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<document id="A2B4525C561BC00B3C294F177D95DA9E" ID-CLB-Dataset="45708" ID-DOI="10.5281/zenodo.279573" ID-GBIF-Dataset="7359830e-3896-4dc8-91bc-1c15c4ee2993" ID-ISSN="1175-5326" ID-Zenodo-Dep="279573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460423643735" checkinUser="plazi" docAuthor="Holleman, Wouter &amp; Bogorodsky, Sergey V." docDate="2012" docId="038C87863F61FFDCFF3661C34A99632C" docLanguage="en" docName="zt03152p060.pdf" docOrigin="Zootaxa 3152" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enneapterygius pusillus Ruppell 1835" docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="FFB5FFFE3F6CFFD2FFA16231482A6756" masterDocTitle="A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980" masterLastPageNumber="60" masterPageNumber="36" pageNumber="49" updateTime="1698330352715" updateUser="ExternalLinkService">
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<mods:title id="905A362D5282F003D9AC4B006E9B632F">A review of the blennioid fish family Tripterygiidae (Perciformes) in the Red Sea, with description of Enneapterygius qirmiz, and reinstatement of Enneapterygius altipinnis Clark, 1980</mods:title>
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<mods:namePart id="CA3050589ECDCF74AF4F4DC15C2E409E">Holleman, Wouter</mods:namePart>
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<mods:namePart id="DA01AB24EA0399BC4A8C193593AAC21D">Bogorodsky, Sergey V.</mods:namePart>
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<mods:date id="ADB8470D42E3BE11ADBEA4F176E35B76">2012</mods:date>
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<treatment id="038C87863F61FFDCFF3661C34A99632C" ID-DOI="http://doi.org/10.5281/zenodo.5626528" ID-GBIF-Taxon="119428008" ID-Zenodo-Dep="5626528" LSID="urn:lsid:plazi:treatment:038C87863F61FFDCFF3661C34A99632C" httpUri="http://treatment.plazi.org/id/038C87863F61FFDCFF3661C34A99632C" lastPageId="14" lastPageNumber="50" pageId="13" pageNumber="49">
<subSubSection id="C33F651B3F61FFDFFF3661C34B5C6338" pageId="13" pageNumber="49" type="nomenclature">
<paragraph id="8B9A36903F61FFDFFF3661C34A44635A" blockId="13.[151,622,1010,1069]" box="[151,622,1010,1036]" pageId="13" pageNumber="49">
<heading id="D0D281FC3F61FFDFFF3661C34A44635A" bold="true" box="[151,622,1010,1036]" fontSize="11" level="1" pageId="13" pageNumber="49" reason="1">
<taxonomicName id="4C254D133F61FFDFFF3661C34A44635A" ID-CoL="6FBJQ" authority="Ruppell, 1835" authorityName="Ruppell" authorityYear="1835" box="[151,622,1010,1036]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="49" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F61FFDFFF3661C34A44635A" bold="true" box="[151,622,1010,1036]" pageId="13" pageNumber="49">
<emphasis id="B951EA823F61FFDFFF3661C34996635A" bold="true" box="[151,444,1010,1036]" italics="true" pageId="13" pageNumber="49">Enneapterygius pusillus</emphasis>
<bibRefCitation id="EFB44B613F61FFDFFE6361C34A44635A" author="Ruppell" box="[450,622,1010,1036]" pageId="13" pageNumber="49" refString="Ruppell, E. (1835 - 1838) Fische des Rothen Meeres. Neue Wirbelthiere der Fauna Abyssiniens. Frankfurt am Main, [1835, 1 - 28, pls. 1 - 7]." type="book chapter" year="1835">Rüppell, 1835</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9A36903F61FFDFFF3666244980637B" blockId="13.[151,622,1010,1069]" box="[151,426,1045,1069]" pageId="13" pageNumber="49">
<figureCitation id="131E2A153F61FFDFFF36662448FD637B" box="[151,215,1045,1069]" captionStart="FIGURE 2" captionStartId="6.[151,250,1651,1674]" captionTargetBox="[184,1390,372,1603]" captionTargetId="figure@6.[151,1436,349,1608]" captionTargetPageId="6" captionText="FIGURE 2. Dentary pore patterns of Red Sea species of Enneapterygius: (A) E. abeli; (B) E. altipinnis; (C) E. clarkae; (D) E. destai; (E) E. obscurus; (F) E. pallidus (diagrammatic - J. T. Williams); (G) E. pusillus; (H) E. qirmiz; (I) E. ventermaculus." httpUri="https://zenodo.org/record/279575/files/figure.png" pageId="13" pageNumber="49">Fig. 2</figureCitation>
G, 8C &amp; D; table 1
</paragraph>
<paragraph id="8B9A36903F61FFDFFF3666684B5C6338" blockId="13.[151,1436,1112,1256]" box="[151,886,1112,1134]" pageId="13" pageNumber="49">
<taxonomicName id="4C254D133F61FFDFFF3666684A0E6338" authority="Ruppell, 1935" authorityName="Ruppell" authorityYear="1935" box="[151,548,1112,1134]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="49" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F61FFDFFF36666849A16338" box="[151,395,1112,1134]" italics="true" pageId="13" pageNumber="49">Enneapterygius pusillus</emphasis>
Rüppell, 1935
</taxonomicName>
: (original description, Massawa).
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F61FFDFFF3666494A5963BE" pageId="13" pageNumber="49" type="reference_group">
<paragraph id="8B9A36903F61FFDFFF3666494A4363DA" blockId="13.[151,1436,1112,1256]" box="[151,617,1142,1165]" pageId="13" pageNumber="49">
<treatmentCitationGroup id="AB3511BE3F61FFDFFF3666494A4363DA" box="[151,617,1142,1165]" pageId="13" pageNumber="49">
<taxonomicName id="4C254D133F61FFDFFF3666494A4363DA" authority="Klunzinger, 1871: 497" authorityName="Klunzinger" authorityPageNumber="497" authorityYear="1871" box="[151,617,1142,1165]" class="Magnoliopsida" family="Celastraceae" genus="Tripterygium" kingdom="Animalia" order="Celastrales" pageId="13" pageNumber="49" phylum="Tracheophyta" rank="species" species="pusillum">
<emphasis id="B951EA823F61FFDFFF366649495263DB" box="[151,376,1143,1165]" italics="true" pageId="13" pageNumber="49">Tripterygium pusillum</emphasis>
:
<treatmentCitation id="0A8410813F61FFDFFE2466474A4363DA" author="Klunzinger" box="[389,617,1142,1164]" page="497" pageId="13" pageNumber="49" year="1871">
<bibRefCitation id="EFB44B613F61FFDFFE2466474A1363DA" author="Klunzinger" box="[389,569,1142,1164]" pageId="13" pageNumber="49" refString="Klunzinger, C. B. (1871) Synopsis der Fische de Rothen Meeres. Verhandellungen der Kaiserlich-Konichlichen Zoologish-botanischen Gesellschaft in Wien, 21, 441 - 688." type="journal article" year="1871">Klunzinger, 1871</bibRefCitation>
: 497
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</taxonomicName>
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</paragraph>
<paragraph id="8B9A36903F61FFDFFF3666A7490D639F" blockId="13.[151,1436,1112,1256]" pageId="13" pageNumber="49">
<treatmentCitationGroup id="AB3511BE3F61FFDFFF3666A7490D639F" pageId="13" pageNumber="49">
<taxonomicName id="4C254D133F61FFDFFF3666A74A6363FD" ID-CoL="6FBJQ" authority="Smith, 1949: 359" authorityName="Smith" authorityPageNumber="359" authorityYear="1949" box="[151,585,1173,1195]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="49" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F61FFDFFF3666A749A063FD" box="[151,394,1173,1195]" italics="true" pageId="13" pageNumber="49">Enneapterygius pusillus</emphasis>
:
<treatmentCitation id="0A8410813F61FFDFFE3966A44A6363FD" author="Smith" box="[408,585,1173,1195]" page="359" pageId="13" pageNumber="49" year="1949">
<bibRefCitation id="EFB44B613F61FFDFFE3966A44A3D63FD" author="Smith" box="[408,535,1173,1195]" pageId="13" pageNumber="49" refString="Smith, J. L. B. (1949) The Sea Fishes of Southern Africa. Central News Agency Ltd., South Africa, xvi + 550 pp, 111 pls." type="book" year="1949">Smith, 1949</bibRefCitation>
: 359
</treatmentCitation>
</taxonomicName>
; Holleman in
<bibRefCitation id="EFB44B613F61FFDFFD7666A44BFD63FD" author="Smith" box="[727,983,1173,1195]" pageId="13" pageNumber="49" refString="Smith, M. M. &amp; Heemstra, P. C. (Eds.) (1986) Smiths' Sea Fishes. Macmillan, Johannesburg, xx + 1047 pp. 144 pls." type="book" year="1986">Smith &amp; Heemstra, 1986</bibRefCitation>
:
<treatmentCitation id="0A8410813F61FFDFFC4166A44C8663FD" author="Holleman" box="[992,1196,1173,1195]" page="19" pageId="13" pageNumber="49" year="2005">
<bibRefCitation id="EFB44B613F61FFDFFC4166A44CAD63FD" author="Holleman" box="[992,1159,1173,1195]" pageId="13" pageNumber="49" refString="Holleman, W. (2005) A review of the triplefin fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the Western Indian Ocean, with descriptions of four new species. Smithiana Bulletin, 5, 27 pp, 2 pls." type="book" year="2005">Holleman, 2005</bibRefCitation>
: 19
</treatmentCitation>
;
<treatmentCitation id="0A8410813F61FFDFFB1666A44909639F" author="Golani" page="44" pageId="13" pageNumber="49" year="2010">
<bibRefCitation id="EFB44B613F61FFDFFB1666A448D7639F" author="Golani" pageId="13" pageNumber="49" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The Fishes of the Red Sea-Reappraisal and Updated Checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky, 2010</bibRefCitation>
: 44
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A36903F61FFDFFF3666E34A5963BE" blockId="13.[151,1436,1112,1256]" box="[151,627,1234,1256]" pageId="13" pageNumber="49">
<treatmentCitationGroup id="AB3511BE3F61FFDFFF3666E34A5963BE" box="[151,627,1234,1256]" pageId="13" pageNumber="49">
<taxonomicName id="4C254D133F61FFDFFF3666E34A4563BE" ID-CoL="3K47R" authority="Lal Mohan, 1971: 222" authorityName="Lal Mohan" authorityPageNumber="222" authorityYear="1971" box="[151,623,1234,1256]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="49" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F61FFDFFF3666E3495763BE" box="[151,381,1234,1256]" italics="true" pageId="13" pageNumber="49">Helcogramma pusillus</emphasis>
:
<treatmentCitation id="0A8410813F61FFDFFE2B66E34A4563BE" author="Lal" box="[394,623,1234,1256]" page="222" pageId="13" pageNumber="49" year="1971">
<bibRefCitation id="EFB44B613F61FFDFFE2B66E34A1663BE" author="Lal" box="[394,572,1234,1256]" pageId="13" pageNumber="49" refString="Lal Mohan, R. S. (1971) Helcogramma shinglensis, a new species of tripterygiid fish from the Gulf of Mannar with a key to the fishes of the family Tripterygiidae of the eastern and central Indian Ocean. Senckenbergiana Biologica 52 (3 / 5), 219 - 223." type="journal article" year="1971">Lal Mohan, 1971</bibRefCitation>
: 222
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F61FFDCFF3667234A54647C" lastPageId="14" lastPageNumber="50" pageId="13" pageNumber="49" type="diagnosis">
<paragraph id="8B9A36903F61FFDFFF366723497462B3" blockId="13.[151,1436,1298,2032]" pageId="13" pageNumber="49">
<emphasis id="B951EA823F61FFDFFF366723493B627D" bold="true" box="[151,273,1298,1323]" pageId="13" pageNumber="49">Diagnosis.</emphasis>
Dorsal fins III + XIIXIV (usually XIII) + 1011; anal fin I, 2021; pectoral fin 1315 (usually 14), some rays branched; LL 912 pored scales and 2528 notched scales; longitudinal scale series 2930; nape and abdomen naked; supratemporal sensory canal deeply U-shaped; dentary pores 2 + 2 + 2; eye diameter 3.0
<quantity id="4CDD9B753F61FFDFFAFD676C4DB66223" box="[1372,1436,1373,1397]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="13" pageNumber="49" unit="in" value="3.4">3.4 in</quantity>
head length; supraorbital cirrus small and lobate; body depth 5.36.0 in standard length; first dorsal fin about equal in height to second fin in females and distinctly higher than second in males, sometimes more than twice height of second dorsal fin.
</paragraph>
<paragraph id="8B9A36903F61FFDFFF6467C24B626193" blockId="13.[151,1436,1298,2032]" pageId="13" pageNumber="49">
<emphasis id="B951EA823F61FFDFFF6467C2494B615D" bold="true" box="[197,353,1523,1547]" pageId="13" pageNumber="49">Fresh colour.</emphasis>
Another species in which mature / sexually active males are much more intensely coloured than other males, coupled with a much taller first dorsal fin. Underwater or photographed against a white background fish have largely transparent bodies and fins, colour being concentrated on the head and chest. Against a coloured background these fish have a white body against which pigmentation patterns stand out more clearly. Females are almost entirely transparent against a white background or underwater, with some pigment on the head, a little on body and fins, and the viscera encased in a silver peritoneum.
</paragraph>
<paragraph id="8B9A36903F61FFDCFF6464E349CD663C" blockId="13.[151,1436,1298,2032]" lastBlockId="14.[151,1437,151,1146]" lastPageId="14" lastPageNumber="50" pageId="13" pageNumber="49">
Body of
<emphasis id="B951EA823F61FFDFFE8C64E2494561BD" box="[301,367,1747,1771]" italics="true" pageId="13" pageNumber="49">males</emphasis>
translucent, pale cream with four irregular darkish bars; scales on upper half with scattered orange and yellow-green spots and bordered with a thin, deep orange line. Lower half of body with up to 10 irregular orange blotches interspersed with silver-grey spots. Abdomen covered anteriorly with dark brown rosettes and posteriorly with pale red blotches; dark brown or black pre-anal spot — often V-shaped — usually present. Top of head laced with subcutaneous green and cream, with red surface spots. Upper margin of eye and orbital cirrus speckled with crimson, iris golden-red. Snout and lips pale yellow with orange spots; cheeks with yellow, red and black spots; iridescent blue spot below eye. First dorsal fin orange-yellow with many small pale red spots and melanophores; second dorsal fin heavily marked with dark brown to black melanophores interspersed with red spots and with 3 silvery-transparent vertical bands, and red and orange marks along base of fin; third dorsal fin with three reddish bands, colour on rays only and scattered melanophores on margin. Upper half of caudal fin pale orange, lower half pale yellowish green. Anal fin with red and black spots, tending to form 6 or more indistinct bars. Pectoral fins yellow distally, orange proximally, with large circular orange-red mark with green centre on fin base. Pelvic fin bases and surrounding skin heavily mottled with deep red and black, rays black for proximal half, distally with red stipples; membrane black.
</paragraph>
<paragraph id="8B9A36903F62FFDCFF6463494A436539" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF646349490E66C6" box="[197,292,376,400]" italics="true" pageId="14" pageNumber="50">Females</emphasis>
less intensely pigmented and lack the melanin of large males, but with an olive streak on side of snout, broadening onto lips; membrane between first two dorsal-fin spines olive green, the first spine banded in pale green and orange, membrane between second and third spines whitish, and between third spine and dorsum transparent; second and third dorsal fins transparent with some olive green spots anteriorly on the first dorsal fin, spines and rays with orange bands; caudal-fin rays with pale orange bands; anal fin transparent, but with some rays orange proximally, colour continuous with darker spots at base of rays; pectoral fins white at base and with band of white marks on rays; pelvic fins without colour.
</paragraph>
<paragraph id="8B9A36903F62FFDCFF64604C4CCD6589" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF64604C499965C3" bold="true" box="[197,435,637,662]" pageId="14" pageNumber="50">Colour underwater.</emphasis>
Body of
<emphasis id="B951EA823F62FFDCFD82604C4A4E65C3" box="[547,612,637,661]" italics="true" pageId="14" pageNumber="50">males</emphasis>
(
<figureCitation id="131E2A153F62FFDCFDD7604C4A9065C3" box="[630,698,637,661]" captionStart="FIGURE 8" captionStartId="19.[151,250,1883,1906]" captionTargetBox="[179,1414,193,1852]" captionTargetId="figure@19.[179,1415,193,1853]" captionTargetPageId="19" captionText="FIGURE 8. (A) Enneapterygius obscurus, Marsa Alam; (B) Enneapterygius pallidus, Farasan Is.; (C) Enneapterygius pusillus, M, Sharm el Sheik; (D) Enneapterygius pusillus, M, Oman; (E) Enneapterygius ventermaculus, Farasan Is.; (E) Helcogramma obtusirostris, M, Sharm el Sheil; (F) Helcogramma obtusirostris, F, Sharm el Sheik; (G) Helcogramma steinitzi, F, Marsa Alam; (H) Norfolkia brachylepis, Bali. Photos by R. Field (B, E), J. E. Randall (D); G. R. Allen (H); S. V. Bogorodsky (the others)." httpUri="https://zenodo.org/record/279581/files/figure.png" pageId="14" pageNumber="50">Fig. 8</figureCitation>
D) mottled dark green with splashes of brown and white, scales edged in brown, all fin elements greenish gold, except first dorsal fin which is golden yellow with black stippling, and pelvic fins which are black; head reddish with many melanophores, snout grey, eye ridges red.
</paragraph>
<paragraph id="8B9A36903F62FFDCFF6460DC4A54647C" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF6460DC490E6453" box="[197,292,749,773]" italics="true" pageId="14" pageNumber="50">Females</emphasis>
(
<figureCitation id="131E2A153F62FFDCFE9260DC495E6453" box="[307,372,749,773]" captionStart="FIGURE 8" captionStartId="19.[151,250,1883,1906]" captionTargetBox="[179,1414,193,1852]" captionTargetId="figure@19.[179,1415,193,1853]" captionTargetPageId="19" captionText="FIGURE 8. (A) Enneapterygius obscurus, Marsa Alam; (B) Enneapterygius pallidus, Farasan Is.; (C) Enneapterygius pusillus, M, Sharm el Sheik; (D) Enneapterygius pusillus, M, Oman; (E) Enneapterygius ventermaculus, Farasan Is.; (E) Helcogramma obtusirostris, M, Sharm el Sheil; (F) Helcogramma obtusirostris, F, Sharm el Sheik; (G) Helcogramma steinitzi, F, Marsa Alam; (H) Norfolkia brachylepis, Bali. Photos by R. Field (B, E), J. E. Randall (D); G. R. Allen (H); S. V. Bogorodsky (the others)." httpUri="https://zenodo.org/record/279581/files/figure.png" pageId="14" pageNumber="50">Fig. 8</figureCitation>
C) are almost invisible, with only white and green spots on the body, the banded first dorsal-fin spine and white on the pectoral fins visible.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F62FFDCFF64610949D364E9" pageId="14" pageNumber="50" type="key">
<paragraph id="8B9A36903F62FFDCFF64610949D364E9" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF646109494F6406" bold="true" box="[197,357,823,848]" pageId="14" pageNumber="50">Key features.</emphasis>
Very tall first dorsal fin, in males to twice height of second dorsal fin; body of both males and females translucent, males with black chest and pelvic fins, yellow first dorsal fin, red at corner of mouth and deep blue spots below eye. Females with brown bar below eye and on pectoral-fin base, first dorsal fin equal in height to second and yellowish.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F62FFDCFF6461FC4BFD635C" pageId="14" pageNumber="50" type="distribution">
<paragraph id="8B9A36903F62FFDCFF6461FC4BFD635C" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF6461FC497464B0" bold="true" box="[197,350,973,998]" pageId="14" pageNumber="50">Distribution.</emphasis>
Restricted to the Western Indian Ocean, east to Persian Gulf and southern
<collectingCountry id="F33276003F62FFDCFB1161FC4CC664B3" box="[1200,1260,973,997]" name="India" pageId="14" pageNumber="50">India</collectingCountry>
, south to
<collectingCountry id="F33276003F62FFDCFAFA61FC48C8635C" name="South Africa" pageId="14" pageNumber="50">South Africa</collectingCountry>
, described from the Red Sea (
<typeStatus id="549E88323F62FFDCFD9161C34A48635C" box="[560,610,1010,1034]" pageId="14" pageNumber="50">type</typeStatus>
locality, Massawa,
<collectingCountry id="F33276003F62FFDCFC9E61C34BBA635C" box="[831,912,1010,1034]" name="Eritrea" pageId="14" pageNumber="50">Eritrea</collectingCountry>
); rare.
</paragraph>
</subSubSection>
<subSubSection id="C33F651B3F62FFDCFF6466294A99632C" pageId="14" pageNumber="50" type="discussion">
<paragraph id="8B9A36903F62FFDCFF6466294A99632C" blockId="14.[151,1437,151,1146]" pageId="14" pageNumber="50">
<emphasis id="B951EA823F62FFDCFF64662949136366" bold="true" box="[197,313,1048,1072]" pageId="14" pageNumber="50">Remarks.</emphasis>
Like
<taxonomicName id="4C254D133F62FFDCFEDA66284A296366" box="[379,515,1048,1072]" class="Actinopterygii" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="species" species="altipinnis">
<emphasis id="B951EA823F62FFDCFEDA66284A296366" box="[379,515,1048,1072]" italics="true" pageId="14" pageNumber="50">E. altipinnis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D133F62FFDCFDB366284AA36366" box="[530,649,1048,1072]" class="Actinopterygii" family="Tripterygiidae" genus="Helcogramma" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="50" phylum="Chordata" rank="species" species="pusillus">
<emphasis id="B951EA823F62FFDCFDB366284AA36366" box="[530,649,1048,1072]" italics="true" pageId="14" pageNumber="50">E. pusillus</emphasis>
</taxonomicName>
can be distinguished by the very tall first dorsal fin. Both species live on a silty sand or mud substrate, close to the base of reefs, often on stones and dead corals of the same color. Found in bays or sheltered lagoons. Attains
<quantity id="4CDD9B753F62FFDCFD9466534AAE632C" box="[565,644,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="50" unit="mm" value="30.0">30 mm</quantity>
SL.
</paragraph>
</subSubSection>
</treatment>
</document>