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<document id="C28CD79F7C295A0DD4F2B609BCC4995D" ID-DOI="10.5281/zenodo.183222" ID-GBIF-Dataset="5f96ed66-37a9-429d-bd7d-0cb8486df61e" ID-ISSN="1175-5326" ID-Zenodo-Dep="183222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459898289836" checkinUser="plazi" docAuthor="Ellenrieder, Natalia Von &amp; Garrison, Rosser W." docDate="2008" docId="038087FCFFC0491CFF66D716704EF93F" docLanguage="en" docName="zt01842p034.pdf" docOrigin="Zootaxa 1842" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Epipleoneura demarmelsi Ellenrieder &amp; Garrison, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FFB9FF84FFDB4900FFF1D13C7357FFC4" masterDocTitle="Drepanoneura gen. nov. for Epipleoneura letitia and Protoneura peruviensis, with descriptions of eight new Protoneuridae from South America (Odonata: Protoneuridae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="28" updateTime="1698228552848" updateUser="plazi">
<mods:mods id="9A88897B26F6F6FB6DD48A2CA6F48191" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0A9CE7D2F2D0DBA67B9B3163477F3EE6">
<mods:title id="07728E083B787E4E4387A925B389A60C">Drepanoneura gen. nov. for Epipleoneura letitia and Protoneura peruviensis, with descriptions of eight new Protoneuridae from South America (Odonata: Protoneuridae)</mods:title>
</mods:titleInfo>
<mods:name id="FB8DA113E1A404C93194DAD1D0125346" type="personal">
<mods:role id="1E645C8858B43752A18389827F5E4BD2">
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</mods:role>
<mods:namePart id="5E826D802041CF0ECE41881115D5E4D2">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:name id="49154573D3625142BF3FAE7A9A8E0AB3" type="personal">
<mods:role id="8B51FDBD9B4172321BD4D7853E691C60">
<mods:roleTerm id="8979C6BDF56B91ADA72612E5D84CC824">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D8B522008236FFB02F52E361F79AB35">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="93372B3812AD89D1E01F1F8A4E5F257B">text</mods:typeOfResource>
<mods:relatedItem id="09C5BEA9E4619BF85D27A7676845749B" type="host">
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<mods:title id="19E3C9C71726EFB9AAECC10F37247DB8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9C6A2903523C042AAB5FE926DE158AFE">
<mods:date id="566904EF61F62D5C93119F2830E58FFB">2008</mods:date>
<mods:detail id="B6FB068C23E16AB265DFCFE1D0EC686D" type="volume">
<mods:number id="FC9C73717C4083FF0808FD807E7097B4">1842</mods:number>
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<mods:classification id="C961860884C689B2EF3319C971ED0B9A">journal article</mods:classification>
<mods:identifier id="81639D5BCB105F2CCC0481CB883EB582" type="DOI">10.5281/zenodo.183222</mods:identifier>
<mods:identifier id="50F88365A5136E8869F3460D635AAFF0" type="GBIF-Dataset">5f96ed66-37a9-429d-bd7d-0cb8486df61e</mods:identifier>
<mods:identifier id="E3079345E889B97C361D327E995F5EBA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3A522838ACD3BE0AA7E3887556272C32" type="Zenodo-Dep">183222</mods:identifier>
</mods:mods>
<treatment id="038087FCFFC0491CFF66D716704EF93F" ID-DOI="http://doi.org/10.5281/zenodo.6231958" ID-GBIF-Taxon="119372614" ID-Zenodo-Dep="6231958" LSID="urn:lsid:plazi:treatment:038087FCFFC0491CFF66D716704EF93F" httpUri="http://treatment.plazi.org/id/038087FCFFC0491CFF66D716704EF93F" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C3336561FFC0491BFF66D71672F4F9A3" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B9636EAFFC0491BFF66D7167164F980" blockId="27.[151,563,1578,1639]" box="[151,563,1578,1604]" pageId="27" pageNumber="28">
<heading id="D0DE8186FFC0491BFF66D7167164F980" bold="true" box="[151,563,1578,1604]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B95DEAF8FFC0491BFF66D7167164F980" bold="true" box="[151,563,1578,1604]" pageId="27" pageNumber="28">
<taxonomicName id="4C294D69FFC0491BFF66D7167298F980" ID-CoL="6FV7C" box="[151,463,1578,1604]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="B95DEAF8FFC0491BFF66D7167298F980" bold="true" box="[151,463,1578,1604]" italics="true" pageId="27" pageNumber="28">Epipleoneura demarmelsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E5783FFC0491BFE27D7167164F980" box="[470,563,1578,1604]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9636EAFFC0491BFF66D77172F4F9A3" blockId="27.[151,563,1578,1639]" box="[151,419,1613,1639]" pageId="27" pageNumber="28">
<figureCitation id="13122A6FFFC0491BFF66D7717244F9A3" box="[151,275,1613,1639]" captionStart="FIGURE 11" captionStartId="9.[151,254,1816,1840]" captionTargetBox="[165,1341,228,1755]" captionTargetId="figure@9.[151,1436,194,1792]" captionTargetPageId="9" captionText="FIGURE 11. Wings. (a) Drepanoneura donnellyi sp. nov., ♂ paratype Colombia, Cristalina; (b) D. loutoni sp. nov., ♂ paratype Peru, Pakitza; (c) Eplipleoneura demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel. An: Antenodal space." httpUri="https://zenodo.org/record/183226/files/figure.png" pageId="27" pageNumber="28">Figures 11</figureCitation>
c, 22, 32, 38
</paragraph>
</subSubSection>
<subSubSection id="C3336561FFC0491CFF66D7A17754FF1E" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8B9636EAFFC0491BFF66D7A171ECF8C3" blockId="27.[151,1437,1693,1999]" pageId="27" pageNumber="28">
<emphasis id="B95DEAF8FFC0491BFF66D7A17273F973" bold="true" box="[151,292,1693,1719]" pageId="27" pageNumber="28">Etymology.</emphasis>
We name this species
<taxonomicName id="4C294D69FFC0491BFDDAD7A171E4F972" box="[555,691,1693,1718]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="B95DEAF8FFC0491BFDDAD7A171E4F972" box="[555,691,1693,1718]" italics="true" pageId="27" pageNumber="28">demarmelsi</emphasis>
</taxonomicName>
(noun in the genitive case) in honor of our good friend and colleague Jürg De Marmels, in recognition of his invaluable contributions to the knowledge of New World
<taxonomicName id="4C294D69FFC0491BFA91D7F9739FF8C3" class="Insecta" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
and his continuing kind aid in our studies.
</paragraph>
<paragraph id="8B9636EAFFC0491CFF37D6297754FF1E" blockId="27.[151,1437,1693,1999]" lastBlockId="28.[151,1437,151,1787]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B95DEAF8FFC0491BFF37D6297284F8EB" bold="true" box="[198,467,1813,1839]" pageId="27" pageNumber="28">Specimens examined.</emphasis>
Total 25 ɗ.—
<typeStatus id="54928848FFC0491BFD7DD62971A9F8EB" box="[652,766,1813,1839]" pageId="27" pageNumber="28" type="holotype">
<emphasis id="B95DEAF8FFC0491BFD7DD62971A9F8EB" bold="true" box="[652,766,1813,1839]" pageId="27" pageNumber="28">Holotype</emphasis>
</typeStatus>
ɗ:
<collectingCountry id="F33E767AFFC0491BFCDCD62970FFF8EB" box="[813,936,1813,1839]" name="Venezuela" pageId="27" pageNumber="28">Venezuela</collectingCountry>
, Bolívar State, Río Churún at entrance to Salto Ángel (
<geoCoordinate id="EE1D502DFFC0491BFEC4D60172C3F893" box="[309,404,1853,1879]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="-5.9666667">05°58'S</geoCoordinate>
,
<geoCoordinate id="EE1D502DFFC0491BFE53D601715EF893" box="[418,521,1853,1879]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="-62.516666">62°31'W</geoCoordinate>
,
<quantity id="4CD19B0FFFC0491BFDE6D6017132F893" box="[535,613,1853,1879]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="28" unit="m" value="800.0">800 m</quantity>
), 10/
<date id="FF97102AFFC0491BFD52D6017064F893" box="[675,819,1853,1879]" pageId="27" pageNumber="28" value="1990-08-11">11 viii 1990</date>
, leg. T.W. Donnelly (FSCA).
<emphasis id="B95DEAF8FFC0491BFB66D6017643F893" bold="true" box="[1175,1300,1853,1879]" pageId="27" pageNumber="28">
<typeStatus id="54928848FFC0491BFB66D6017643F893" box="[1175,1300,1853,1879]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
: 5 ɗ, same data as
<typeStatus id="54928848FFC0491BFF00D659720FF8BB" box="[241,344,1893,1919]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(TWD); 9 ɗ, same data (RWG); 2 ɗ, Río Chaverú,
<quantity id="4CD19B0FFFC0491BFC34D6597755F8BB" box="[965,1026,1893,1919]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="km" value="4.0">4 km</quantity>
W of El Paují (
<geoCoordinate id="EE1D502DFFC0491BFB4ED6597674F8BB" box="[1215,1315,1893,1919]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="4.4666667">04°28'N</geoCoordinate>
,
<geoCoordinate id="EE1D502DFFC0491BFAC1D65976C0F8BB" box="[1328,1431,1893,1919]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="-61.616665">61°37'W</geoCoordinate>
,
<quantity id="4CD19B0FFFC0491BFF66D6B173B2F863" box="[151,229,1933,1959]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="m" value="900.0">900 m</quantity>
), 0
<date id="FF97102AFFC0491BFEFAD6B172D8F863" box="[267,399,1933,1959]" pageId="27" pageNumber="28" value="1990-08-06">6 viii 1990</date>
, leg. T.W. Donnelly (TWD); 3 ɗ, same data but leg. R.W. Garrison (RWG); 1 ɗ, same data but
<collectingCountry id="F33E767AFFC0491BFF0CD68972DEF80B" box="[253,393,1973,1999]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="27" pageNumber="28">Santa Elena</collectingCountry>
de Uairén, El Paují, streams in forest N of village,
<date id="FF97102AFFC0491BFC16D6897727F80B" box="[999,1136,1973,1999]" pageId="27" pageNumber="28" value="1991-07-30">30 vii 1991</date>
, leg. G.S. Vick (RWG); 1 ɗ, same data (GSV); 1 ɗ, Río Cinco Ranchos,
<quantity id="4CD19B0FFFC7491CFD33D1A4705BFF76" box="[706,780,152,178]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="km" value="15.0">15 km</quantity>
E of El Paují (
<geoCoordinate id="EE1D502DFFC7491CFC4CD1A47775FF76" box="[957,1058,152,178]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="4.5666666">04°34'N</geoCoordinate>
,
<geoCoordinate id="EE1D502DFFC7491CFBDFD1A477C2FF76" box="[1070,1173,152,178]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-61.516666">61°31'W</geoCoordinate>
,
<quantity id="4CD19B0FFFC7491CFB53D1A477A7FF76" box="[1186,1264,152,178]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="900.0">900 m</quantity>
), 0
<date id="FF97102AFFC7491CFAE4D1A476C0FF76" box="[1301,1431,152,178]" pageId="28" pageNumber="29" value="1990-08-07">7 viii 1990</date>
, leg. R.W. Garrison (RWG); 2 ɗ, same data but leg. T.W. Donnelly (TWD).
</paragraph>
</subSubSection>
<subSubSection id="C3336561FFC7491CFF37D1DB774CFA27" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9636EAFFC7491CFF37D1DB76D2FE96" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF37D1DB729EFEC5" bold="true" box="[198,457,231,257]" pageId="28" pageNumber="29">
Male
<typeStatus id="54928848FFC7491CFEFCD1DB722DFEC5" box="[269,378,231,257]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
. Head
</emphasis>
. Labium, anteclypeus, and genae pale yellow; labrum black except for narrow marginal yellow rim; base of mandibles, postclypeus, and antefrons glossy black; remainder including rear of head matte black; anterior surface of antennifer dark brown laterally, antennal pedicel and flagellum brown.
</paragraph>
<paragraph id="8B9636EAFFC7491CFF34D063707AFC43" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D0637275FEBD" bold="true" box="[197,290,351,377]" pageId="28" pageNumber="29">Thorax</emphasis>
. Matte black; hind margin of prothorax evenly rounded with lateral rim becoming dark brown, prosternum light yellow with medio-ventral black spot; mesothorax with thin pale hairline along humeral suture interrupted at mesopleural fossa, mesostigmal plates flat and simple, with transverse length shorter than width of medial disc; metathorax becoming paler latero-ventrally and smudged with black along anterior half of metinfraepisternum; venter of thorax pale. Coxae pale smudged with black externally; trochanters yellow except for narrow margin of black between sutures; femora dark brown, becoming black distally; tibiae yellow with black base and apex, inner surface brown; tarsi and pretarsi yellow with black apices; 4 spurs on metafemora, 6 on metatibiae. Wings (as in
<figureCitation id="13122A6FFFC7491CFD6FD34471A5FD56" box="[670,754,632,658]" captionStart="FIGURE 11" captionStartId="9.[151,254,1816,1840]" captionTargetBox="[165,1341,228,1755]" captionTargetId="figure@9.[151,1436,194,1792]" captionTargetPageId="9" captionText="FIGURE 11. Wings. (a) Drepanoneura donnellyi sp. nov., ♂ paratype Colombia, Cristalina; (b) D. loutoni sp. nov., ♂ paratype Peru, Pakitza; (c) Eplipleoneura demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel. An: Antenodal space." httpUri="https://zenodo.org/record/183226/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
c) 5.4 as long as wide; 12 Px in Fw,
<quantity id="4CD19B0FFFC7491CFB59D34477B2FD56" box="[1192,1253,632,658]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="28" pageNumber="29" unit="in" value="10.0">10 in</quantity>
Hw; antenodal space 1 shorter than twice the length of 2, and about as long as 3; CuA and CuP&amp;AA completely fused to wing margin; MP reaching wing margin at basal third of first cell posterior to vein descending from subnodus; IR2 arising at vein descending from subnodus; IR2 and
<collectingCountry id="F33E767AFFC7491CFCEED3CC7014FCCE" box="[799,835,752,778]" name="Poland" pageId="28" pageNumber="29">RP</collectingCountry>
3 separated by a short crossvein; divergence of RP- RA (arculus) distal to Ax 2;
<collectingCountry id="F33E767AFFC7491CFE14D226715EFCF0" box="[485,521,794,820]" name="Poland" pageId="28" pageNumber="29">RP</collectingCountry>
<quantity id="4CD19B0FFFC7491CFDF8D2177178FCF0" box="[521,559,794,825]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="28" pageNumber="29" unit="in" value="2.0">2 in</quantity>
Fw beginning at Px 5 and in Hw at Px 3; pt pale brown with marginal yellow hairline, in Fw as long as underlying cell with costal side shorter than posterior side, in Hw longer than underlying cell with costal side as long as posterior side.
</paragraph>
<paragraph id="8B9636EAFFC7491CFF34D2A97761FB0C" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D2A97268FC6B" bold="true" box="[197,319,917,943]" pageId="28" pageNumber="29">Abdomen</emphasis>
. Dorso-laterally matte black except for narrow yellow bands interrupted medio-dorsally by black on base of S37; latero-ventral portion of terga and sterna pale brown to yellow. Genital ligula distal segment markedly constricted at mid-length, with a u-shaped cleft at apex (as in
<figureCitation id="13122A6FFFC7491CFB93D2D977ECFC3B" box="[1122,1211,997,1023]" captionStart="FIGURES 21 26" captionStartId="13.[151,269,1908,1932]" captionTargetBox="[151,1407,389,1882]" captionTargetId="figure@13.[151,1436,371,1890]" captionTargetPageId="13" captionText="FIGURES 21 26. ♂ genital ligula: (a) ectal view, (b) lateral view, (c) latero-ental view. (21) Eplipleoneura capilliformis Brazil, Obidos, in RWG; (22) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (23) E. haroldoi Brazil, Manaus, in RWG; (24) Phasmoneura exigua Brazil, Reserva Ducke, in RWG; (25) Epipotoneura machadoi sp. nov., ♂ holotype Brazil, Rio Xingu; (26) E. nehalennia Brazil, Manaus. ip: inner process; lp: latero-posterior lobe." httpUri="https://zenodo.org/record/183229/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
a); no inner fold or internal processes; a pair of latero-posterior flap-like pedunculate projections externally at level of flexure (as in
<figureCitation id="13122A6FFFC7491CFF47D5097259FB8B" box="[182,270,1077,1103]" captionStart="FIGURES 21 26" captionStartId="13.[151,269,1908,1932]" captionTargetBox="[151,1407,389,1882]" captionTargetId="figure@13.[151,1436,371,1890]" captionTargetPageId="13" captionText="FIGURES 21 26. ♂ genital ligula: (a) ectal view, (b) lateral view, (c) latero-ental view. (21) Eplipleoneura capilliformis Brazil, Obidos, in RWG; (22) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (23) E. haroldoi Brazil, Manaus, in RWG; (24) Phasmoneura exigua Brazil, Reserva Ducke, in RWG; (25) Epipotoneura machadoi sp. nov., ♂ holotype Brazil, Rio Xingu; (26) E. nehalennia Brazil, Manaus. ip: inner process; lp: latero-posterior lobe." httpUri="https://zenodo.org/record/183229/files/figure.png" pageId="28" pageNumber="29">Fig. 22</figureCitation>
a). Dorso-posterior margin of S10 not projected posteriorly (
<figureCitation id="13122A6FFFC7491CFC24D5097760FB8B" box="[981,1079,1077,1103]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 32</figureCitation>
ab). Cercus shorter than S10, with a dorso-apical hook and a short ventro-basal branch directed medially and adjacent to that of opposite cercus (
<figureCitation id="13122A6FFFC7491CFF05D5BA720FFB64" box="[244,344,1158,1184]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 32</figureCitation>
bc). Paraproct as long as cercus; epiproct forming a sclerotized, decumbent plate between cerci, with a medio-dorsal blunt tooth and distal margin trilobate (
<figureCitation id="13122A6FFFC7491CFC6AD59270ABFB0C" box="[923,1020,1198,1224]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 32</figureCitation>
bc).
</paragraph>
<paragraph id="8B9636EAFFC7491CFF34D5E9771AFB34" blockId="28.[151,1437,151,1787]" box="[197,1101,1237,1264]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D5E97201FB2B" bold="true" box="[197,342,1237,1263]" pageId="28" pageNumber="29">Dimensions</emphasis>
. Total length
<quantity id="4CD19B0FFFC7491CFE08D5EA7133FB34" box="[505,612,1238,1264]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="28" pageNumber="29" unit="mm" value="36.1">36.1 mm</quantity>
; abdomen length
<quantity id="4CD19B0FFFC7491CFCC2D5EA70C8FB34" box="[819,927,1238,1264]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="28" pageNumber="29" unit="mm" value="31.1">31.1 mm</quantity>
; Hw
<quantity id="4CD19B0FFFC7491CFC2CD5EA771FFB34" box="[989,1096,1238,1264]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="28" pageNumber="29" unit="mm" value="18.5">18.5 mm</quantity>
.
</paragraph>
<paragraph id="8B9636EAFFC7491CFF34D5C176D5FA54" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D5C1721DFAD3" bold="true" box="[197,330,1277,1303]" pageId="28" pageNumber="29">
<typeStatus id="54928848FFC7491CFF34D5C17213FAD3" box="[197,324,1277,1303]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Some
<typeStatus id="54928848FFC7491CFE6CD5C2715AFADC" box="[413,525,1278,1304]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
are young specimens with slight metallic luster on black of thorax and head, and femora lighter brown. Variation is as follows: 35 spurs on metafemora and 57 on metatibiae, 1011 Px in Hw, MP reaching wing margin at half to entire length of first cell posterior to vein descending from subnodus,
<collectingCountry id="F33E767AFFC7491CFF3FD44A73A5FA54" box="[206,242,1398,1424]" name="Poland" pageId="28" pageNumber="29">RP</collectingCountry>
<quantity id="4CD19B0FFFC7491CFF03D4B9724FFA54" box="[242,280,1398,1427]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="28" pageNumber="29" unit="in" value="2.0">2 in</quantity>
Fw beginning at Px 4 or 5 (mostly at 5), and in Hw at Px 3 or 4 (mostly at 3). Female unknown.
</paragraph>
<paragraph id="8B9636EAFFC7491CFF34D49C774CFA27" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D49C7201FA7E" bold="true" box="[197,342,1440,1466]" pageId="28" pageNumber="29">Dimensions</emphasis>
. (n = 10): total length 34.0
<quantity id="4CD19B0FFFC7491CFD5FD49D704EFA7F" box="[686,793,1441,1467]" metricMagnitude="-2" metricUnit="m" metricValue="3.59" pageId="28" pageNumber="29" unit="mm" value="35.9">35.9 mm</quantity>
[mean
<quantity id="4CD19B0FFFC7491CFC84D49D70B5FA7F" box="[885,994,1441,1467]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="28" pageNumber="29" unit="mm" value="35.1">35.1 mm</quantity>
; SD 0.6]; abdomen length 29.0
<quantity id="4CD19B0FFFC7491CFA98D49D7392FA27" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="28" pageNumber="29" unit="mm" value="30.6">30.6 mm</quantity>
[mean 30.0 mm; SD 0.6]; Hw
<quantity id="4CD19B0FFFC7491CFDC1D4F5718CFA27" box="[560,731,1481,1507]" metricMagnitude="-2" metricUnit="m" metricValue="1.82" metricValueMax="1.86" metricValueMin="1.78" pageId="28" pageNumber="29" unit="mm" value="18.2" valueMax="18.6" valueMin="17.8">17.818.6 mm</quantity>
[mean
<quantity id="4CD19B0FFFC7491CFCC3D4F570FBFA27" box="[818,940,1481,1507]" metricMagnitude="-2" metricUnit="m" metricValue="1.842" pageId="28" pageNumber="29" unit="mm" value="18.42">18.42 mm</quantity>
; SD 0.4].
</paragraph>
</subSubSection>
<subSubSection id="C3336561FFC7491CFF34D4CC72CDF947" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B9636EAFFC7491CFF34D4CC72CDF947" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D4CC7212F9CE" bold="true" box="[197,325,1520,1546]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
<taxonomicName id="4C294D69FFC7491CFEBCD4CD7129F9CE" box="[333,638,1521,1546]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="B95DEAF8FFC7491CFEBCD4CD7129F9CE" box="[333,638,1521,1546]" italics="true" pageId="28" pageNumber="29">Epipleoneura demarmelsi</emphasis>
</taxonomicName>
is most similar in morphology to
<taxonomicName id="4C294D69FFC7491CFBE0D4CD779FF9CE" box="[1041,1224,1521,1546]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="capilliformis">
<emphasis id="B95DEAF8FFC7491CFBE0D4CD779FF9CE" box="[1041,1224,1521,1546]" italics="true" pageId="28" pageNumber="29">E. capilliformis</emphasis>
</taxonomicName>
. The shape of the cerci and paraprocts is essentially the same (
<figureCitation id="13122A6FFFC7491CFD49D725701DF9F7" box="[696,842,1561,1587]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 3132</figureCitation>
), but in
<taxonomicName id="4C294D69FFC7491CFC44D7257735F9F6" box="[949,1122,1561,1586]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="B95DEAF8FFC7491CFC44D7257735F9F6" box="[949,1122,1561,1586]" italics="true" pageId="28" pageNumber="29">E. demarmelsi</emphasis>
</taxonomicName>
the ventral margin of the epiproct is convexly curved (
<figureCitation id="13122A6FFFC7491CFE04D77D710EF99F" box="[501,601,1601,1627]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 32</figureCitation>
bc), while in
<taxonomicName id="4C294D69FFC7491CFCF9D77D7096F99E" box="[776,961,1601,1626]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="capilliformis">
<emphasis id="B95DEAF8FFC7491CFCF9D77D7096F99E" box="[776,961,1601,1626]" italics="true" pageId="28" pageNumber="29">E. capilliformis</emphasis>
</taxonomicName>
it forms a bluntly shaped quadrangular process (
<figureCitation id="13122A6FFFC7491CFF0ED7557209F947" box="[255,350,1641,1667]" captionStart="FIGURES 31 33" captionStartId="16.[151,269,1865,1889]" captionTargetBox="[160,1434,210,1825]" captionTargetId="figure@16.[151,1436,194,1841]" captionTargetPageId="16" captionText="FIGURES 31 33. ♂ S 10: (a) lateral view, (b) medio-dorsal view, (c) posterior view. (31) Epipleoneura capilliformis Brazil, Manaus, in RWG; (32) E. demarmelsi sp. nov., ♂ paratype Venezuela, Salto Ángel; (36) E. haroldoi Brazil, Manaus, in RWG. ah: apical hook; ce: cercus; ep: epiproct; pa: paraproct; vb: ventro-basal branch." httpUri="https://zenodo.org/record/183234/files/figure.png" pageId="28" pageNumber="29">Figs. 31</figureCitation>
bc).
</paragraph>
</subSubSection>
<subSubSection id="C3336561FFC7491CFF34D7AC727DF917" pageId="28" pageNumber="29" type="biology_ecology">
<paragraph id="8B9636EAFFC7491CFF34D7AC727DF917" blockId="28.[151,1437,151,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D7AC727EF96E" bold="true" box="[197,297,1680,1706]" pageId="28" pageNumber="29">Biology.</emphasis>
Adults were inconspicuous. They were collected as they hovered near margins of forest rivers and streams.
</paragraph>
</subSubSection>
<subSubSection id="C3336561FFC7491CFF34D7DC704EF93F" box="[197,793,1760,1787]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8B9636EAFFC7491CFF34D7DC704EF93F" blockId="28.[151,1437,151,1787]" box="[197,793,1760,1787]" pageId="28" pageNumber="29">
<emphasis id="B95DEAF8FFC7491CFF34D7DC7231F93E" bold="true" box="[197,358,1760,1786]" pageId="28" pageNumber="29">Distribution.</emphasis>
Bolívar State in
<collectingCountry id="F33E767AFFC7491CFDDDD7DD71F3F93F" box="[556,676,1761,1787]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
(
<figureCitation id="13122A6FFFC7491CFD45D7DD705BF93F" box="[692,780,1761,1787]" captionStart="FIGURE 38" captionStartId="29.[151,254,1534,1558]" captionTargetBox="[152,1360,188,1533]" captionTargetId="figure@29.[150,1435,184,1536]" captionTargetPageId="29" captionText="FIGURE 38. Distribution map of Epipotoneura and Epipleoneura demarmelsi in the Neotropical region." httpUri="https://zenodo.org/record/183238/files/figure.png" pageId="28" pageNumber="29">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>