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<document id="1E3E678C28D8D7F5AAE24885B94960BF" ID-CLB-Dataset="7899" ID-DOI="10.11646/zootaxa.4938.2.1" ID-GBIF-Dataset="e1daacda-87e9-479f-abeb-2431532b98d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4563736" ID-ZooBank="DFCB5C2C-1E70-4628-B65A-4E5322C57F3B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614324477351" checkinUser="plazi" docAuthor="León-Espinosa, Angel De, León-Gonzalez, Jesus Angel De &amp; Gómez-Gutiérrez, Jaime" docDate="2021" docId="038087FFFFD2C64CB6A084915632F816" docLanguage="en" docName="zootaxa.4938.2.1.pdf" docOrigin="Zootaxa 4938 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nymphon apheles Child 1979" docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="FFB9FF87FFC9C657B63784065265FFCE" masterDocTitle="Pycnogonids from marine docks located along the west coast of the Gulf of California, Mexico" masterLastPageNumber="195" masterPageNumber="151" pageNumber="178" updateTime="1698931148356" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EED4D0CD16CD574B554E03FD061781AB">Pycnogonids from marine docks located along the west coast of the Gulf of California, Mexico</mods:title>
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<mods:namePart id="41B1DE2EFC57784AB43CD461F290F627">León-Espinosa, Angel De</mods:namePart>
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<mods:namePart id="033D8EA6BF38FC5E227F5D80ECE045D5">León-Gonzalez, Jesus Angel De</mods:namePart>
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<mods:namePart id="6E225BE5BE7848D928A5BE8A16E0FF9E">Gómez-Gutiérrez, Jaime</mods:namePart>
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<mods:date id="6FA71F7316CB3C5ABC1A8B2D75EAC198">2021</mods:date>
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<treatment id="038087FFFFD2C64CB6A084915632F816" ID-DOI="http://doi.org/10.5281/zenodo.4579381" ID-GBIF-Taxon="179987448" ID-Zenodo-Dep="4579381" LSID="urn:lsid:plazi:treatment:038087FFFFD2C64CB6A084915632F816" httpUri="http://treatment.plazi.org/id/038087FFFFD2C64CB6A084915632F816" lastPageNumber="178" pageId="27" pageNumber="178">
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<paragraph id="8B9636E9FFD2C64CB6A08491506DFF7F" blockId="27.[151,520,151,214]" box="[151,520,151,178]" pageId="27" pageNumber="178">
<heading id="D0DE8185FFD2C64CB6A08491506DFF7F" bold="true" box="[151,520,151,178]" fontSize="11" level="1" pageId="27" pageNumber="178" reason="1">
<taxonomicName id="4C294D6AFFD2C64CB6A08491506DFF7F" ID-CoL="486SQ" authority="Child, 1979" authorityName="Child" authorityYear="1979" box="[151,520,151,178]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="B95DEAFBFFD2C64CB6A08491506DFF7F" bold="true" box="[151,520,151,178]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6A084915315FF7C" bold="true" box="[151,368,151,178]" italics="true" pageId="27" pageNumber="178">Nymphon apheles</emphasis>
<bibRefCitation id="EFB84B18FFD2C64CB74F8491506DFF7F" author="Child, C. A." box="[376,520,151,177]" pageId="27" pageNumber="178" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child, 1979</bibRefCitation>
</emphasis>
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<subSubSection id="C3336562FFD2C64CB6A084BD53C4FF1B" box="[151,417,187,214]" pageId="27" pageNumber="178" type="description">
<paragraph id="8B9636E9FFD2C64CB6A084BD53C4FF1B" blockId="27.[151,520,151,214]" box="[151,417,187,214]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6A084BD52B6FF1B" bold="true" box="[151,211,187,213]" pageId="27" pageNumber="178">
(
<figureCitation id="13122A6CFFD2C64CB6A884BD52B6FF1B" box="[159,211,187,213]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Figs</figureCitation>
</emphasis>
.
<emphasis id="B95DEAFBFFD2C64CB6D784BD535FFF1B" bold="true" box="[224,314,187,214]" pageId="27" pageNumber="178">14 A–G</emphasis>
;
<emphasis id="B95DEAFBFFD2C64CB77E84BD53C4FF1B" bold="true" box="[329,417,187,214]" pageId="27" pageNumber="178">
<figureCitation id="13122A6CFFD2C64CB77E84BD53FDFF1B" box="[329,408,187,214]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">15 A–I</figureCitation>
)
</emphasis>
</paragraph>
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<subSubSection id="C3336562FFD2C64CB6A085065009FED6" box="[151,620,255,280]" pageId="27" pageNumber="178" type="reference_group">
<paragraph id="8B9636E9FFD2C64CB6A085065009FED6" blockId="27.[151,620,255,280]" box="[151,620,255,280]" pageId="27" pageNumber="178">
<treatmentCitationGroup id="AB3911C7FFD2C64CB6A085065009FED6" box="[151,620,255,280]" pageId="27" pageNumber="178">
<treatmentCitation id="0A8810F8FFD2C64CB6A08506507CFED9" author="Child, C. A." box="[151,537,255,280]" page="37" pageId="27" pageNumber="178" year="1979">
<taxonomicName id="4C294D6AFFD2C64CB6A08506507CFED9" ID-CoL="486SQ" authority="Child, 1979: 35 - 37" authorityName="Child" authorityPageNumber="35 - 37" authorityYear="1979" box="[151,537,255,280]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="B95DEAFBFFD2C64CB6A085065329FED6" box="[151,332,256,280]" italics="true" pageId="27" pageNumber="178">Nymphon apheles</emphasis>
<bibRefCitation id="EFB84B18FFD2C64CB7658506507CFED9" author="Child, C. A." box="[338,537,255,280]" pageId="27" pageNumber="178" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child, 1979: 35–37</bibRefCitation>
</taxonomicName>
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, fig. 12.
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</paragraph>
</subSubSection>
<subSubSection id="C3336562FFD2C64CB6A085435042FE4A" pageId="27" pageNumber="178" type="materials_examined">
<paragraph id="8B9636E9FFD2C64CB6A085435042FE4A" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6A0854353EFFE91" bold="true" box="[151,394,325,351]" pageId="27" pageNumber="178">Material examined.</emphasis>
Four specimens: 4³, (UANL-FCB-PYCNO-0068), Cantamar Marine,
<geoCoordinate id="EE1D502EFFD2C64CB2DB8540571BFEAE" box="[1260,1406,326,352]" degrees="24" direction="north" minutes="16" orientation="latitude" pageId="27" pageNumber="178" precision="15" seconds="44" value="24.278889">24°16´44´´N</geoCoordinate>
, -
<geoCoordinate id="EE1D502EFFD2C64CB6A0856C535CFE4B" box="[151,313,362,389]" degrees="110" direction="west" minutes="19" orientation="longitude" pageId="27" pageNumber="178" precision="15" seconds="51" value="-110.33083">110°19´51´´W</geoCoordinate>
,
<date id="FF971029FFD2C64CB772856C53A0FE4A" box="[325,453,362,388]" pageId="27" pageNumber="178" value="2017-06-26">26/06/2017</date>
(
<figureCitation id="13122A6CFFD2C64CB7E3856C5070FE4A" box="[468,533,362,388]" captionStart="FIGURE 1" captionStartId="3.[151,250,1426,1450]" captionTargetBox="[473,1106,721,1395]" captionTargetId="figure-199@3.[473,1113,718,1400]" captionTargetPageId="3" captionText="FIGURE 1. Area of study showing the nine marine docks located along the peninsular coast of the Gulf of California, Mexico. 1 = Cabo San Lucas; 2 = Puerto Los Cabos; 3 = FONATUR-Fidepaz; 4 = La Paz; 5 = Fiscal Pier, 6 = Palmira, 7 = Cantamar; 8 = Puerto Escondido (located about 50 km south of Loreto), and 9 = FONATUR-Santa Rosalía, Baja California Sur, Mexico. A magnification of the map shown marine docks 3–7 located at Bahía de La Paz." figureDoi="http://doi.org/10.5281/zenodo.4563740" httpUri="https://zenodo.org/record/4563740/files/figure.png" pageId="27" pageNumber="178">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFD2C64CB6F0858B5776F982" pageId="27" pageNumber="178" type="description">
<paragraph id="8B9636E9FFD2C64CB6F0858B51FCFD06" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6F0858B533CFE69" bold="true" box="[199,345,397,423]" pageId="27" pageNumber="178">Description.</emphasis>
Male. Trunk completely segmented. Neck short, cylindrical, widened anteriorly; cephalon Yshaped, with a pair of conical projections at the anterior end (
<figureCitation id="13122A6CFFD2C64CB57785B451F5FE02" box="[832,912,434,460]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14</figureCitation>
A–B). Lateral processes separated by less than their own diameter in all segments (
<figureCitation id="13122A6CFFD2C64CB41085D050F6FE3E" box="[551,659,470,496]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 A</figureCitation>
;
<figureCitation id="13122A6CFFD2C64CB4A885D0516EFE3E" box="[671,779,470,497]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 A</figureCitation>
). Ocular tubercle inserted in front of the first pair of lateral processes, conical, with two small papilliform lateral projections and two pairs of oval eyes (
<figureCitation id="13122A6CFFD2C64CB2AF85FC5682FDDA" box="[1176,1255,506,532]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14</figureCitation>
A–B;
<figureCitation id="13122A6CFFD2C64CB31985FC52CAFDF6" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 A, D</figureCitation>
). Abdomen extends past lateral processes of fourth pair of legs, rises at a 45° angle upward, has two pairs of lateral setae and several rows (in groups of two or three) of smaller setae (
<figureCitation id="13122A6CFFD2C64CB5A486445180FD92" box="[915,997,578,604]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14</figureCitation>
A–B,
<figureCitation id="13122A6CFFD2C64CB218864456F7FD92" box="[1071,1170,578,605]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 I</figureCitation>
). Proboscis cylindrical, 1.5 times longer than wide, with a downward inclination of 40° with respect to the horizontal axis of the body (
<figureCitation id="13122A6CFFD2C64CB358866052AAFD6A" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 B</figureCitation>
,
<figureCitation id="13122A6CFFD2C64CB6ED868C53E6FD6A" box="[218,387,650,677]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 A, C, F</figureCitation>
), covered by groups of two setae each in longitudinal rows (
<figureCitation id="13122A6CFFD2C64CB213868C56F4FD6A" box="[1060,1169,650,677]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 C</figureCitation>
). Mouth with three lips, each lip with a row of three papillae, and a band of setae (
<figureCitation id="13122A6CFFD2C64CB52886A851EEFD06" box="[799,907,686,713]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 B</figureCitation>
).
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6F086D456ABFCFA" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
Two-articled chelifore formed by two articles, scape slightly longer than chela, cylindrical, as long as the proboscis, with a mid-dorsal seta (
<figureCitation id="13122A6CFFD2C64CB7C686F05038FCDE" box="[497,605,758,784]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 B</figureCitation>
). Thin chelae, each with a row of four dorsal setae (
<figureCitation id="13122A6CFFD2C64CB29686F0574BFCDE" box="[1185,1326,758,784]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 A, C</figureCitation>
). Immovable and movable fingers each with nine teeth, distal ends slightly curved (
<figureCitation id="13122A6CFFD2C64CB5EB871C562CFCFA" box="[988,1097,794,820]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 C</figureCitation>
;
<figureCitation id="13122A6CFFD2C64CB262871C56A4FCFA" box="[1109,1217,794,821]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 C</figureCitation>
).
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6F0873851EDFC0A" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
Palp with five articles, second one is the longest. First article short, cylindrical, glabrous. Second and third articles 2.5 and 2.3 times longer than the first article, each one with a ventro- distal seta. Fourth article1.2 times longer than the first one, with two ventro-distal setae. Fifth article1.8 times longer than the first one, with a mid-ventral and a lateral seta, and seven to ten distal setae (
<figureCitation id="13122A6CFFD2C64CB44F87AC5082FC0A" box="[632,743,938,964]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 D</figureCitation>
;
<figureCitation id="13122A6CFFD2C64CB4C487AC511EFC0A" box="[755,891,938,965]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15 E, F</figureCitation>
).
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6F087C85708FB52" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
Ovigers insert below the first pair of lateral processes (
<figureCitation id="13122A6CFFD2C64CB51F87C851F6FC26" box="[808,915,974,1000]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 B</figureCitation>
). First to third articles short, glabrous, equal in size. Fourth article 2.5 times longer than first, glabrous. Fifth article longest, 2.8 times longer than the first, with a dorso-distal seta. Sixth article twice as long as first one, with a dorso-distal seta. The four terminal article that form the strigilis each have one or two dorso-distal setae, and are armed with ventral denticulate compound spines in the formula: 5: 4: 2: 3. These compound spines are composed of 3–4 pairs of lateral teeth depending on the size. The terminal claw is the same length as article ten, curved, and armed with three teeth (
<figureCitation id="13122A6CFFD2C64CB200808456EDFB52" box="[1079,1160,1154,1180]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14</figureCitation>
F–G;
<figureCitation id="13122A6CFFD2C64CB2F88084577AFB53" box="[1231,1311,1154,1181]" captionStart="FIGURE 15" captionStartId="28.[151,250,1860,1884]" captionTargetBox="[153,1436,181,1836]" captionTargetId="figure-23@28.[151,1436,181,1836]" captionTargetPageId="28" captionText="FIGURE 15. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, proboscis, showing mouth detail and papillae (pap), distal end; C, proboscis, palp, and chelae, ventral view; D, ocular tubercle, lateral view; E, palp; F, trunk, ventral view; G, oviger; H, strigilis, enlarged; I, abdomen, dorsal view. Scale bars: A, F = 0.5 mm; C, E–G = 0.2 mm; H–I = 0.1 mm; B, D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4563770" httpUri="https://zenodo.org/record/4563770/files/figure.png" pageId="27" pageNumber="178">Fig. 15</figureCitation>
G–H).
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6F080A0569AFABA" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
Legs thin. Coxa 1 short, with single latero-distal seta. Coxae 2 and 3, are 2 and 1.5 times longer than coxa 1, both with a ventro-distal seta. Femur longer than coxae 1 to 3 combined, with two dorsal setae, distal one on a low protuberance, and two mid-ventral setae. Tibia 1 with a row of five dorsal setae and a mid-ventral seta. Tibia 2 longest, with a row of six setae, the distal-most one is the longest, and two ventro-distal setae. Tarsus with a mid-ventral seta and a ventro-distal seta. Propodus slightly curved, with six dorsal spines and seven ventral sole spines. Main claw short, 1/4 the length of the propodus, auxiliary claws ¾ the length of the main claw (
<figureCitation id="13122A6CFFD2C64CB2B1815C5697FABA" box="[1158,1266,1370,1396]" captionStart="FIGURE 14" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1436,205,1886]" captionTargetId="figure-22@25.[151,1436,181,1895]" captionTargetPageId="25" captionText="FIGURE 14. Nymphon apheles, UANL-FCB-PYCNO-0068, male: A, trunk, dorsal view; B, trunk, lateral view; C, chelae, showing finger details, enlarged; D, palp; E, third leg; F, oviger; G, strigilis, enlarged. Scale bars: A–B, D–E = 0.5 mm; C, F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4563768" httpUri="https://zenodo.org/record/4563768/files/figure.png" pageId="27" pageNumber="178">Fig. 14 E</figureCitation>
).
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6F0817850E2FA2E" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6F081785169FA59" bold="true" box="[199,780,1405,1432]" pageId="27" pageNumber="178">Measurements of the illustrated specimen (mm):</emphasis>
Trunk:
<quantity id="4CD19B0CFFD2C64CB551817851D8FA56" box="[870,957,1406,1432]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="178" unit="mm" value="1.1">1.1 mm</quantity>
long from the anterior part of the cephalic segment to the insertion of the fourth pair of lateral processes, 0.49 wide between the second pair of lateral processes. Proboscis:
<quantity id="4CD19B0CFFD2C64CB75081C053AEFA2E" box="[359,459,1478,1504]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="27" pageNumber="178" unit="mm" value="0.36">0.36 mm</quantity>
long, 0.22 wide.
</paragraph>
<paragraph id="8B9636E9FFD2C64CB6A081EC53E7F9E6" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">Third leg: Coxa 1, 0.13 coxa 2, 0.27; coxa 3, 0.15; femur, 0.56; tibia 1, 0.62; tibia 2, 0.88; tarsus, 0.09; propodus, 0.4; main claw, 0.12.</paragraph>
<paragraph id="8B9636E9FFD2C64CB6A082345776F982" blockId="27.[151,1437,325,2008]" box="[151,1299,1586,1613]" pageId="27" pageNumber="178">Oviger: 1) 0.12; 2) 0.12; 3) 0.31; 4) 0.36; 5) 0.24; 6) 0.19; 7) 0.14; 8) 0.12; 9) 0.12; 10) 0.13; 11) 0.12.</paragraph>
</subSubSection>
<subSubSection id="C3336562FFD2C64CB6F082535048F976" pageId="27" pageNumber="178" type="distribution">
<paragraph id="8B9636E9FFD2C64CB6F082535048F976" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6F08253533CF9A1" bold="true" box="[199,345,1621,1647]" pageId="27" pageNumber="178">Distribution</emphasis>
.
<bibRefCitation id="EFB84B18FFD2C64CB75082505390F9BE" author="Child, C. A." box="[359,501,1622,1648]" pageId="27" pageNumber="178" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child (1979)</bibRefCitation>
described
<taxonomicName id="4C294D6AFFD2C64CB45A82505150F9BE" authorityName="Child" authorityYear="1979" box="[621,821,1622,1648]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="B95DEAFBFFD2C64CB45A82505150F9BE" box="[621,821,1622,1648]" italics="true" pageId="27" pageNumber="178">Nymphon apheles</emphasis>
</taxonomicName>
for Punta Paitilla, Pacific coast of
<collectingCountry id="F33E7679FFD2C64CB28382505775F9BE" box="[1204,1296,1622,1648]" name="Panama" pageId="27" pageNumber="178">Panama</collectingCountry>
. The present record of
<taxonomicName id="4C294D6AFFD2C64CB735827D531DF95A" authorityName="Child" authorityYear="1979" box="[258,376,1658,1684]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="B95DEAFBFFD2C64CB735827D531DF95A" box="[258,376,1658,1684]" italics="true" pageId="27" pageNumber="178">N. apheles</emphasis>
</taxonomicName>
in the Gulf of California is the first report since 1979 and extends the distribution of the species about
<quantity id="4CD19B0CFFD2C64CB6EB82985323F976" box="[220,326,1694,1720]" metricMagnitude="6" metricUnit="m" metricValue="3.9" pageId="27" pageNumber="178" unit="km" value="3900.0">3,900 km</quantity>
north from
<collectingCountry id="F33E7679FFD2C64CB7FA8298504CF976" box="[461,553,1694,1720]" name="Panama" pageId="27" pageNumber="178">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3336562FFD2C64CB6F082C45632F816" pageId="27" pageNumber="178" type="discussion">
<paragraph id="8B9636E9FFD2C64CB6F082C45632F816" blockId="27.[151,1437,325,2008]" pageId="27" pageNumber="178">
<emphasis id="B95DEAFBFFD2C64CB6F082C45351F912" bold="true" box="[199,308,1730,1756]" pageId="27" pageNumber="178">Remarks</emphasis>
. The genus
<taxonomicName id="4C294D6AFFD2C64CB78A82C4504DF912" authorityName="Fabricius" authorityYear="1794" box="[445,552,1730,1756]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAFBFFD2C64CB78A82C4504DF912" box="[445,552,1730,1756]" italics="true" pageId="27" pageNumber="178">Nymphon</emphasis>
</taxonomicName>
currently includes 255 nominal species in the world, of which only three have been reported in
<collectingCountry id="F33E7679FFD2C64CB76582E053C9F8CE" box="[338,428,1766,1792]" name="Mexico" pageId="27" pageNumber="178">Mexico</collectingCountry>
:
<taxonomicName id="4C294D6AFFD2C64CB78282E05162F8CE" authority="Scott, 1912" authorityName="Scott" authorityYear="1912" box="[437,775,1766,1792]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="pixellae">
<emphasis id="B95DEAFBFFD2C64CB78282E050E5F8CE" box="[437,640,1766,1792]" italics="true" pageId="27" pageNumber="178">Nymphon pixellae</emphasis>
<bibRefCitation id="EFB84B18FFD2C64CB4B182E05162F8CE" author="Scott, F. M." box="[646,775,1766,1792]" pageId="27" pageNumber="178" pagination="206 - 208" refId="ref23748" refString="Scott, F. M. (1912) On a species of Nymphon from the North Pacific. Annals and Magazine of Natural History, Series 8, 10 (56), 206 - 208. https: // doi. org / 10.1080 / 00222931208693219" type="journal article" year="1912">Scott, 1912</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C294D6AFFD2C64CB52682E056DDF8CE" authority="Hedgpeth, 1948" authorityName="Hedgpeth" authorityYear="1948" box="[785,1208,1766,1792]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="floridanum">
<emphasis id="B95DEAFBFFD2C64CB52682E0519AF8CE" box="[785,1023,1766,1792]" italics="true" pageId="27" pageNumber="178">Nymphon floridanum</emphasis>
<bibRefCitation id="EFB84B18FFD2C64CB23182E056DDF8CE" author="Hedgpeth, J. W." box="[1030,1208,1766,1792]" pageId="27" pageNumber="178" pagination="157 - 342" refId="ref21840" refString="Hedgpeth, J. W. (1948) The Pycnogonida of the western North Atlantic and the Caribbean. Proceedings of the United States National Museum, 97 (3216), 157 - 342. https: // doi. org / 10.5479 / si. 00963801.97 - 3216.157" type="journal article" year="1948">Hedgpeth, 1948</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C294D6AFFD2C64CB2DB82E05344F8EA" authority="Child, 1979" authorityName="Child" authorityYear="1979" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="lituus">
<emphasis id="B95DEAFBFFD2C64CB2DB82E057F9F8CE" box="[1260,1436,1766,1792]" italics="true" pageId="27" pageNumber="178">Nymphon lituus</emphasis>
<bibRefCitation id="EFB84B18FFD2C64CB6A0830C5344F8EA" author="Child, C. A." box="[151,289,1802,1828]" pageId="27" pageNumber="178" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child, 1979</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFB84B18FFD2C64CB718830C53A7F8EA" author="Child, C. A." box="[303,450,1802,1828]" pageId="27" pageNumber="178" pagination="1 - 86" refId="ref20895" refString="Child, C. A. (1979). Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of middle America. Smithsonian Contributions to Zoology, 293, 1 - 86. https: // doi. org / 10.5479 / si. 00810282.293" type="journal article" year="1979">Child (1979)</bibRefCitation>
described it on the basis of a juvenile female specimen. Of the four adult specimens collected in the present study, two were males with egg masses attached to the ovigers and two were females. The present study includes the first description of a male of
<taxonomicName id="4C294D6AFFD2C64CB52D835551F6F8A2" authorityName="Child" authorityYear="1979" box="[794,915,1874,1900]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" kingdom="Animalia" order="Pantopoda" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="B95DEAFBFFD2C64CB52D835551F6F8A2" box="[794,915,1874,1900]" italics="true" pageId="27" pageNumber="178">N. apheles</emphasis>
</taxonomicName>
. Specimens from the Gulf of California have only three morphological differences compared to the
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from
<collectingCountry id="F33E7679FFD2C64CB59A8370566FF85E" box="[941,1034,1910,1936]" name="Panama" pageId="27" pageNumber="178">Panama</collectingCountry>
: presence of two proximal and two distal setae on the abdomen, as well as, the strigilis formula; which is 5: 5: 2:
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the
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and 5: 4: 2:
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the specimens of the present work. The rest of the morphological characters were similar.
</paragraph>
</subSubSection>
</treatment>
</document>
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