageNumber="50" value="2011-04-10">10 April 2011</date>
;
<collectionCode id="ED34AE553F62FFDCFDE265654A51603A" LSID="urn:lsid:biocol.org:col:34838" box="[579,635,1876,1900]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="14" pageNumber="50">SMF</collectionCode>
33592 (20.0 mm SL), Rabigh,
<collectingCountry id="F33276003F62FFDCFC7E65654C58603A" box="[991,1138,1876,1900]" name="Saudi Arabia" pageId="14" pageNumber="50">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFB2365654D36603A" box="[1154,1308,1876,1900]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="1" value="22.785383">22°47.123' N</geoCoordinate>
, 38°
<geoCoordinate id="EE1150573F62FFDCFAED6565488160C7" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="55" value="57.507">57.507 E</geoCoordinate>
), collected S. Bogorodsky, 0
<date id="FF9B10503F62FFDCFE4A65484A5160C7" box="[491,635,1913,1937]" pageId="14" pageNumber="50" value="2011-04-06">6 April 2011</date>
; NHCY-P1 (
<quantity id="4CDD9B753F62FFDCFCA865484B4460C7" box="[777,878,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="50" unit="mm" value="17.5">17.5 mm</quantity>
SL), Kadaman Isl, Kamaran Archipelago,
<collectingCountry id="F33276003F62FFDCFAEF654B4DB660C7" box="[1358,1436,1914,1937]" name="Yemen" pageId="14" pageNumber="50">Yemen</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFF0165AF490460E0" box="[160,302,1950,1974]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="15" value="15.570278">15°34'13&quot; N</geoCoordinate>
, 42°32'156&quot; E), collected by U. Zajonz and P.C. Heemstra,
<date id="FF9B10503F62FFDCFC6965AF4C7060E0" box="[968,1114,1950,1974]" pageId="14" pageNumber="50" value="1998-04-03">3 April 1998</date>
;
<collectionCode id="ED34AE553F62FFDCFBC465AF4C9360E0" LSID="urn:lsid:biocol.org:col:34984" box="[1125,1209,1950,1974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="50">SAIAB</collectionCode>
88720 (14.9 &amp;
<quantity id="4CDD9B753F62FFDCFACA65AF48E9608A" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="50" unit="mm" value="16.4">16.4 mm</quantity>
SL), Shaab Rumi,
<collectingCountry id="F33276003F62FFDCFE3665F549F7608A" box="[407,477,1988,2012]" name="Sudan" pageId="14" pageNumber="50">Sudan</collectingCountry>
(
<geoCoordinate id="EE1150573F62FFDCFE4D65F54A78608A" box="[492,594,1988,2012]" direction="north" orientation="latitude" pageId="14" pageNumber="50" precision="925" value="19.816668">19°49' N</geoCoordinate>
,
<geoCoordinate id="EE1150573F62FFDCFDFC65F54AEA608A" box="[605,704,1988,2012]" direction="east" orientation="longitude" pageId="14" pageNumber="50" precision="925" value="37.35">37°21' E</geoCoordinate>
), depth
<quantity id="4CDD9B753F62FFDCFCB965F54B6F608A" box="[792,837,1988,2012]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="50" unit="m" value="8.0">8 m</quantity>
, S. Bogorodsky,
<date id="FF9B10503F62FFDCFBA365F54CEA608A" box="[1026,1216,1988,2012]" pageId="14" pageNumber="50" value="2009-10-10">10 October 2009</date>
;
<collectionCode id="ED34AE553F62FFDCFB6A65F54D36608A" LSID="urn:lsid:biocol.org:col:34871" box="[1227,1308,1988,2012]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="50">USNM</collectionCode>
205805 (6: 12.018,7 mm SL); Ras Muhammad, Sinai Peninsula, Red Sea, depth
<quantity id="4CDD9B753F63FFDDFC6062A64C2467F9" box="[961,1038,151,175]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="9.0" metricValueMin="0.0" pageId="15" pageNumber="51" unit="m" value="4.5" valueMax="9.0" valueMin="0.0">09 m</quantity>
; collected V. G. Springer
<emphasis id="B951EA823F63FFDDFA9462A84D5A67E6" box="[1333,1392,152,176]" italics="true" pageId="15" pageNumber="51">et al.</emphasis>
,
<date id="FF9B10503F63FFDDFA2162A6497A6783" pageId="15" pageNumber="51" value="1969-09-16">16 September 1969</date>
.
</paragraph>
<paragraph id="8B9A36903F63FFDDFF6462D24A7D6649" blockId="15.[151,1437,151,922]" pageId="15" pageNumber="51">
<emphasis id="B951EA823F63FFDDFF6462D2498C67AD" bold="true" box="[197,422,226,251]" pageId="15" pageNumber="51">Non-type material.</emphasis>
<collectionCode id="ED34AE553F63FFDDFE0D62D34A2A67AC" LSID="urn:lsid:biocol.org:col:34984" box="[428,512,226,250]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="15" pageNumber="51">SAIAB</collectionCode>
96002 (
<quantity id="4CDD9B753F63FFDDFDFA62D34AEA67AC" box="[603,704,226,250]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="15" pageNumber="51" unit="mm" value="16.4">16.4 mm</quantity>
SL), Abingdon Reef,
<collectingCountry id="F33276003F63FFDDFC1762D34BD667AC" box="[950,1020,226,250]" name="Sudan" pageId="15" pageNumber="51">Sudan</collectingCountry>
(
<geoCoordinate id="EE1150573F63FFDDFBAA62D34C4167AC" box="[1035,1131,226,250]" direction="north" orientation="latitude" pageId="15" pageNumber="51" precision="925" value="20.833334">20°50'N</geoCoordinate>
,
<geoCoordinate id="EE1150573F63FFDDFBD762D34CF967AC" box="[1142,1235,226,250]" direction="east" orientation="longitude" pageId="15" pageNumber="51" precision="925" value="37.416668">37°25'E</geoCoordinate>
), depth
<quantity id="4CDD9B753F63FFDDFA8A62D34D4F67AC" box="[1323,1381,226,250]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="51" unit="m" value="14.0">14 m</quantity>
, collected S. Bogorodsky,
<date id="FF9B10503F63FFDDFE3263364A796649" box="[403,595,263,287]" pageId="15" pageNumber="51" value="2009-10-15">15 October 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F63FFDDFF64631C493C66A9" pageId="15" pageNumber="51" type="diagnosis">
<paragraph id="8B9A36903F63FFDDFF64631C493C66A9" blockId="15.[151,1437,151,922]" pageId="15" pageNumber="51">
<emphasis id="B951EA823F63FFDDFF64631C49156610" bold="true" box="[197,319,301,326]" pageId="15" pageNumber="51">Diagnosis.</emphasis>
A small species of
<taxonomicName id="4C254D133F63FFDDFDBD631F4AE66613" box="[540,716,302,325]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="B951EA823F63FFDDFDBD631F4AE66613" box="[540,716,302,325]" italics="true" pageId="15" pageNumber="51">Enneapterygius</emphasis>
</taxonomicName>
with a maximum recorded SL of
<quantity id="4CDD9B753F63FFDDFBE7631C4C846613" box="[1094,1198,301,325]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="15" pageNumber="51" unit="mm" value="25.3">25.3 mm</quantity>
. Dorsal fins III + XII or XIII + 9, anal fin I, 17, pectoral fin 14; first dorsal fin about the same height as the second; nape scaled; both sexes with characteristic oval patch of melanophores on dorsum between third and fourth dorsal-fin spines, males with large rectangular blotch of melanophores and brown pigment on side below posterior of second dorsal fin; red in life, with first dorsal of males crimson in front with golden yellow marks, white behind, base of fin a white patch on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F63FFC2FF64603C4D0465EC" lastPageId="16" lastPageNumber="52" pageId="15" pageNumber="51" type="description">
<paragraph id="8B9A36903F63FFDDFF64603C4C3D6589" blockId="15.[151,1437,151,922]" pageId="15" pageNumber="51">
<emphasis id="B951EA823F63FFDDFF64603C497D6570" bold="true" box="[197,343,525,550]" pageId="15" pageNumber="51">Description.</emphasis>
Dorsal fins III + XII or XIII + 9; anal fin I, 17 (one of 13 with 18 rays); pectoral fin 14 (rarely 15), upper 2 and lower 6 simple, others divided once. (Clark recorded lower and upper 5 unbranched). LL with 810 pored scales followed two scale rows lower by 22 notched scales, some specimens with 8 pored scales followed by 2 notched scales in anterior series; total lateral scales about 33, with single row of scales at base of caudal fin; transverse scales 2/5. Vertebrae 10 precaudal + 23 (2 of 13 with 24) caudal, including urostyle. Dentary pores 2 + 2 + 2 (
<figureCitation id="131E2A153F63FFDDFF6A60F649246589" box="[203,270,711,735]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="15" pageNumber="51">Fig. 2</figureCitation>
H). Head length 3.13.7 (3.4) in SL; eye 2.32.7 (2.6) in head length.
</paragraph>
<paragraph id="8B9A36903F63FFDDFF6460DF4AEB64CC" blockId="15.[151,1437,151,922]" pageId="15" pageNumber="51">Nape scaled, belly naked; eye relatively large, orbital cirrus a rounded flap, its length equal to orbit diameter. First dorsal fin of males about same height as second, a little lower than second in females. Pectoral fins long, longest ray reaching to first spine of third dorsal fin; pelvic-fin rays not united by membrane, longest ray reaching first anal-fin ray. Supratemporal canal a broad, flat U-shape rather than a crescent. Mouth small and pointed, maxilla reaching vertical though anterior margin of pupil.</paragraph>
<caption id="DF5A66183F63FFDDFF3664FF4C1761B3" box="[151,1085,1742,1765]" httpUri="https://zenodo.org/record/279579/files/figure.png" pageId="15" pageNumber="51" targetBox="[151,1436,977,1721]" targetPageId="15">
<paragraph id="8B9A36903F63FFDDFF3664FF4C1761B3" blockId="15.[151,1085,1742,1765]" box="[151,1085,1742,1765]" pageId="15" pageNumber="51">
<emphasis id="B951EA823F63FFDDFF3664FF493861B2" bold="true" box="[151,274,1742,1765]" pageId="15" pageNumber="51">FIGURE 6.</emphasis>
<taxonomicName id="4C254D133F63FFDDFEB864E149D461B3" box="[281,510,1744,1765]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="51" phylum="Chordata" rank="species" species="qirmiz" status="sp. nov.">
<emphasis id="B951EA823F63FFDDFEB864E149D461B3" box="[281,510,1744,1765]" italics="true" pageId="15" pageNumber="51">Enneapterygius qirmiz</emphasis>
</taxonomicName>
<emphasis id="B951EA823F63FFDDFDA464FE4A7F61B2" bold="true" box="[517,597,1743,1764]" pageId="15" pageNumber="51">
<taxonomicNameLabel id="A26257F93F63FFDDFDA464FE4A7F61B2" box="[517,597,1743,1764]" pageId="15" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, M, Angarosh Reef. Photo by S.V. Bogorodsky.
</paragraph>
</caption>
<paragraph id="8B9A36903F63FFC2FF6465204CBE668C" blockId="15.[151,1436,1809,2020]" lastBlockId="16.[151,1437,151,1445]" lastPageId="16" lastPageNumber="52" pageId="15" pageNumber="51">
<emphasis id="B951EA823F63FFDDFF6465204937607C" bold="true" box="[197,285,1809,1834]" pageId="15" pageNumber="51">Colour.</emphasis>
<emphasis id="B951EA823F63FFDDFE8665234946607C" box="[295,364,1810,1834]" italics="true" pageId="15" pageNumber="51">Males</emphasis>
(from underwater photographs by Helmut Debelius and the second author) body translucent, scales with double, crimson-red line on edge, except for black borders to scales immediately below end of second dorsal fin forming an indistinct narrow, oblique bar, anterior to which is large, rectangular patch of scattered melanophores extending from just below the dorsal fin almost to the base of the anal fin. Some specimens with another cluster of scattered melanophores anterior to main blotch. Nape and opercle deep crimson; deep crimson blotch on mid-side below first two or three spines of second dorsal fin; crimson blotches along vertebral column and along base of anal fin; preanal triangular cluster of melanophores present in large specimens; head red above and pale below, iris crimson, with a thin whitish ring around the iris, and with white patches behind and between eyes, orbital cirri yellow, nasal cirri red, red on lips. First dorsal fin with a broad, inverted deep red triangle, apex at base of first spine, with golden yellow marks, remainder of fin white as is base of fin and a narrow saddle extending from base of first spine nearly to base of fourth spine; second dorsal-fin membranes transparent with red on spines, and with black and white spots along margin and at base, and a narrow line of melanophores along margin which continues onto body as black spots on scale margins; third dorsal fin similar; caudal fin with crimson lines on edges of rays; anal fin rays crimson and patches of black and red spots on membranes; pectoral-fin base with patch of crimson spots anteriorly and white spots below, rays red with patches of bright white spots.
</paragraph>
<paragraph id="8B9A36903F7CFFC2FF6463D64A9A6573" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
Live colour of
<emphasis id="B951EA823F7CFFC2FED263D949E26556" box="[371,456,488,512]" italics="true" pageId="16" pageNumber="52">females</emphasis>
unknown, but probably also crimson, but without the blotch of melanophores on the side and conspicuously coloured first dorsal fin.
</paragraph>
<paragraph id="8B9A36903F7CFFC2FF6460034D0465EC" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">Body of preserved fishes pale yellow, males with large blotch of melanophores on side below pectoral fin, line of scale pockets outlined in black extending from posterior end of third dorsal fin to midline and oval patch of melanophores on dorsum between third and fourth dorsal-fin spines, patch of melanophores on peduncle just behind end of third dorsal fin; small pre-anal patch of melanophores; some clusters of melanophores on anal fin.</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F7CFFC2FF6460F94A2A6419" pageId="16" pageNumber="52" type="key">
<paragraph id="8B9A36903F7CFFC2FF6460F94A2A6419" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
<emphasis id="B951EA823F7CFFC2FF6460F9494E65B6" bold="true" box="[197,356,711,736]" pageId="16" pageNumber="52">Key features.</emphasis>
Translucent crimson; first dorsal fin lower than second, red anteriorly with yellow marks, white posteriorly and with bright white patch on dorsum at base of fin; narrow line of melanophores along margin of second dorsal fin which continues onto body as black spots on scale margins; males with broad patch of melanophores on side below second dorsal fin.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F7CFFC2FF64616C4C4F632C" pageId="16" pageNumber="52" type="etymology">
<paragraph id="8B9A36903F7CFFC2FF64616C49F164CC" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
<emphasis id="B951EA823F7CFFC2FF64616C49666423" bold="true" box="[197,332,861,885]" pageId="16" pageNumber="52">Etymology.</emphasis>
Qirmiz is the Arabic word for deep red or crimson, and is given for the colour of the species. It is used as a noun in apposition.
</paragraph>
<paragraph id="8B9A36903F7CFFC2FF6461964C4F632C" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
<emphasis id="B951EA823F7CFFC2FF64619649416496" bold="true" box="[197,363,935,960]" pageId="16" pageNumber="52">Comparisons.</emphasis>
<collectingCountry id="F33276003F7CFFC2FED3619949BB6496" box="[370,401,936,960]" name="American Samoa" pageId="16" pageNumber="52">As</collectingCountry>
can be seen from
<tableCitation id="C6A7032B3F7CFFC2FDC561964A9264E9" box="[612,696,935,959]" captionStart="TABLE 1" captionStartId="2.[151,235,154,175]" captionTargetBox="[151,1401,203,996]" captionTargetPageId="2" captionText="TABLE 1. Comparative characters for Red Sea tripterygiid species." httpUri="http://table.plazi.org/id/DF5A66183F6EFFD0FF3662AB4B6C67F9" pageId="16" pageNumber="52" tableUuid="DF5A66183F6EFFD0FF3662AB4B6C67F9">Table 1</tableCitation>
, there is little variation in morphometric characters between species of
<taxonomicName id="4C254D133F7CFFC2FF4B61FF49B064B3" box="[234,410,974,997]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B951EA823F7CFFC2FF4B61FF49B064B3" box="[234,410,974,997]" italics="true" pageId="16" pageNumber="52">Enneapterygius</emphasis>
</taxonomicName>
. Three species tend to have 17 anal-fin rays:
<taxonomicName id="4C254D133F7CFFC2FC0061FF4C0164B3" box="[929,1067,973,997]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="altipinnis">
<emphasis id="B951EA823F7CFFC2FC0061FF4C0164B3" box="[929,1067,973,997]" italics="true" pageId="16" pageNumber="52">E. altipinnis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D133F7CFFC2FB9A61FF4CEA64B3" box="[1083,1216,973,997]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F7CFFC2FB9A61FF4CEA64B3" box="[1083,1216,973,997]" italics="true" pageId="16" pageNumber="52">E. obscurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D133F7CFFC2FB5861FF4D4B64B3" box="[1273,1377,974,997]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="qirmiz">
<emphasis id="B951EA823F7CFFC2FB5861FF4D4B64B3" box="[1273,1377,974,997]" italics="true" pageId="16" pageNumber="52">E. qirmiz</emphasis>
</taxonomicName>
. The first has a tall first dorsal fin and the characteristic black blotch on the second dorsal fin; the last two species are both red in colour, but males of
<taxonomicName id="4C254D133F7CFFC2FDA666284A456366" box="[519,623,1049,1072]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="qirmiz">
<emphasis id="B951EA823F7CFFC2FDA666284A456366" box="[519,623,1049,1072]" italics="true" pageId="16" pageNumber="52">E. qirmiz</emphasis>
</taxonomicName>
have a very characteristic red, white and yellow first dorsal fin, and
<taxonomicName id="4C254D133F7CFFC2FA25662848D66303" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F7CFFC2FA25662848D66303" italics="true" pageId="16" pageNumber="52">E. obscurus</emphasis>
</taxonomicName>
an intense black spot on the upper caudal peduncle. These two species can also be separated on LL scale counts,
<taxonomicName id="4C254D133F7CFFC2FF4F6655497F632D" box="[238,341,1124,1147]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="qirmiz">
<emphasis id="B951EA823F7CFFC2FF4F6655497F632D" box="[238,341,1124,1147]" italics="true" pageId="16" pageNumber="52">E. qirmiz</emphasis>
</taxonomicName>
with 9 pored scales in the anterior series and
<taxonomicName id="4C254D133F7CFFC2FCF566554BF2632D" box="[852,984,1123,1147]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F7CFFC2FCF566554BF2632D" box="[852,984,1123,1147]" italics="true" pageId="16" pageNumber="52">E. obscurus</emphasis>
</taxonomicName>
with 1012.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F7CFFC2FF6466B64AAA63BC" pageId="16" pageNumber="52" type="distribution">
<paragraph id="8B9A36903F7CFFC2FF6466B64AAA63BC" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
<emphasis id="B951EA823F7CFFC2FF6466B6497463F6" bold="true" box="[197,350,1159,1184]" pageId="16" pageNumber="52">Distribution.</emphasis>
Clarks original specimens were collected in the Gulf of Aqaba, at Ras Mohammad and
<collectingCountry id="F33276003F7CFFC2FA9766B64DB363C9" box="[1334,1433,1159,1183]" name="Ethiopia" pageId="16" pageNumber="52">Ethiopia</collectingCountry>
. The
<typeStatus id="549E88323F7CFFC2FF6B669F492F6390" box="[202,261,1198,1222]" pageId="16" pageNumber="52">types</typeStatus>
here described were collected in similar localities as well as off
<collectingCountry id="F33276003F7CFFC2FC43669F4C196393" box="[994,1075,1198,1221]" name="Yemen" pageId="16" pageNumber="52">Yemen</collectingCountry>
, suggesting the species is widespread, though not common in the Red Sea.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F7CFFC2FF6466C94B0E62F3" pageId="16" pageNumber="52" type="discussion">
<paragraph id="8B9A36903F7CFFC2FF6466C94B0E62F3" blockId="16.[151,1437,151,1445]" pageId="16" pageNumber="52">
<emphasis id="B951EA823F7CFFC2FF6466C949136246" bold="true" box="[197,313,1272,1296]" pageId="16" pageNumber="52">Remarks.</emphasis>
Debelius photographs in
<emphasis id="B951EA823F7CFFC2FDC466C94B6D6246" box="[613,839,1272,1296]" italics="true" pageId="16" pageNumber="52">Red Sea Reef Guide</emphasis>
(1998) were incorrectly identified as
<taxonomicName id="4C254D133F7CFFC2FB5266C84D526246" box="[1267,1400,1272,1296]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F7CFFC2FB5266C84D526246" box="[1267,1400,1272,1296]" italics="true" pageId="16" pageNumber="52">E. obscurus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB44B613F7CFFC2FF36672C49706263" author="Holleman" box="[151,346,1309,1333]" pageId="16" pageNumber="52" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman (2005)</bibRefCitation>
. The species is found in environments very similar to those of
<taxonomicName id="4C254D133F7CFFC2FBB4672F4CB36263" box="[1045,1177,1309,1333]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B951EA823F7CFFC2FBB4672F4CB36263" box="[1045,1177,1309,1333]" italics="true" pageId="16" pageNumber="52">E. obscurus</emphasis>
</taxonomicName>
. However, in comparison with other
<taxonomicName id="4C254D133F7CFFC2FEE2677549D9620D" box="[323,499,1348,1371]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B951EA823F7CFFC2FEE2677549D9620D" box="[323,499,1348,1371]" italics="true" pageId="16" pageNumber="52">Enneapterygius</emphasis>
</taxonomicName>
spp. which are mostly known from shallow coastal waters, this species is surprisingly common on some isolated reefs far from the coast, for example, on the isolated Abingdon Reef located about
<quantity id="4CDD9B753F7CFFC2FF3667BC48F462F3" box="[151,222,1421,1445]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="52" unit="km" value="20.0">20 km</quantity>
from the coast, where no other triplefins were seen.
</paragraph>
</subSubSection>
</treatment>
</document>