154 lines
16 KiB
XML
154 lines
16 KiB
XML
<document id="6982E4A2491F808DC4FE99D69D800E0C" ID-DOI="10.5252/g2015n2a5" ID-ISSN="1638-9395" ID-Zenodo-Dep="4535125" ID-ZooBank="urn:lsid:zoobank.org:pub:0117CBA4-4CE0-4431-B5F6-721F998C72C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613058030568" checkinUser="felipe" docAuthor="Fourvel, Jean-Baptiste, Fosse, Philippe, Fernandez, Philippe & Antoine, Pierre-Olivier" docDate="2015" docId="038087D7FFEDFF93FEC1FC9BFE1334EB" docLanguage="en" docName="Geodiversitas.37.2.237-266.pdf" docOrigin="Geodiversitas 37 (2)" docSource="http://dx.doi.org/10.5252/g2015n2a5" docStyle="DocumentStyle:4DA100580BA89F561A6D37CADC50A8FD.3:Geodiversitas.2015-2017.journal_article.type2" docStyleId="4DA100580BA89F561A6D37CADC50A8FD" docStyleName="Geodiversitas.2015-2017.journal_article.type2" docStyleVersion="3" docTitle="Canis lupus Linnaeus 1758" docType="treatment" docVersion="3" lastPageNumber="242" masterDocId="FFB9FFAFFFE8FF96FFEEFFAEFFA53307" masterDocTitle="Large mammals of Fouvent-Saint-Andoche (Haute-Saône, France): a glimpse into a Late Pleistocene hyena den" masterLastPageNumber="266" masterPageNumber="237" pageNumber="242" updateTime="1698870744442" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="8077F30C0CFDF46DB2C1182583015C01">Large mammals of Fouvent-Saint-Andoche (Haute-Saône, France): a glimpse into a Late Pleistocene hyena den</mods:title>
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<mods:namePart id="33F1AF427B8DB58F0AA0D9F70962CD1E">Fourvel, Jean-Baptiste</mods:namePart>
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<mods:namePart id="0CA8E14A858C55DAEAB28CBF210686FE">Fosse, Philippe</mods:namePart>
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<mods:namePart id="7B8DD0B966A50CED993CFD0FB3626827">Fernandez, Philippe</mods:namePart>
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<mods:namePart id="C8404CD9BDCF56E87AABAA292F46FAC4">Antoine, Pierre-Olivier</mods:namePart>
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<mods:title id="F799D16C7FE39C0296C334E2CB12C359">Geodiversitas</mods:title>
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<mods:date id="116038599ED6A6D41CF3946838F47EF5">2015</mods:date>
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<mods:number id="BF34346D3F390917FAC3E7826BF1D074">2015-06-26</mods:number>
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<mods:number id="F49FD21C45C4D8536E3252F963219517">37</mods:number>
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<treatment id="038087D7FFEDFF93FEC1FC9BFE1334EB" LSID="urn:lsid:plazi:treatment:038087D7FFEDFF93FEC1FC9BFE1334EB" httpUri="http://treatment.plazi.org/id/038087D7FFEDFF93FEC1FC9BFE1334EB" lastPageNumber="242" pageId="5" pageNumber="242">
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<subSubSection id="C333654AFFEDFF93FEC1FC9BFDF83057" box="[303,605,821,848]" pageId="5" pageNumber="242" type="nomenclature">
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<paragraph id="8B9636C1FFEDFF93FEC1FC9BFDF83057" blockId="5.[303,605,821,848]" box="[303,605,821,848]" pageId="5" pageNumber="242">
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<heading id="D0DE81ADFFEDFF93FEC1FC9BFDF83057" box="[303,605,821,848]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="242" reason="2">
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<taxonomicName id="4C294D42FFEDFF93FEC1FC9BFDF83057" ID-CoL="QLXL" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[303,605,821,848]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="species" species="lupus">
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<emphasis id="B95DEAD3FFEDFF93FEC1FC9BFE0B3057" bold="true" box="[303,430,821,848]" italics="true" pageId="5" pageNumber="242">Canis lupus</emphasis>
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Linnaeus, 1758
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C333654AFFEDFF93FF6AFCC1FDAA370D" pageId="5" pageNumber="242" type="materials_examined">
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<paragraph id="8B9636C1FFEDFF93FF6AFCC1FDE23081" blockId="5.[131,775,878,1034]" box="[132,583,878,902]" pageId="5" pageNumber="242">
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MATERIAL EXAMINED. —
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<collectionCode id="ED38AE04FFEDFF93FE93FCC0FE133081" box="[381,438,878,902]" pageId="5" pageNumber="242">NISP</collectionCode>
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=37;
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<collectionCode id="ED38AE04FFEDFF93FE08FCC0FDB93081" box="[486,540,878,902]" pageId="5" pageNumber="242">MNI</collectionCode>
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=12.
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</paragraph>
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<paragraph id="8B9636C1FFEDFF93FF6AFC26FDAA370D" blockId="5.[131,775,878,1034]" pageId="5" pageNumber="242">
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<emphasis id="B95DEAD3FFEDFF93FF6AFC26FF5630A7" bold="true" box="[132,243,904,928]" pageId="5" pageNumber="242">1989-1992</emphasis>
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sample: 1 left maxillary; 2 left I3; 1 right C; 1 left P4; 1 left M2; 3 right i1; 6 left i1; 1 i1?; 1 right i2; 1 left i2; 1 left i2-3; 1 right c; 1 left c; 1 left p3; 1 right p4; 1 right m1; 1 right m2; 1 left m3; 1 canine?; 1 right ulna; 1 left ulna; 1 left tibia; 1 left metatarsal II; 3 first phalanges; 3 second phalanges.
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</paragraph>
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</subSubSection>
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<subSubSection id="C333654AFFEDFF93FF6AFB9BFE1334EB" pageId="5" pageNumber="242" type="description">
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<paragraph id="8B9636C1FFEDFF93FF6AFB9BFEB0374A" blockId="5.[130,777,1077,2028]" box="[132,277,1077,1103]" pageId="5" pageNumber="242">DESCRIPTION</paragraph>
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<paragraph id="8B9636C1FFEDFF93FF6AFBFBFE1334EB" blockId="5.[130,777,1077,2028]" pageId="5" pageNumber="242">
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In Europe, two medium-to-large-sized canids coexist during the Late Pleistocene: wolf
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<taxonomicName id="4C294D42FFEDFF93FE7DFBDAFDAF3789" authorityName="Linnaeus" authorityYear="1758" box="[403,522,1140,1166]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="species" species="lupus">
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<emphasis id="B95DEAD3FFEDFF93FE7DFBDAFDAF3789" box="[403,522,1140,1166]" italics="true" pageId="5" pageNumber="242">Canis lupus</emphasis>
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</taxonomicName>
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and dhole
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<taxonomicName id="4C294D42FFEDFF93FD93FBDAFEAE37A9" authority="Pallas, 1811" authorityName="Pallas" authorityYear="1811" class="Mammalia" family="Canidae" genus="Cuon" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="species" species="alpinus">
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<emphasis id="B95DEAD3FFEDFF93FD93FBDAFCA03789" box="[637,773,1140,1166]" italics="true" pageId="5" pageNumber="242">Cuon alpinus</emphasis>
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Pallas, 1811
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</taxonomicName>
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. Their association in a same geochronological context suggests their potential identification in palaeontological samples (Perez-Ripoll
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<emphasis id="B95DEAD3FFEDFF93FE7AFB7AFE6337E9" box="[404,454,1236,1262]" italics="true" pageId="5" pageNumber="242">et al.</emphasis>
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2010; Pionnier-Capitan
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<emphasis id="B95DEAD3FFEDFF93FD3AFB7AFCA237E9" box="[724,775,1236,1262]" italics="true" pageId="5" pageNumber="242">et al.</emphasis>
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2011). Thirty seven bone remains recovered from the recent excavations have been related to large canids. The right lower carnassial (m1 – F9.C.264) has a large talonid with two cusps which is typical of wolves (
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<figureCitation id="13122A44FFEDFF93FE7CFAFDFE593669" box="[402,508,1363,1390]" captionStart="FIG" captionStartId="8.[132,143,1760,1778]" captionTargetBox="[241,1335,215,1717]" captionTargetId="figure-98@8.[744,1289,692,1248]" captionTargetPageId="8" captionText="FIG. 4.— Carnivores (other than hyenas) remains from Fouvent-Saint-Andoche, exclusive of hyenas: A, B, Panthera (Leo) spelaea (Goldfüss, 1810) right upper canines E11.224 (A) and H8.C.241(B); C, Panthera (Leo) spelaea (Goldfüss, 1810) left lower carnassial F10.A.48; D, E, Canis lupus Linnaeus, 1758 right lower carnassial F9.C.264; F, Vulpes vulpesLinnaeus, 1758 left mandible E11.A.15; G, H, Mustela (Putorius) eversmannii left mandible F11.B.214; I, Gulo gulo Linnaeus, 1758 left innominate; J, K, Ursus spelaeus Rosenmüller & Heinroth, 1794 right M2 G9.B.558. Scale bar: 20 mm. Pictures by JBF." figureDoi="http://doi.org/10.5281/zenodo.4535133" httpUri="https://zenodo.org/record/4535133/files/figure.png" pageId="5" pageNumber="242">Fig.4D, E</figureCitation>
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||
). The general proportions of this carnassial (breadth=
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||
<quantity id="4CD19B24FFEDFF93FE4AFADDFDA23689" box="[420,519,1395,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="242" unit="mm" value="12.5">12.5 mm</quantity>
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, length=
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<quantity id="4CD19B24FFEDFF93FD8BFADDFD173689" box="[613,690,1395,1422]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="242" unit="mm" value="27.0">27 mm</quantity>
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) and of the left upper carnassial (P4 – H8.C.240) (breadth=
|
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<quantity id="4CD19B24FFEDFF93FD5AFA3DFCA736A9" box="[692,770,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="242" unit="mm" value="13.0">13 mm</quantity>
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, length=
|
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<quantity id="4CD19B24FFEDFF93FF36FA1DFE8236CA" box="[216,295,1459,1486]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="242" unit="mm" value="25.0">25 mm</quantity>
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||
) match the range recorded for Late Pleistocene wolves (breadth and length variations in m1 11.1-13.2 by
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<quantity id="4CD19B24FFEDFF93FF4BFA5CFEE6350A" box="[165,323,1522,1549]" metricMagnitude="-2" metricUnit="m" metricValue="2.995" metricValueMax="3.34" metricValueMin="2.65" pageId="5" pageNumber="242" unit="mm" value="29.95" valueMax="33.4" valueMin="26.5">26.5-33.4 mm</quantity>
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and in P4 12.3-15.3 by
|
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<quantity id="4CD19B24FFEDFF93FDA2FA5DFD4F350A" box="[588,746,1523,1549]" metricMagnitude="-2" metricUnit="m" metricValue="2.685" metricValueMax="2.9899999999999998" metricValueMin="2.38" pageId="5" pageNumber="242" unit="mm" value="26.85" valueMax="29.9" valueMin="23.8">23.8-29.9 mm</quantity>
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<emphasis id="B95DEAD3FFEDFF93FD1FFA5DFCA2350B" box="[753,775,1523,1548]" italics="true" pageId="5" pageNumber="242">in</emphasis>
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||
<bibRefCitation id="EFB84B30FFEDFF93FF6AF9BCFEB5352B" author="SCHUTT G." box="[132,272,1554,1580]" pageId="5" pageNumber="242" pagination="79 - 125" refId="ref27735" refString="SCHUTT G. 1969. - Die jungpleistozane Fauna der Hohlen bei Rubeland im Harz. Quartar 20: 79 - 125." type="journal article" year="1969">Schütt 1969</bibRefCitation>
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; Boudadi-Maligne 2010; Brugal & Boudadi- Maligne 2011). Yet, these carnassials are smaller than the ones referred to as
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<taxonomicName id="4C294D42FFEDFF93FEB6F9FDFF1A358B" authority="Boudadi-Maligne, 2012" authorityName="Boudadi-Maligne" authorityYear="2012" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="subSpecies" species="lupus" subSpecies="maximus">
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<emphasis id="B95DEAD3FFEDFF93FEB6F9FDFD9F356B" box="[344,570,1618,1645]" italics="true" pageId="5" pageNumber="242">Canis lupus maximus</emphasis>
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Boudadi-Maligne, 2012
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</taxonomicName>
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, described at Jaurens (OIS3; Boudadi-Maligne 2012). Postcranial remains are also referable to
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<taxonomicName id="4C294D42FFEDFF93FDDBF93DFD2C35AB" authorityName="Linnaeus" authorityYear="1758" box="[565,649,1682,1709]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="species" species="lupus">
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<emphasis id="B95DEAD3FFEDFF93FDDBF93DFD2C35AB" box="[565,649,1682,1709]" italics="true" pageId="5" pageNumber="242">C. lupus</emphasis>
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</taxonomicName>
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. In particular, the morphological features of the fragmentary left tibia G11.B.109 are diagnostic of
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<taxonomicName id="4C294D42FFEDFF93FE2FF97CFE5935EB" authorityName="Linnaeus" authorityYear="1758" box="[449,508,1746,1772]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="genus">
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<emphasis id="B95DEAD3FFEDFF93FE2FF97CFE5935EB" box="[449,508,1746,1772]" italics="true" pageId="5" pageNumber="242">Canis</emphasis>
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||
</taxonomicName>
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||
, as defined by Pionnier-
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<bibRefCitation id="EFB84B30FFEDFF93FF6AF95CFEC9340B" author="CAPITAN M. & BEMILLI C. & BODU P. & CELERIER G. & RIE J. - G. & FOSSE P. & GARCIA, M. & VIGNE J. - D." box="[132,364,1778,1804]" pageId="5" pageNumber="242" pagination="2123 - 2140" refId="ref27395" refString="PIONNIER- CAPITAN M., BEMILLI C., BODU P., CELERIER G., FER- RIE J. - G., FOSSE P., GARCIA, M. & VIGNE J. - D. 2011. - New evidence for Upper Palaeolithic small domestic dogs in South- Western Europe. Journal of Archaeological Science 38: 2123 - 2140." type="journal article" year="2011">
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Capitan
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<emphasis id="B95DEAD3FFEDFF93FF0DF95DFEB2340B" box="[227,279,1778,1804]" italics="true" pageId="5" pageNumber="242">et al.</emphasis>
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||
(2011)
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</bibRefCitation>
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: the border of the medial malleolus is salient and prominent, whereas the distal border of the cranial side is straight and regular; moreover, the proximo caudal tuberosity of the ulnar olecranon E11.B.112 is more prominent in
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<taxonomicName id="4C294D42FFEDFF93FF39F8DCFEB6348B" authorityName="Linnaeus" authorityYear="1758" box="[215,275,1906,1932]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="genus">
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||
<emphasis id="B95DEAD3FFEDFF93FF39F8DCFEB6348B" box="[215,275,1906,1932]" italics="true" pageId="5" pageNumber="242">Canis</emphasis>
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</taxonomicName>
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as observed by Pionnier-Capitan
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<emphasis id="B95DEAD3FFEDFF93FD93F8DCFD15348C" box="[637,688,1905,1931]" italics="true" pageId="5" pageNumber="242">et al.</emphasis>
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(2011). To sum up, the large canid from Fouvent is unambiguously referable to
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<taxonomicName id="4C294D42FFEDFF93FF12F81CFE9234CB" authorityName="Linnaeus" authorityYear="1758" box="[252,311,1970,1996]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="genus">
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<emphasis id="B95DEAD3FFEDFF93FF12F81CFE9234CB" box="[252,311,1970,1996]" italics="true" pageId="5" pageNumber="242">Canis</emphasis>
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</taxonomicName>
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(morphological features) and more precisely to
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<taxonomicName id="4C294D42FFEDFF93FF4FF87FFEBE34EC" authorityName="Linnaeus" authorityYear="1758" box="[161,283,2001,2027]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="242" phylum="Chordata" rank="species" species="lupus">
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<emphasis id="B95DEAD3FFEDFF93FF4FF87FFEBE34EC" box="[161,283,2001,2027]" italics="true" pageId="5" pageNumber="242">Canis lupus</emphasis>
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</taxonomicName>
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(dimensions).
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</paragraph>
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||
</subSubSection>
|
||
</treatment>
|
||
</document> |