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<document id="E217C5C2FF4ECA628B8E92DEE3B2D626" ID-CLB-Dataset="301817" ID-DOI="10.5281/zenodo.13407183" ID-GBIF-Dataset="b8ad0eea-c84c-41f2-8a61-3f0ef6d45687" ID-ISSN="1732-2421" ID-Zenodo-Dep="13407183" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1723062243441" checkinUser="felipe" docAuthor="Salesa, Manuel J., Sánchez, Israel M. &amp; Morales, Jorge" docDate="2004" docId="038C1A2E72084256B7263D9BFAF1FDD0" docLanguage="en" docName="ActaPalaeontolPol.49.2.189-196.pdf" docOrigin="Acta Palaeontologica Polonica 49 (2)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Sinohippus sampelayoi" docType="treatment" docVersion="3" lastPageNumber="192" masterDocId="FFB5625672094255B40A3846FFE1FF9C" masterDocTitle="Presence of the Asian horse Sinohippus in the Miocene of Europe" masterLastPageNumber="196" masterPageNumber="189" pageNumber="190" updateTime="1724889626526" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FEE3ADD7B3F6E5255437EA2325D2F12F">Presence of the Asian horse Sinohippus in the Miocene of Europe</mods:title>
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<treatment id="038C1A2E72084256B7263D9BFAF1FDD0" ID-DOI="http://doi.org/10.5281/zenodo.13407193" ID-Zenodo-Dep="13407193" LSID="urn:lsid:plazi:treatment:038C1A2E72084256B7263D9BFAF1FDD0" httpUri="http://treatment.plazi.org/id/038C1A2E72084256B7263D9BFAF1FDD0" lastPageId="3" lastPageNumber="192" pageId="1" pageNumber="190">
<subSubSection id="C33FF8B372084254B7263D9BFC9EF980" pageId="1" pageNumber="190" type="nomenclature">
<paragraph id="8B9AAB3872084254B7263D9BFC9EF980" blockId="1.[812,1419,1319,1597]" pageId="1" pageNumber="190">
<heading id="D0D21C5472084254B7263D9BFC9EF980" fontSize="12" level="2" pageId="1" pageNumber="190" reason="9">
<taxonomicName id="4C25D0BB72084254B7263D9BFC9AF980" authority="(Villalta and Crusafont, 1945)" baseAuthorityName="Villalta and Crusafont" baseAuthorityYear="1945" class="Mammalia" family="Equidae" genus="Sinohippus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A72084254B7263D9BFBB5FA65" box="[812,1108,1501,1529]" italics="true" pageId="1" pageNumber="190">Sinohippus sampelayoi</emphasis>
(
<bibRefCitation id="EFB4D6C972084254B06F3D9AFC90F980" author="Villalta, J. F. &amp; Crusafont, M." pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont, 1945</bibRefCitation>
)
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33FF8B372084256B7263E6EFAF1FDD0" lastPageId="3" lastPageNumber="192" pageId="1" pageNumber="190" type="description">
<paragraph id="8B9AAB3872084254B7263E6EFC6DF9A1" blockId="1.[812,1419,1319,1597]" box="[812,908,1576,1597]" pageId="1" pageNumber="190">
<figureCitation id="131EB7BD72084254B7263E6EFC69F9A1" box="[812,904,1576,1597]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[102,135,1363,1383]" captionStartId-1="2.[102,135,1977,1997]" captionStartId-2="3.[102,135,1975,1995]" captionStartId-3="4.[102,135,758,778]" captionTargetBox-0="[128,747,237,1321]" captionTargetBox-1="[123,1464,225,1950]" captionTargetBox-2="[102,774,223,1948]" captionTargetBox-3="[102,775,227,720]" captionTargetId-0="figure-526@1.[123,753,233,1325]" captionTargetId-1="figure-10@2.[123,1464,225,1950]" captionTargetId-2="figure-471@3.[102,775,223,1948]" captionTargetId-3="figure-640@4.[102,775,227,720]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="Fig. 1. Cladogram proposed by Sánchez et al. (1998) for the Iberian Anchitheriinae, showing the relationships between the two lineages of Anchitherium. Abbreviations: A., Anchitherium; S., Sinohippus." captionText-1="Fig. 2. Sinohippus sampelayoi, IGME 1175M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (A), buccal (B), and lingual (C) views." captionText-2="Fig. 3. Sinohippus sampelayoi, IPS 2001. Fragment of left hemimandible with p4, m1 and m2. In occlusal (A), lingual (B), and buccal (C) views." captionText-3="Fig. 4. Sinohippus sampelayoi, MNCN 50110. Occlusal view of fragment of right maxilla with P3P4." figureDoi-0="http://doi.org/10.5281/zenodo.13407189" figureDoi-1="http://doi.org/10.5281/zenodo.13407199" figureDoi-2="http://doi.org/10.5281/zenodo.13407203" figureDoi-3="http://doi.org/10.5281/zenodo.13407212" httpUri-0="https://zenodo.org/record/13407189/files/figure.png" httpUri-1="https://zenodo.org/record/13407199/files/figure.png" httpUri-2="https://zenodo.org/record/13407203/files/figure.png" httpUri-3="https://zenodo.org/record/13407212/files/figure.png" pageId="1" pageNumber="190">Figs. 14</figureCitation>
.
</paragraph>
<paragraph id="8B9AAB3872084254B7263E09FA2FF9F8" blockId="1.[812,1486,1615,2025]" box="[812,1486,1615,1636]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263E09FC6DF9F8" box="[812,908,1615,1636]" italics="true" pageId="1" pageNumber="190">Synonyms</emphasis>
:
<taxonomicName id="4C25D0BB72084254B7943E09FA2BF9F8" authority="Villalta and Crusafont, 1945" authorityName="Villalta and Crusafont" authorityYear="1945" box="[926,1482,1615,1636]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A72084254B7943E09FB78F9F8" box="[926,1177,1615,1636]" italics="true" pageId="1" pageNumber="190">Anchitherium sampelayoi</emphasis>
<bibRefCitation id="EFB4D6C972084254B0AF3E09FA2BF9F8" author="Villalta, J. F. &amp; Crusafont, M." box="[1189,1482,1615,1636]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont, 1945</bibRefCitation>
</taxonomicName>
:
</paragraph>
<paragraph id="8B9AAB3872084254B7583E2DFA2FF91C" blockId="1.[812,1486,1615,2025]" box="[850,1486,1643,1664]" pageId="1" pageNumber="190">
figs. 14;
<taxonomicName id="4C25D0BB72084254B7B73E2DFA2FF91C" authority="Abusch-Siewert, 1983" authorityName="Abusch-Siewert" authorityYear="1983" box="[957,1486,1643,1664]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="subSpecies" species="ezquerrae" subSpecies="sampelayoi">
<emphasis id="B951772A72084254B7B73E2DFAC3F91C" box="[957,1314,1643,1664]" italics="true" pageId="1" pageNumber="190">Anchitherium ezquerrae sampelayoi</emphasis>
<bibRefCitation id="EFB4D6C972084254B1273E2DFA2FF91C" author="Abusch-Siewert, S." box="[1325,1486,1643,1664]" pageId="1" pageNumber="190" pagination="1 - 361" refId="ref4018" refString="Abusch-Siewert, S. 1983. Gebissmorphologische Untersuchungen an eurasiatischen Anchitherien (Equidae, Mammalia) unter besonderer Berucksichtigung der Fundstelle Sandelzhausen. Courier Forschungsinstitut Senckenberg 62: 1 - 361." type="journal article" year="1983">AbuschSiewert,</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B9AAB3872084254B7583EC1FC39F900" blockId="1.[812,1486,1615,2025]" box="[850,984,1671,1692]" pageId="1" pageNumber="190">1983: pl. 3: 2.</paragraph>
<paragraph id="8B9AAB3872084254B7263EEDFA20F940" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<materialsCitation id="3B4DA16572084254B7263EEDFA20F940" collectionCode="IGME" pageId="1" pageNumber="190" specimenCode="IGME 1175" specimenCount="1" typeStatus="holotype">
<emphasis id="B951772A72084254B7263EEDFC65F95C" box="[812,900,1707,1728]" italics="true" pageId="1" pageNumber="190">
<typeStatus id="549E159A72084254B7263EEDFC65F95C" box="[812,900,1707,1728]" pageId="1" pageNumber="190" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DB83034372084254B79A3EEDFBE3F95C" box="[912,1026,1707,1728]" collectionCode="IGME" pageId="1" pageNumber="190">IGME 1175</specimenCode>
M (
<figureCitation id="131EB7BD72084254B02E3EEDFBBFF95C" box="[1060,1118,1707,1728]" captionStart="Fig" captionStartId="2.[102,135,1977,1997]" captionTargetBox="[123,1464,225,1950]" captionTargetId="figure-10@2.[123,1464,225,1950]" captionTargetPageId="2" captionText="Fig. 2. Sinohippus sampelayoi, IGME 1175M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (A), buccal (B), and lingual (C) views." figureDoi="http://doi.org/10.5281/zenodo.13407199" httpUri="https://zenodo.org/record/13407199/files/figure.png" pageId="1" pageNumber="190">Fig. 2</figureCitation>
), Right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3 (see
<bibRefCitation id="EFB4D6C972084254B0593E81FABAF940" author="Villalta, J. F. &amp; Crusafont, M." box="[1107,1371,1735,1756]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont 1945</bibRefCitation>
:figs. 13).
</materialsCitation>
</paragraph>
<paragraph id="8B9AAB3872084254B7263EAFFB64F886" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<materialsCitation id="3B4DA16572084254B7263EAFFB60F886" collectionCode="IPS-" pageId="1" pageNumber="190" specimenCode="IPS-70" specimenCount="1" typeStatus="paratype">
<emphasis id="B951772A72084254B7263EAFFC65F962" box="[812,900,1769,1790]" italics="true" pageId="1" pageNumber="190">
<typeStatus id="549E159A72084254B7263EAFFC65F962" box="[812,900,1769,1790]" pageId="1" pageNumber="190" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCode id="DB83034372084254B79B3EAFFC36F962" box="[913,983,1769,1790]" collectionCode="IPS-" pageId="1" pageNumber="190">IPS70</specimenCode>
, Fragment of left hemimandible with d2 and d3 (see
<bibRefCitation id="EFB4D6C972084254B7263F43FBDAF886" author="Villalta, J. F. &amp; Crusafont, M." box="[812,1083,1797,1818]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont 1945</bibRefCitation>
: fig. 4)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9AAB3872084254B7263F61FBCFF837" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263F61FBC0F8A0" box="[812,1057,1831,1852]" italics="true" pageId="1" pageNumber="190">Other material examined</emphasis>
: IPS2001, fragment of hemimandible with p4, m1 and m2 (
<figureCitation id="131EB7BD72084254B7C13F04FBE4F8CB" box="[971,1029,1858,1879]" captionStart="Fig" captionStartId="3.[102,135,1975,1995]" captionTargetBox="[102,774,223,1948]" captionTargetId="figure-471@3.[102,775,223,1948]" captionTargetPageId="3" captionText="Fig. 3. Sinohippus sampelayoi, IPS 2001. Fragment of left hemimandible with p4, m1 and m2. In occlusal (A), lingual (B), and buccal (C) views." figureDoi="http://doi.org/10.5281/zenodo.13407203" httpUri="https://zenodo.org/record/13407203/files/figure.png" pageId="1" pageNumber="190">Fig. 3</figureCitation>
). There is a cast of an unpublished fragment of maxilla with P3 and P4 (
<figureCitation id="131EB7BD72084254B01A3F18FBA8F8EF" box="[1040,1097,1886,1907]" captionStart="Fig" captionStartId="4.[102,135,758,778]" captionTargetBox="[102,775,227,720]" captionTargetId="figure-640@4.[102,775,227,720]" captionTargetPageId="4" captionText="Fig. 4. Sinohippus sampelayoi, MNCN 50110. Occlusal view of fragment of right maxilla with P3P4." figureDoi="http://doi.org/10.5281/zenodo.13407212" httpUri="https://zenodo.org/record/13407212/files/figure.png" pageId="1" pageNumber="190">Fig. 4</figureCitation>
), deposited in the MNCN, which give the only information about the upper dentition of this species; the original specimen belongs to MLS.
</paragraph>
<paragraph id="8B9AAB3872084254B7263FFEFC36F875" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263FFEFC4CF851" box="[812,941,1976,1997]" italics="true" pageId="1" pageNumber="190">Type locality</emphasis>
: Nombrevilla1 (CalatayudDaroca Basin, province of Zaragoza,
<collectingCountry id="F332EBA872084254B79B3F92FC2DF875" box="[913,972,2004,2025]" name="Spain" pageId="1" pageNumber="190">Spain</collectingCountry>
).
</paragraph>
<caption id="DF5AFBB0720B4257B46C3FFFFE7DF875" ID-DOI="http://doi.org/10.5281/zenodo.13407199" ID-Zenodo-Dep="13407199" httpUri="https://zenodo.org/record/13407199/files/figure.png" pageId="2" pageNumber="191" startId="2.[102,135,1977,1997]" targetBox="[123,1464,225,1950]" targetPageId="2" targetType="figure">
<paragraph id="8B9AAB38720B4257B46C3FFFFE7DF875" blockId="2.[102,1485,1977,2025]" pageId="2" pageNumber="191">
Fig. 2.
<taxonomicName id="4C25D0BB720B4257B4AD3FFFFE15F851" authority=", IGME 1175" authorityName=", IGME" authorityYear="1175" box="[167,500,1977,1997]" class="Mammalia" family="Equidae" genus="Sinohippus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="2" pageNumber="191" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A720B4257B4AD3FFFFE98F851" box="[167,377,1977,1997]" italics="true" pageId="2" pageNumber="191">Sinohippus sampelayoi</emphasis>
, IGME 1175
</taxonomicName>
M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (
<emphasis id="B951772A720B4257B1A53FFFFA5EF851" bold="true" box="[1455,1471,1977,1997]" pageId="2" pageNumber="191">A</emphasis>
), buccal (
<emphasis id="B951772A720B4257B4A43F93FF5CF874" bold="true" box="[174,189,2005,2024]" pageId="2" pageNumber="191">B</emphasis>
), and lingual (
<emphasis id="B951772A720B4257B5493F93FEB2F875" bold="true" box="[323,339,2005,2025]" pageId="2" pageNumber="191">C</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8B9AAB38720A4256B72638A1FA5BFF60" blockId="3.[812,1466,231,252]" box="[812,1466,231,252]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B72638A1FCB0FF60" box="[812,849,231,252]" italics="true" pageId="3" pageNumber="192">Age</emphasis>
: Early Vallesian (basal Late Miocene) (zone MN 9 of
<bibRefCitation id="EFB4D6C9720A4256B14F38A1FA4EFF60" author="Mein, P." box="[1349,1455,231,252]" pageId="3" pageNumber="192" pagination="77 - 81" refId="ref5509" refString="Mein, P. 1975. Biozonation du neogene Mediterranee a partir des mammiferes. Proceedings of the VI Congress of the Regional Committee on Mediterranean Neogene Stratigraphy, Bratislava, 77 - 81." type="book chapter" year="1975">
<collectingRegion id="49E165DA720A4256B14F38A1FA99FF60" box="[1349,1400,231,252]" country="United States of America" name="Maine" pageId="3" pageNumber="192">Mein</collectingRegion>
1975
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9AAB38720A4256B726394AFC01FEA1" blockId="3.[812,1485,268,317]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B726394AFC27FEBD" box="[812,966,268,289]" italics="true" pageId="3" pageNumber="192">Other localities</emphasis>
: Soblay and DouélaFontaine (
<collectingCountry id="F332EBA8720A4256B102394AFAAFFEBD" box="[1288,1358,268,289]" name="France" pageId="3" pageNumber="192">France</collectingCountry>
), and EçmeAkçaköi (
<collectingCountry id="F332EBA8720A4256B780396EFC34FEA1" box="[906,981,296,317]" name="Turkey" pageId="3" pageNumber="192">Turkey</collectingCountry>
).
</paragraph>
<paragraph id="8B9AAB38720A4256B7263912FAF1FDD0" blockId="3.[812,1486,339,588]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B7263912FC7AFEF7" box="[812,923,340,363]" italics="true" pageId="3" pageNumber="192">Diagnosis</emphasis>
.—In
<bibRefCitation id="EFB4D6C9720A4256B7D73915FAB2FEF0" author="Villalta, J. F. &amp; Crusafont, M." box="[989,1363,339,364]" pageId="3" pageNumber="192" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont (1945: 78)</bibRefCitation>
: “terminal, derived
<taxonomicName id="4C25D0BB720A4256B78C3932FBFDFE10" authorityName="von Meyer" authorityYear="1844" box="[902,1052,372,396]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="192" phylum="Chordata" rank="genus">
<emphasis id="B951772A720A4256B78C3932FBFDFE10" box="[902,1052,372,396]" italics="true" pageId="3" pageNumber="192">Anchitherium</emphasis>
</taxonomicName>
of large size, with marked tendency to hypsodonty, basal cingulum almost obsolete, mainly in the juvenile teeth; more evident homeodonty than in the primitive forms, milk teeth highly reduced, special shape of d2”; additionally: upper premolars with marked buccal cingulum and absence of lingual cingulum, hypostyle without connection with the metalophe, and absence of crochet.
</paragraph>
</subSubSection>
</treatment>
</document>