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<document id="CD16A545BDF7F822BD3ECFB0E97D2E32" ID-DOI="10.5281/zenodo.173409" ID-GBIF-Dataset="35be9144-74a5-4a2d-be11-42c5f112283e" ID-ISSN="1175­5326" ID-Zenodo-Dep="173409" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459774222433" checkinUser="plazi" docAuthor="Framenau, Volker W." docDate="2006" docId="038C8784FFAFC04BFECFFA6FE88DE369" docLanguage="en" docName="zt01281p067.pdf" docOrigin="Zootaxa 1281" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Knoelle Framenau, 2006, gen. nov." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFB5FFFCFFADC048FFC7FFEEEB6DE537" masterDocTitle="Knoelle, a new monotypic wolf spider genus from Australia (Araneae: Lycosidae)" masterLastPageNumber="67" masterPageNumber="55" pageNumber="57" updateTime="1698217622538" updateUser="plazi">
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<mods:title id="F043179739B6354B96BD9D056824062A">Knoelle, a new monotypic wolf spider genus from Australia (Araneae: Lycosidae)</mods:title>
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<mods:namePart id="FBB0984F276A2C35BA1B3D4A3D920176">Framenau, Volker W.</mods:namePart>
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<mods:date id="68A99E27B5615D2A41985CA41EBF8582">2006</mods:date>
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<treatment id="038C8784FFAFC04BFECFFA6FE88DE369" ID-DOI="http://doi.org/10.5281/zenodo.6262156" ID-GBIF-Taxon="119347674" ID-Zenodo-Dep="6262156" LSID="urn:lsid:plazi:treatment:038C8784FFAFC04BFECFFA6FE88DE369" httpUri="http://treatment.plazi.org/id/038C8784FFAFC04BFECFFA6FE88DE369" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
<subSubSection id="C33F6519FFAFC04AFECFFA6FEAB7E0AC" box="[264,474,1409,1435]" pageId="2" pageNumber="57" type="nomenclature">
<paragraph id="8B9A3692FFAFC04AFECFFA6FEAB7E0AC" blockId="2.[264,474,1409,1435]" box="[264,474,1409,1435]" pageId="2" pageNumber="57">
<heading id="D0D281FEFFAFC04AFECFFA6FEAB7E0AC" bold="true" box="[264,474,1409,1435]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
<emphasis id="B951EA80FFAFC04AFECFFA6FEAB7E0AC" bold="true" box="[264,474,1409,1435]" pageId="2" pageNumber="57">
<taxonomicName id="4C254D11FFAFC04AFECFFA6FEA08E0AC" ID-CoL="58L7" box="[264,357,1409,1435]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EA80FFAFC04AFECFFA6FEA08E0AC" bold="true" box="[264,357,1409,1435]" italics="true" pageId="2" pageNumber="57">Knoelle</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257FBFFAFC04AFEABFA6FEAB7E0AC" box="[364,474,1409,1435]" pageId="2" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33F6519FFAFC04AFECFFA2AE8EAE0EC" box="[264,903,1476,1499]" pageId="2" pageNumber="57" type="materials_examined">
<paragraph id="8B9A3692FFAFC04AFECFFA2AE8EAE0EC" blockId="2.[264,903,1476,1499]" box="[264,903,1476,1499]" pageId="2" pageNumber="57">
<emphasis id="B951EA80FFAFC04AFECFFA2AEAE0E0EC" box="[264,397,1476,1499]" italics="true" pageId="2" pageNumber="57">
<typeStatus id="549E8830FFAFC04AFECFFA2AEA57E0EC" box="[264,314,1476,1499]" pageId="2" pageNumber="57">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C254D11FFAFC04AFE5BFA2AE9A7E0EC" authority="L. Koch, 1877" authorityName="L. Koch" authorityYear="1877" box="[412,714,1476,1499]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="B951EA80FFAFC04AFE5BFA2AE948E0EC" box="[412,549,1476,1499]" italics="true" pageId="2" pageNumber="57">Lycosa clara</emphasis>
<bibRefCitation id="EFB44B63FFAFC04AFDEBFA2AE9A7E0EC" author="Koch" box="[556,714,1476,1499]" pageId="2" pageNumber="66" refString="Koch, L. (1877) Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Bauer and Raspe, Nurnberg, pp. 889 - 968." type="book chapter" year="1877">L. Koch, 1877</bibRefCitation>
</taxonomicName>
. Here designated.
</paragraph>
</subSubSection>
<subSubSection id="C33F6519FFAFC04AFECFF9E2E972E341" pageId="2" pageNumber="57" type="etymology">
<paragraph id="8B9A3692FFAFC04AFECFF9E2E972E341" blockId="2.[264,1323,1548,1894]" pageId="2" pageNumber="57">
<emphasis id="B951EA80FFAFC04AFECFF9E2EAF8E311" bold="true" box="[264,405,1548,1574]" pageId="2" pageNumber="57">Etymology.</emphasis>
The generic name honours my mother, the late Hildegard Framenau. Knölle was a nickname she used frequently, and may refer to the
<collectingCountry id="F3327602FFAFC04AFC71F9DAEF79E379" box="[950,1044,1588,1614]" name="Germany" pageId="2" pageNumber="57">German</collectingCountry>
word Knolle (= spud). The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C33F6519FFAFC04BFEFFF96AEAF0E499" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="diagnosis">
<paragraph id="8B9A3692FFAFC04BFEFFF96AEAF0E499" blockId="2.[264,1323,1548,1894]" lastBlockId="3.[264,1324,284,1630]" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
<emphasis id="B951EA80FFAFC04AFEFFF96AEADDE3A9" bold="true" box="[312,432,1668,1694]" pageId="2" pageNumber="57">Diagnosis</emphasis>
.
<taxonomicName id="4C254D11FFAFC04AFE07F96AE976E3AA" box="[448,539,1668,1693]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EA80FFAFC04AFE07F96AE976E3AA" box="[448,539,1668,1693]" italics="true" pageId="2" pageNumber="57">Knoelle</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAFC04AFDE4F96AE9FCE3A9" bold="true" box="[547,657,1668,1694]" pageId="2" pageNumber="57">
<taxonomicNameLabel id="A26257FBFFAFC04AFDE4F96AE9FCE3A9" box="[547,657,1668,1694]" pageId="2" pageNumber="57" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is similar to representatives of the genus
<emphasis id="B951EA80FFAFC04AFBB8F96AEFBAE3AA" box="[1151,1239,1668,1693]" italics="true" pageId="2" pageNumber="57">Ve na t or</emphasis>
Hogg,
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respect to somatic characters, in particular its body colouration with a ventral black patch on the opisthosoma. However, it differs considerably by the presence of a large patch of macrosetae at the tip of the cymbium in males which is absent in
<taxonomicName id="4C254D11FFAFC04AFBBDF912EFB9E222" box="[1146,1236,1788,1813]" class="Arachnida" family="Lycosidae" genus="Venator" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA80FFAFC04AFBBDF912EFB9E222" box="[1146,1236,1788,1813]" italics="true" pageId="2" pageNumber="57">Venator</emphasis>
</taxonomicName>
. Males in the genus
<taxonomicName id="4C254D11FFAFC04AFE77F8CAE942E20A" box="[432,559,1828,1853]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA80FFAFC04AFE77F8CAE942E20A" box="[432,559,1828,1853]" italics="true" pageId="2" pageNumber="57">Hoggicosa</emphasis>
</taxonomicName>
also have a patch of macrosetae at the tip of the cymbium, however it is much smaller. In addition, the tegular apophysis is very different in
<taxonomicName id="4C254D11FFAEC04BFECFFEF2EAEAE402" box="[264,391,284,309]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA80FFAEC04BFECFFEF2EAEAE402" box="[264,391,284,309]" italics="true" pageId="3" pageNumber="58">Hoggicosa</emphasis>
</taxonomicName>
as it is of a simple, triangular
<typeStatus id="549E8830FFAEC04BFCCBFEF2E853E401" box="[780,830,284,310]" pageId="3" pageNumber="58">type</typeStatus>
lacking the two hooks and the lamellar structure characteristic for
<taxonomicName id="4C254D11FFAEC04BFD8CFEAAE9CBE46A" box="[587,678,324,349]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EA80FFAEC04BFD8CFEAAE9CBE46A" box="[587,678,324,349]" italics="true" pageId="3" pageNumber="58">Knoelle</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAEC04BFD77FEADE84DE46A" bold="true" box="[688,800,323,349]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26257FBFFAEC04BFD77FEADE84DE46A" box="[688,800,323,349]" pageId="3" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C254D11FFAEC04BFCEDFEAAE8C4E46A" box="[810,937,324,349]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA80FFAEC04BFCEDFEAAE8C4E46A" box="[810,937,324,349]" italics="true" pageId="3" pageNumber="58">Hoggicosa</emphasis>
</taxonomicName>
also have a very different body colouration with a distinct sexual dimorphism that is absent in
<taxonomicName id="4C254D11FFAEC04BFC28FE82EF27E4B2" box="[1007,1098,364,389]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EA80FFAEC04BFC28FE82EF27E4B2" box="[1007,1098,364,389]" italics="true" pageId="3" pageNumber="58">Knoelle</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAEC04BFB95FE85EFACE4B2" bold="true" box="[1106,1217,363,389]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26257FBFFAEC04BFB95FE85EFACE4B2" box="[1106,1217,363,389]" pageId="3" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EFB44B63FFAEC04BFB17FE82EA28E499" author="McKay" pageId="3" pageNumber="66" refString="McKay, R. J. (1973) The wolf spiders of Australia (Araneae: Lycosidae): 1. The bicolor group. Memoirs of the Queensland Museum, 16, 375 - 398." type="journal article" year="1973">McKay 1973</bibRefCitation>
,
<bibRefCitation id="EFB44B63FFAEC04BFE96FE7AEAFDE499" author="McKay" box="[337,400,404,430]" lastPageId="3" lastPageNumber="67" pageId="3" pageNumber="66" refString="McKay, R. J. (1975) The wolf spiders of Australia (Araneae: Lycosidae): 5. Two new species of the bicolor group. Memoirs of the Queensland Museum, 17, 313 - 318." type="journal article" year="1975">1975</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6519FFAEC04BFEFFFE55EF4AE1F9" pageId="3" pageNumber="58" type="description">
<paragraph id="8B9A3692FFAEC04BFEFFFE55E879E131" blockId="3.[264,1324,284,1630]" pageId="3" pageNumber="58">
<emphasis id="B951EA80FFAEC04BFEFFFE55EAA4E4E2" bold="true" box="[312,457,443,469]" pageId="3" pageNumber="58">Description</emphasis>
. Medium sized
<taxonomicName id="4C254D11FFAEC04BFD4BFE52E86EE4E1" box="[652,771,444,470]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
(TL ca. 10.0 25.0 mm). Males smaller than females. Prosoma longer than wide (
<figureCitation id="131E2A17FFAEC04BFD70FE0AE996E4C9" box="[695,763,484,510]" captionStart="FIGURE 1 3" captionStartId="7.[264,367,1710,1734]" captionTargetBox="[290,1296,729,1684]" captionTargetId="figure@7.[290,1296,729,1684]" captionTargetPageId="7" captionText="FIGURE 1 3. Knoelle clara (L. Koch) comb. nov. 1 2 male from Drysdale River Station, Western Australia (WAM T 56241). 1 Dorsal view (body length 12.23 mm); 2 right male pedipalp, prolateral view (cymbium length 2.22 mm). 3 Female from Kapalga, Northern Territory (WAM T 58372), epigyne, ventral view (posterior width of median septum 0.69 mm)." httpUri="https://zenodo.org/record/173410/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
), dorsal profile straight in lateral view (
<figureCitation id="131E2A17FFAEC04BFB14FE0AEE70E4C9" box="[1235,1309,484,510]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">Fig. 4</figureCitation>
). Head flanks comparatively steep in males (
<figureCitation id="131E2A17FFAEC04BFD39FDE2E828E711" box="[766,837,524,550]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
). Dorsal shield of prosoma brown to dark brown, with distinct light brown median and submarginal bands; submarginal and brown marginal bands irregular; white setae in median and submarginal bands. Anterior median eyes ca. twice as large as anterior lateral eyes, row of anterior eyes narrower than row of posterior median eyes; row of anterior eyes slightly procurved (
<figureCitation id="131E2A17FFAEC04BFBCCFD42EF3BE7F1" box="[1035,1110,684,710]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
). Chelicerae with three promarginal and three retromarginal teeth. Labium shorter than or as long as wide. Opisthosoma mottled olive­grey, centrally lighter and with dark, irregular heart mark (
<figureCitation id="131E2A17FFAEC04BFB3DFD12EA7BE609" captionStart="FIGURE 1 3" captionStartId="7.[264,367,1710,1734]" captionTargetBox="[290,1296,729,1684]" captionTargetId="figure@7.[290,1296,729,1684]" captionTargetPageId="7" captionText="FIGURE 1 3. Knoelle clara (L. Koch) comb. nov. 1 2 male from Drysdale River Station, Western Australia (WAM T 56241). 1 Dorsal view (body length 12.23 mm); 2 right male pedipalp, prolateral view (cymbium length 2.22 mm). 3 Female from Kapalga, Northern Territory (WAM T 58372), epigyne, ventral view (posterior width of median septum 0.69 mm)." httpUri="https://zenodo.org/record/173410/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
). Venter with a large black patch covering about three quarters of venter from epigastric furrow. Leg formula VI&gt;I&gt;II&gt;III. Spination of legs: Femur: 3 dorsal, 2 apicoprolateral, 2 retrolateral; patella: 1 prolateral (male only), 1 retrolateral; tibia: 2 dorsal (male only), 3 ventral pairs, 2 prolateral (male only), 2 retrolateral; metatarsus: 3 ventral pairs, 2 prolateral (male only), 2 retrolateral (male only), 1 apicoprolateral (male only), 1 apicoretrolateral (male only), 1 apicoventral.
</paragraph>
<paragraph id="8B9A3692FFAEC04BFEFFFBFAEF4AE1F9" blockId="3.[264,1324,284,1630]" pageId="3" pageNumber="58">
Cymbium tip with large patch of macrosetae (
<figureCitation id="131E2A17FFAEC04BFC9DFBFAE8C5E119" box="[858,936,1044,1070]" captionStart="FIGURE 1 3" captionStartId="7.[264,367,1710,1734]" captionTargetBox="[290,1296,729,1684]" captionTargetId="figure@7.[290,1296,729,1684]" captionTargetPageId="7" captionText="FIGURE 1 3. Knoelle clara (L. Koch) comb. nov. 1 2 male from Drysdale River Station, Western Australia (WAM T 56241). 1 Dorsal view (body length 12.23 mm); 2 right male pedipalp, prolateral view (cymbium length 2.22 mm). 3 Female from Kapalga, Northern Territory (WAM T 58372), epigyne, ventral view (posterior width of median septum 0.69 mm)." httpUri="https://zenodo.org/record/173410/files/figure.png" pageId="3" pageNumber="58">Figs 2</figureCitation>
,
<figureCitation id="131E2A17FFAEC04BFC72FBFAE888E119" box="[949,997,1044,1070]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">67</figureCitation>
). Tegulum undivided, large retrolateral tegular lobe; tegular apophysis with two distinct, basally directed hooks. Embolus originating prolaterally on palea and curving ventrally around it, long and slender. Terminal apophysis and pars pendula sickle­shaped (
<figureCitation id="131E2A17FFAEC04BFBD5FB62EF0FE191" box="[1042,1122,1164,1190]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">Fig. 8</figureCitation>
). Epigyne with inverted T­shaped median septum, that widens anteriorly (
<figureCitation id="131E2A17FFAEC04BFC68FB5AE897E1F9" box="[943,1018,1204,1230]" captionStart="FIGURE 1 3" captionStartId="7.[264,367,1710,1734]" captionTargetBox="[290,1296,729,1684]" captionTargetId="figure@7.[290,1296,729,1684]" captionTargetPageId="7" captionText="FIGURE 1 3. Knoelle clara (L. Koch) comb. nov. 1 2 male from Drysdale River Station, Western Australia (WAM T 56241). 1 Dorsal view (body length 12.23 mm); 2 right male pedipalp, prolateral view (cymbium length 2.22 mm). 3 Female from Kapalga, Northern Territory (WAM T 58372), epigyne, ventral view (posterior width of median septum 0.69 mm)." httpUri="https://zenodo.org/record/173410/files/figure.png" pageId="3" pageNumber="58">Figs 3</figureCitation>
,
<figureCitation id="131E2A17FFAEC04BFBC0FB5AEF75E1F9" box="[1031,1048,1204,1230]" captionStart="FIGURES 4 10" captionStartId="8.[264,382,1741,1765]" captionTargetBox="[281,1307,325,1701]" captionTargetId="figure@8.[274,1315,301,1705]" captionTargetPageId="8" captionText="FIGURES 4 10. Knoelle clara (L. Koch) comb. nov. 4 8 Male from Drysdale River Station, Western Australia (WAM T 56241). 4 prosoma profile in lateral view; 5 eye pattern in frontal view; 6 7 left male pedipalp (6 ventral, 7 lateral view); 8 left male pedipalp, apical part of bulbus, ventral view. 9 10 Female from Kapalga, Northern Territory (WAM T 58372). epigyne (9 ventral, 10 dorsal view). Scale bar: 4 5.70 mm; 5 3.80 mm; 6 7 1.05 mm; 8 0.50 mm; 9 10 0.95 mm." httpUri="https://zenodo.org/record/173411/files/figure.png" pageId="3" pageNumber="58">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6519FFAEC04BFEFFFB32E994E1C1" box="[312,761,1244,1270]" pageId="3" pageNumber="58" type="distribution">
<paragraph id="8B9A3692FFAEC04BFEFFFB32E994E1C1" blockId="3.[264,1324,284,1630]" box="[312,761,1244,1270]" pageId="3" pageNumber="58">
<emphasis id="B951EA80FFAEC04BFEFFFB32EAB5E1C1" bold="true" box="[312,472,1244,1270]" pageId="3" pageNumber="58">Distribution.</emphasis>
As for species (
<figureCitation id="131E2A17FFAEC04BFD52FB32E986E1C1" box="[661,747,1244,1270]" captionStart="FIGURE 11" captionStartId="10.[264,367,1853,1877]" captionTargetBox="[268,1316,985,1825]" captionTargetId="figure@10.[264,1323,966,1849]" captionTargetPageId="10" captionText="FIGURE 11. Records (n = 131) of Knoelle clara (L. Koch) comb. nov. in Australia." httpUri="https://zenodo.org/record/173412/files/figure.png" pageId="3" pageNumber="58">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6519FFAEC04BFEFFFAEAE88DE369" pageId="3" pageNumber="58" type="discussion">
<paragraph id="8B9A3692FFAEC04BFEFFFAEAE88DE369" blockId="3.[264,1324,284,1630]" pageId="3" pageNumber="58">
<emphasis id="B951EA80FFAEC04BFEFFFAEAEADCE029" bold="true" box="[312,433,1284,1310]" pageId="3" pageNumber="58">Remarks.</emphasis>
<taxonomicName id="4C254D11FFAEC04BFE78FAEAE90BE02A" box="[447,614,1284,1309]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="clara" status="comb. nov.">
<emphasis id="B951EA80FFAEC04BFE78FAEAE90BE02A" box="[447,614,1284,1309]" italics="true" pageId="3" pageNumber="58">Knoelle clara</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAEC04BFDB3FAEAE86CE029" bold="true" box="[628,769,1284,1310]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26257FBFFAEC04BFDB3FAEAE86CE029" box="[628,769,1284,1310]" pageId="3" pageNumber="58">comb. nov.</taxonomicNameLabel>
</emphasis>
is a typical representative of the subfamily Lycosinae as the tegular apophysis is transverse, with a ventrally directed spur and it has a sinuous channel on the dorsal surface (
<bibRefCitation id="EFB44B63FFAEC04BFCC7FABAE8C0E059" author="Dondale" box="[768,941,1364,1390]" pageId="3" pageNumber="66" refString="Dondale, C. D. (1986) The subfamilies of wolf spiders (Araneae: Lycosidae). Actas X Congreso Internacional de Aracnologia, Jaca, Espana, 1, 327 - 332." type="journal article" year="1986">Dondale 1986</bibRefCitation>
). A recent molecular phylogeny inferred from 12S rRNA and NADH1 that included seventy world­wide
<taxonomicName id="4C254D11FFAEC04BFB9FFA92EFA2E0A1" box="[1112,1231,1404,1430]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
species placed
<taxonomicName id="4C254D11FFAEC04BFE9BFA4AEAD0E08A" box="[348,445,1444,1469]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="clara" status="comb. nov.">
<emphasis id="B951EA80FFAEC04BFE9BFA4AEAD0E08A" box="[348,445,1444,1469]" italics="true" pageId="3" pageNumber="58">K. clara</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAEC04BFE03FA4AE926E089" bold="true" box="[452,587,1444,1470]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26257FBFFAEC04BFE03FA4AE926E089" box="[452,587,1444,1470]" pageId="3" pageNumber="58">comb. nov.</taxonomicNameLabel>
</emphasis>
as sister taxon to
<taxonomicName id="4C254D11FFAEC04BFCD8FA4AE882E08A" box="[799,1007,1444,1469]" class="Arachnida" family="Lycosidae" genus="Allocosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="hasseltii">
<emphasis id="B951EA80FFAEC04BFCD8FA4AE882E08A" box="[799,1007,1444,1469]" italics="true" pageId="3" pageNumber="58">Allocosa hasseltii</emphasis>
</taxonomicName>
(parsimony analysis) or in a polytomy with most other Australian Lycosinae (Bayesian analysis) (
<bibRefCitation id="EFB44B63FFAEC04BFBBAFA22EA25E339" author="Murphy" pageId="3" pageNumber="67" refString="Murphy, N. P., Framenau, V. W., Donellan, S. C., Harvey, M. S., Park, Y. - C. &amp; Austin, A. D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 rRNA and NADH 1 genes: implications for classification, biogeography and the evolution of web building behavior. Molecular Phylogenetics and Evolution, 38, 583 - 602." type="journal article" year="2006">
Murphy
<emphasis id="B951EA80FFAEC04BFB2DFA22EE46E0D2" box="[1258,1323,1484,1509]" italics="true" pageId="3" pageNumber="58">et al.</emphasis>
2006
</bibRefCitation>
). Morphological evidence places
<taxonomicName id="4C254D11FFAEC04BFD1BFA1AE85AE33A" box="[732,823,1524,1549]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EA80FFAEC04BFD1BFA1AE85AE33A" box="[732,823,1524,1549]" italics="true" pageId="3" pageNumber="58">Knoelle</emphasis>
</taxonomicName>
<emphasis id="B951EA80FFAEC04BFC86FA1AE8DFE339" bold="true" box="[833,946,1524,1550]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26257FBFFAEC04BFC86FA1AE8DFE339" box="[833,946,1524,1550]" pageId="3" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
close to
<emphasis id="B951EA80FFAEC04BFBE3FA1AEF11E33A" box="[1060,1148,1524,1549]" italics="true" pageId="3" pageNumber="58">Ve n at o r</emphasis>
or
<taxonomicName id="4C254D11FFAEC04BFB6BFA1AEE46E33A" box="[1196,1323,1524,1549]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA80FFAEC04BFB6BFA1AEE46E33A" box="[1196,1323,1524,1549]" italics="true" pageId="3" pageNumber="58">Hoggicosa</emphasis>
</taxonomicName>
(see generic diagnosis), but a detailed phylogenetic analysis of the Australian Lycosinae is required to ascertain the systematic position of the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>