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<document id="459399E9D48A8D05C1ED3F5C66751926" ID-CLB-Dataset="7393" ID-DOI="10.11646/zootaxa.4944.1.1" ID-GBIF-Dataset="8adfa0f6-06a7-44b5-a3be-351d0bdc7a62" ID-ISSN="1175-5326" ID-Zenodo-Dep="4681813" ID-ZooBank="380D2F75-D4F9-4974-97E2-25E0C62CB3B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615997073071" checkinUser="plazi" docAuthor="Kobayashi, Shigeki, Johns, Chris A. &amp; Kawahara, Akito Y." docDate="2021" docId="038087CBFFFC071CFF7592B2FB14A29E" docLanguage="en" docName="zootaxa.4944.1.1.pdf" docOrigin="Zootaxa 4944 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philodoria Walsingham 1907" docType="key" docVersion="7" lastPageNumber="20" masterDocId="FFB9FFB3FFEE070FFFE29409FF89A136" masterDocTitle="Revision of the Hawaiian endemic leaf-mining moth genus Philodoria Walsingham (Lepidoptera: Gracillariidae): its conservation status, host plants and descriptions of thirteen new species" masterLastPageNumber="175" masterPageNumber="1" pageNumber="19" updateTime="1698933850732" updateUser="ExternalLinkService">
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<mods:title id="2ADDC1BCA42D8B37366FD7E576F50485">Revision of the Hawaiian endemic leaf-mining moth genus Philodoria Walsingham (Lepidoptera: Gracillariidae): its conservation status, host plants and descriptions of thirteen new species</mods:title>
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<mods:namePart id="DA29BABA94A8B9911AD371AF6C155608">Kobayashi, Shigeki</mods:namePart>
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<treatment id="038087CBFFFC071CFF7592B2FB14A29E" ID-DOI="http://doi.org/10.5281/zenodo.4968558" ID-GBIF-Taxon="181002370" ID-Zenodo-Dep="4968558" LSID="urn:lsid:plazi:treatment:038087CBFFFC071CFF7592B2FB14A29E" httpUri="http://treatment.plazi.org/id/038087CBFFFC071CFF7592B2FB14A29E" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C3336556FFFC071DFF7592B2FE3EA7E2" box="[151,439,1722,1749]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B9636DDFFFC071DFF7592B2FE3EA7E2" blockId="18.[151,1436,1722,1857]" box="[151,439,1722,1749]" pageId="18" pageNumber="19">
<emphasis id="B95DEACFFFFC071DFF7592B2FE3EA7E2" bold="true" box="[151,439,1722,1749]" pageId="18" pageNumber="19">
Key to female
<taxonomicName id="4C294D5EFFFC071DFEA992B3FE3EA7E2" ID-CoL="8V6FQ" authorityName="Walsingham" authorityYear="1907" box="[331,439,1722,1748]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">genitalia</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFFC071DFF7592D7FE78A677" pageId="18" pageNumber="19" type="description">
<paragraph id="8B9636DDFFFC071DFF7592D7FE78A677" blockId="18.[151,1436,1722,1857]" pageId="18" pageNumber="19">
(Females of
<taxonomicName id="4C294D5EFFFC071DFEC392D6FE58A7CE" box="[289,465,1758,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="limahuliensis" status="sp. nov.">
<emphasis id="B95DEACFFFFC071DFEC392D6FE58A7CE" box="[289,465,1758,1784]" italics="true" pageId="18" pageNumber="19">P. limahuliensis</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFFC071DFE3592D6FD9FA7CF" bold="true" box="[471,534,1759,1785]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A26E57B4FFFC071DFE3592D6FD9FA7CF" box="[471,534,1759,1785]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C294D5EFFFC071DFDC092D6FD34A7CE" box="[546,701,1758,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="napaliensis" status="sp. nov.">
<emphasis id="B95DEACFFFFC071DFDC092D6FD34A7CE" box="[546,701,1758,1784]" italics="true" pageId="18" pageNumber="19">P. napaliensis</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFFC071DFD2092D6FC88A7CF" bold="true" box="[706,769,1759,1785]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A26E57B4FFFC071DFD2092D6FC88A7CF" box="[706,769,1759,1785]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C294D5EFFFC071DFCEF92D6FC09A7CE" authorityName="Walsingham" authorityYear="1907" box="[781,896,1758,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis id="B95DEACFFFFC071DFCEF92D6FC09A7CE" box="[781,896,1758,1784]" italics="true" pageId="18" pageNumber="19">P. nigrella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFC071DFC6F92D6FB94A7CE" authorityName="Swezey" authorityYear="1923" box="[909,1053,1759,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pipturiana">
<emphasis id="B95DEACFFFFC071DFC6F92D6FB94A7CE" box="[909,1053,1759,1784]" italics="true" pageId="18" pageNumber="19">P. pipturiana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFC071DFBC892D6FB5AA7CE" baseAuthorityName="Swezey" baseAuthorityYear="1928" box="[1066,1235,1758,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pittosporella">
<emphasis id="B95DEACFFFFC071DFBC892D6FB5AA7CE" box="[1066,1235,1758,1784]" italics="true" pageId="18" pageNumber="19">P. pittosporella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFC071DFB0292D6FA15A7CE" box="[1248,1436,1758,1784]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="platyphylliella" status="sp. nov.">
<emphasis id="B95DEACFFFFC071DFB0292D6FA15A7CE" box="[1248,1436,1758,1784]" italics="true" pageId="18" pageNumber="19">P. platyphylliella</emphasis>
</taxonomicName>
<emphasis id="B95DEACFFFFC071DFF75930AFF5FA62B" bold="true" box="[151,214,1795,1821]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A26E57B4FFFC071DFF75930AFF5FA62B" box="[151,214,1795,1821]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFC071DFF06930AFF7CA62A" box="[228,245,1795,1820]" italics="true" pageId="18" pageNumber="19">P</emphasis>
. sp. 2, 3, 7, and
<emphasis id="B95DEACFFFFC071DFE49930AFE35A62A" box="[427,444,1795,1820]" italics="true" pageId="18" pageNumber="19">P</emphasis>
. sp. B are unknown. The female of
<taxonomicName id="4C294D5EFFFC071DFCAE930AFC64A62A" authorityName="Walsingham" authorityYear="1907" box="[844,1005,1794,1820]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="micropetala">
<emphasis id="B95DEACFFFFC071DFCAE930AFC64A62A" box="[844,1005,1794,1820]" italics="true" pageId="18" pageNumber="19">P. micropetala</emphasis>
</taxonomicName>
is known except that its corpus bursae and ductus bursae are missing)
</paragraph>
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<key id="FFB861ECFFFC071CFF759365FA15A38B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<paragraph id="8B9636DDFFFC071DFF759365FA15A6B6" blockId="18.[151,1436,1899,2039]" box="[151,1436,1899,1921]" pageId="18" pageNumber="19">
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1 Signa present with spines...............................................................................
<emphasis id="B95DEACFFFFC071DFA729362FA15A6B6" bold="true" box="[1424,1436,1899,1920]" pageId="18" pageNumber="19">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFC071DFF759380FA15A6A8" blockId="18.[151,1436,1899,2039]" box="[151,1436,1929,1951]" pageId="18" pageNumber="19">
<keyLead id="30D895E8FFFC071DFF759380FA15A6A8" box="[151,1436,1929,1951]" pageId="18" pageNumber="19">
- Signa absent; corpus bursae smooth or with some lines consisting of wrinkles running longitudinally, some sclerotized.....
<emphasis id="B95DEACFFFFC071DFA729380FA15A6A8" bold="true" box="[1424,1436,1929,1950]" pageId="18" pageNumber="19">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFC071DFF7593AFFA15A6EE" pageId="18" pageNumber="19">
<paragraph id="8B9636DDFFFC071DFF7593AFFA15A68D" blockId="18.[151,1436,1899,2039]" box="[151,1436,1958,1980]" pageId="18" pageNumber="19">
<keyLead id="30D895E8FFFC071DFF7593AFFA15A68D" box="[151,1436,1958,1980]" pageId="18" pageNumber="19">
2 Lamella antevaginalis large sclerotized plate................................................................
<emphasis id="B95DEACFFFFC071DFA7293AFFA15A68D" bold="true" box="[1424,1436,1958,1979]" pageId="18" pageNumber="19">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFC071DFF7593CDFA15A6EE" blockId="18.[151,1436,1899,2039]" box="[151,1436,1987,2009]" pageId="18" pageNumber="19">
<keyLead id="30D895E8FFFC071DFF7593CDFA15A6EE" box="[151,1436,1987,2009]" pageId="18" pageNumber="19">
- Lamella antevaginalis weakly sclerotized; antrum cup-shaped with a slender pair of lateral lobes......................
<emphasis id="B95DEACFFFFC071DFA7293CAFA15A6EE" bold="true" box="[1424,1436,1987,2008]" pageId="18" pageNumber="19">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFC071CFF7593E8FA15A1D1" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<paragraph id="8B9636DDFFFC071DFF7593E8FA15A6C0" blockId="18.[151,1436,1899,2039]" box="[151,1436,2016,2039]" pageId="18" pageNumber="19">
<keyLead id="30D895E8FFFC071DFF7593E8FA15A6C0" box="[151,1436,2016,2039]" pageId="18" pageNumber="19">
<keyStep id="30DD2E78FFFC071DFF7593E8FA15A6C0" box="[151,1436,2016,2039]" pageId="18" pageNumber="19">
3 Spines on signa large, triangular; lamella antevaginalis trapezoid (
<figureCitation id="13122A58FFFC071DFCBE93E8FC36A6C0" box="[860,959,2017,2039]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="18" pageNumber="19">Fig. 56I, J</figureCitation>
)..........................
<taxonomicName id="4C294D5EFFFC071DFAE193E8FA15A6C0" ID-CoL="4G8NM" box="[1283,1436,2016,2038]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="molokaiensis">
<emphasis id="B95DEACFFFFC071DFAE193E8FA15A6C0" bold="true" box="[1283,1436,2016,2038]" italics="true" pageId="18" pageNumber="19">P. molokaiensis</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF759491FA15A19A" blockId="19.[151,1437,151,936]" box="[151,1436,151,173]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759491FA15A19A" box="[151,1436,151,173]" pageId="19" pageNumber="20">
- Spines on signa short, linear; lamella antevaginalis trapezoid, rather narrowing at posterior end (
<figureCitation id="13122A58FFFD071CFB9F949EFB5CA19B" box="[1149,1237,151,173]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56H</figureCitation>
).........
<emphasis id="B95DEACFFFFD071CFAAD9491FA15A19A" bold="true" box="[1359,1436,151,173]" pageId="19" pageNumber="20">
<emphasis id="B95DEACFFFFD071CFAAD9491FAD7A19B" bold="true" box="[1359,1374,152,173]" italics="true" pageId="19" pageNumber="20">P</emphasis>
. sp. A
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF7594BCFA14A1FC" blockId="19.[151,1437,151,936]" box="[151,1437,180,203]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7594BCFA14A1FC" box="[151,1437,180,203]" pageId="19" pageNumber="20">
- Spines on signa large claw-shaped (
<figureCitation id="13122A58FFFD071CFDDB94BCFD18A1FD" box="[569,657,181,203]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56G</figureCitation>
)...................................................
<taxonomicName id="4C294D5EFFFD071CFAE094BCFA14A1FC" ID-CoL="4G8NH" box="[1282,1437,180,202]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lysimachiella">
<emphasis id="B95DEACFFFFD071CFAE094BCFA14A1FC" bold="true" box="[1282,1437,180,202]" italics="true" pageId="19" pageNumber="20">P. lysimachiella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF7594DAFA15A1D1" blockId="19.[151,1437,151,936]" box="[151,1436,210,232]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7594DAFA15A1D1" box="[151,1436,210,232]" pageId="19" pageNumber="20">
- Spines on signa oblong and podlike (
<figureCitation id="13122A58FFFD071CFDA794DBFD33A1DE" box="[581,698,210,232]" captionStart="FIGURE 40" captionStartId="75.[151,250,1791,1815]" captionTargetBox="[201,1379,192,1757]" captionTargetId="figure-20@75.[194,1392,181,1766]" captionTargetPageId="75" captionText="FIGURE 40. Genitalia of Philodoria. AD: Male genitalia of P. marginestrigata, Kauai (CJ521/SK784), host: Sida fallax, EH: Female genitalia of P. basalis, Hawaii (Big Island [SKH09-1]), host: Metrosideros polymorpha, EG: SK617, H: SK616. A Phallus, lateral view, B Genitalia except right valva and phallus, ventral view, C Right valva, inside view, D Dorsal flap, inside view, E, G Female genitalia, lateral view, F, H Same, ventral view. aa: apophysis anterioris, an: antrum, ap: apophysis posterioris, cb: corpus bursae, co: coecum, cr: cornuti, db: ductus bursae, de: ductus ejaculatorius, ds: ductus seminalis, la: lamella antevaginalis, ob: ostium bursae, ph: phallus, phb: phallobase, sa: saccus, si: signum, ta: tuba analis, te: tegumen, va: valva, vi: vinculum." figureDoi="http://doi.org/10.5281/zenodo.4681899" httpUri="https://zenodo.org/record/4681899/files/figure.png" pageId="19" pageNumber="20">Fig. 40G, H</figureCitation>
,
<figureCitation id="13122A58FFFD071CFD2694DBFC83A1DE" box="[708,778,210,232]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">56K, L</figureCitation>
)........................
<taxonomicName id="4C294D5EFFFD071CFBCF94DBFB00A1DE" ID-CoL="4G8N7" authorityName="Walsingham" authorityYear="1907" box="[1069,1161,210,232]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B95DEACFFFFD071CFBCF94DBFB00A1DE" bold="true" box="[1069,1161,210,232]" italics="true" pageId="19" pageNumber="20">P. basalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFB7794DBFA85A1DE" ID-CoL="4G8NY" authorityName="Walsingham" authorityYear="1907" box="[1173,1292,210,232]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="B95DEACFFFFD071CFB7794DBFA85A1DE" bold="true" box="[1173,1292,210,232]" italics="true" pageId="19" pageNumber="20">P. splendida</emphasis>
</taxonomicName>
,
<emphasis id="B95DEACFFFFD071CFAF594DBFA15A1D1" bold="true" box="[1303,1436,210,232]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFAF594DBFAE9A1DE" ID-CoL="8VM44" box="[1303,1376,210,232]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFAF594DBFAE9A1DE" bold="true" box="[1303,1376,210,232]" italics="true" pageId="19" pageNumber="20">P. lama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFA8494DBFA15A1D1" box="[1382,1436,210,231]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFD071CFF7594F9FA15A014" pageId="19" pageNumber="20">
<paragraph id="8B9636DDFFFD071CFF7594F9FA15A033" blockId="19.[151,1437,151,936]" box="[151,1436,239,261]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7594F9FA15A033" box="[151,1436,239,261]" pageId="19" pageNumber="20">
4 End of ductus bursae membranous........................................................................
<emphasis id="B95DEACFFFFD071CFA7294F9FA15A033" bold="true" box="[1424,1436,240,261]" pageId="19" pageNumber="20">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF759504FA15A014" blockId="19.[151,1437,151,936]" box="[151,1436,269,291]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759504FA15A014" box="[151,1436,269,291]" pageId="19" pageNumber="20">
- End of ductus bursae sclerotized, bifurcated................................................................
<emphasis id="B95DEACFFFFD071CFA729504FA15A014" bold="true" box="[1424,1436,269,290]" pageId="19" pageNumber="20">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFD071CFF759523FA15A0A1" pageId="19" pageNumber="20">
<paragraph id="8B9636DDFFFD071CFF759523FA15A076" blockId="19.[151,1437,151,936]" box="[151,1436,298,320]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759523FA15A076" box="[151,1436,298,320]" pageId="19" pageNumber="20">
5 Signa small and narrow with minute spines; lateral lobes of antrum short (
<figureCitation id="13122A58FFFD071CFC4B9523FC75A076" box="[937,1020,298,320]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56F</figureCitation>
)............................
<taxonomicName id="4C294D5EFFFD071CFAAD9523FA15A076" ID-CoL="8VLKM" authorityName="Kobayashi, Johns &amp; Kawahara" authorityYear="2018" box="[1359,1436,298,320]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kolea">
<emphasis id="B95DEACFFFFD071CFAAD9523FA15A076" bold="true" box="[1359,1436,298,320]" italics="true" pageId="19" pageNumber="20">P. kolea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF759541FA15A06B" blockId="19.[151,1437,151,936]" box="[151,1436,327,350]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759541FA15A06B" box="[151,1436,327,350]" pageId="19" pageNumber="20">
- Signa spiny pyriform with minute spines (
<figureCitation id="13122A58FFFD071CFD94954EFD45A06B" box="[630,716,327,349]" captionStart="FIGURE 57" captionStartId="107.[151,250,1554,1578]" captionTargetBox="[181,1405,189,1520]" captionTargetId="figure-20@107.[176,1411,181,1530]" captionTargetPageId="107" captionText="FIGURE 57. Female genitalia of Pipturus-mining Philodoria. A P. costalis, paralectotype, SK653 B Same species, paratype, SK652 C P. pipturiella, SK722 D P. pipturicola, paralectotype, lateral view, SK717 E Same species, paralectotype, SK716 F P. haelaauensis, SK792 G P. floscula, SK819." figureDoi="http://doi.org/10.5281/zenodo.4681933" httpUri="https://zenodo.org/record/4681933/files/figure.png" pageId="19" pageNumber="20">Fig. 57F</figureCitation>
;
<bibRefCitation id="EFB84B2CFFFD071CFD37954EFC19A06B" author="Zimmerman, E. C." box="[725,912,327,349]" pageId="19" pageNumber="20" refId="ref103182" refString="Zimmerman, E. C. (1978 a) Gracillarioidea. In: Microlepidoptera. Part 1. Insects of Hawaii, Vol. 9. University of Hawaii Press, Honolulu, pp. 644 - 718." type="book" year="1978">Zimmerman 1978a</bibRefCitation>
: fig. 480)..........
<emphasis id="B95DEACFFFFD071CFB809541FAB1A06B" bold="true" box="[1122,1336,327,349]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFB809541FB72A06B" ID-CoL="8VM3X" box="[1122,1275,327,349]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="haelaauensis" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFB809541FB72A06B" bold="true" box="[1122,1275,327,349]" italics="true" pageId="19" pageNumber="20">P. haelaauensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFAE39541FAB1A06B" box="[1281,1336,328,349]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C294D5EFFFD071CFAA19541FA15A06B" ID-CoL="4G8NX" baseAuthorityName="Walsingham" baseAuthorityYear="1907" box="[1347,1436,327,349]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="spilota">
<emphasis id="B95DEACFFFFD071CFAA19541FA15A06B" bold="true" box="[1347,1436,327,349]" italics="true" pageId="19" pageNumber="20">P. spilota</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF75956CFA14A04C" blockId="19.[151,1437,151,936]" box="[151,1437,356,379]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF75956CFA14A04C" box="[151,1437,356,379]" pageId="19" pageNumber="20">
- Signa rather broad, sinouous, linear; lateral lobes long and slender (
<figureCitation id="13122A58FFFD071CFC88956CFC4BA04D" box="[874,962,357,379]" captionStart="FIGURE 57" captionStartId="107.[151,250,1554,1578]" captionTargetBox="[181,1405,189,1520]" captionTargetId="figure-20@107.[176,1411,181,1530]" captionTargetPageId="107" captionText="FIGURE 57. Female genitalia of Pipturus-mining Philodoria. A P. costalis, paralectotype, SK653 B Same species, paratype, SK652 C P. pipturiella, SK722 D P. pipturicola, paralectotype, lateral view, SK717 E Same species, paralectotype, SK716 F P. haelaauensis, SK792 G P. floscula, SK819." figureDoi="http://doi.org/10.5281/zenodo.4681933" httpUri="https://zenodo.org/record/4681933/files/figure.png" pageId="19" pageNumber="20">Fig. 57G</figureCitation>
)..............................
<taxonomicName id="4C294D5EFFFD071CFAD4956CFA14A04C" ID-CoL="4G8NC" box="[1334,1437,356,378]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="floscula">
<emphasis id="B95DEACFFFFD071CFAD4956CFA14A04C" bold="true" box="[1334,1437,356,378]" italics="true" pageId="19" pageNumber="20">P. floscula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF75958AFA15A0A1" blockId="19.[151,1437,151,936]" box="[151,1436,386,408]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF75958AFA15A0A1" box="[151,1436,386,408]" pageId="19" pageNumber="20">
- Signa with a pair of large spines..........................................................................
<emphasis id="B95DEACFFFFD071CFA72958BFA15A0A1" bold="true" box="[1424,1436,386,407]" pageId="19" pageNumber="20">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFD071CFF759596FA14A0C6" pageId="19" pageNumber="20">
<paragraph id="8B9636DDFFFD071CFF759596FA14A083" blockId="19.[151,1437,151,936]" box="[151,1437,415,437]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759596FA14A083" box="[151,1437,415,437]" pageId="19" pageNumber="20">
6 Spines long and slender (
<figureCitation id="13122A58FFFD071CFE369596FDCEA083" box="[468,583,415,437]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56A, B</figureCitation>
)...........................................................
<taxonomicName id="4C294D5EFFFD071CFAF095A9FA14A083" ID-CoL="4G8NZ" authorityName="Walsingham" authorityYear="1907" box="[1298,1437,415,437]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="succedanea">
<emphasis id="B95DEACFFFFD071CFAF095A9FA14A083" bold="true" box="[1298,1437,415,437]" italics="true" pageId="19" pageNumber="20">P. succedanea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF7595B4FA14A0E4" blockId="19.[151,1437,151,936]" box="[151,1437,444,467]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7595B4FA14A0E4" box="[151,1437,444,467]" pageId="19" pageNumber="20">
- Spines on signa small and rounded (
<figureCitation id="13122A58FFFD071CFDDF95B4FD26A0E4" box="[573,687,445,467]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56D, E</figureCitation>
)................................................
<taxonomicName id="4C294D5EFFFD071CFAE095B4FA14A0E4" ID-CoL="8VLYL" authorityName="Kobayashi, Johns &amp; Kawahara" authorityYear="2018" box="[1282,1437,444,466]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kauaulaensis">
<emphasis id="B95DEACFFFFD071CFAE095B4FA14A0E4" bold="true" box="[1282,1437,444,466]" italics="true" pageId="19" pageNumber="20">P. kauaulaensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF7595D2FA14A0C6" blockId="19.[151,1437,151,936]" box="[151,1437,474,496]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7595D2FA14A0C6" box="[151,1437,474,496]" pageId="19" pageNumber="20">
- Spines on signa blunt (
<figureCitation id="13122A58FFFD071CFE5F95D3FD9AA0C6" box="[445,531,474,496]" captionStart="FIGURE 56" captionStartId="106.[151,250,1881,1905]" captionTargetBox="[170,1411,192,1821]" captionTargetId="figure-23@106.[165,1422,181,1857]" captionTargetPageId="106" captionText="FIGURE 56. Female genitalia of Philodoria. A P. succedanea, paralectotype, Maui, SK714, B Same species, Hawaii (Big Island), SK624, C P. auromagnifica, Hawaii (Big Island), SK623, D P. kauaulaensis, holotype, SK690, E Same species, lateral view, F P. kolea, paratype, Hawaii (Big Island), SK634, G P. lysimachiella, SK654, H P. sp. A, SK753, I P. molokaiensis, SK668, lateral view, J Same ventral view, K P. splendida, SK621, L P. lama, paratype, SK813." figureDoi="http://doi.org/10.5281/zenodo.4681931" httpUri="https://zenodo.org/record/4681931/files/figure.png" pageId="19" pageNumber="20">Fig. 56C</figureCitation>
)............................................................
<taxonomicName id="4C294D5EFFFD071CFB1095D3FA14A0C6" ID-CoL="4G8N6" authorityName="Walsingham" authorityYear="1907" box="[1266,1437,474,496]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="auromagnifica">
<emphasis id="B95DEACFFFFD071CFB1095D3FA14A0C6" bold="true" box="[1266,1437,474,496]" italics="true" pageId="19" pageNumber="20">P. auromagnifica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFD071CFF7595F1FA15A352" pageId="19" pageNumber="20">
<paragraph id="8B9636DDFFFD071CFF7595F1FA14A33B" blockId="19.[151,1437,151,936]" box="[151,1437,503,525]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7595F1FA14A33B" box="[151,1437,503,525]" pageId="19" pageNumber="20">
7 Signa small and podlike with minute spines (
<figureCitation id="13122A58FFFD071CFD7095FEFD6FA33B" box="[658,742,503,525]" captionStart="FIGURE 59" captionStartId="110.[151,250,1714,1738]" captionTargetBox="[166,1426,193,1682]" captionTargetId="figure-23@110.[151,1436,181,1690]" captionTargetPageId="110" captionText="FIGURE 59. Female genitalia of Philodoria. A P. neraudicola, paralectotype, Oahu, SK708, host: Neraudia melastomifalia, B Same species, Maui, SK838, host: Pipturus albidus, C P. obamaorum, paratype, Hawaii (Big Island), SK710, host: Pipturus albidus, D Same species, Hawaii (Big Island), SK844 host: Pipturus sp., E P. urerana, paralectotype, SK672, F Philodoria sp. 1, SK835, G Same species, PHIL0021, H P. keahii, paratype, SK756, I P. funkae, holotype, SK761, J P. touchardiella, SK764, K P. keaensis, holotype, Hawaii (Big Island), SK777, L P. hauicola, Oahu, SK618." figureDoi="http://doi.org/10.5281/zenodo.4681937" httpUri="https://zenodo.org/record/4681937/files/figure.png" pageId="19" pageNumber="20">Fig. 59L</figureCitation>
)................................................
<taxonomicName id="4C294D5EFFFD071CFACC95F1FA14A33B" ID-CoL="4G8ND" baseAuthorityName="Swezey" baseAuthorityYear="1910" box="[1326,1437,503,525]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hauicola">
<emphasis id="B95DEACFFFFD071CFACC95F1FA14A33B" bold="true" box="[1326,1437,503,525]" italics="true" pageId="19" pageNumber="20">P. hauicola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF75961CFA15A31C" blockId="19.[151,1437,151,936]" box="[151,1436,532,555]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF75961CFA15A31C" box="[151,1436,532,555]" pageId="19" pageNumber="20">
- Signa two short lines with minute spines (
<figureCitation id="13122A58FFFD071CFD96961CFD5FA31D" box="[628,726,533,555]" captionStart="FIGURE 58" captionStartId="109.[151,250,1622,1646]" captionTargetBox="[164,1427,193,1591]" captionTargetId="figure-20@109.[152,1435,181,1598]" captionTargetPageId="109" captionText="FIGURE 58. Female genitalia of Philodoria. A P. wilkesiella, SK761, B P. epibathra, SK679, C P. dubauticola, SK645, D Same species, PHIL0008, E P. dubautiella, SK692, F P. alakaiensis, paratype, SK659, G Same species, lateral view, H P. hibiscella, SK723, I P. naenaeiella, SK703, J Same species, SK834, K P. knudseniiella, paratype, SK863, L P. hesperomanniella, paratype, SK832, M Same species, SK620." figureDoi="http://doi.org/10.5281/zenodo.4681935" httpUri="https://zenodo.org/record/4681935/files/figure.png" pageId="19" pageNumber="20">Figs. 58H</figureCitation>
,
<figureCitation id="13122A58FFFD071CFD02961CFC8AA31D" box="[736,771,533,555]" captionStart="FIGURE 59" captionStartId="110.[151,250,1714,1738]" captionTargetBox="[166,1426,193,1682]" captionTargetId="figure-23@110.[151,1436,181,1690]" captionTargetPageId="110" captionText="FIGURE 59. Female genitalia of Philodoria. A P. neraudicola, paralectotype, Oahu, SK708, host: Neraudia melastomifalia, B Same species, Maui, SK838, host: Pipturus albidus, C P. obamaorum, paratype, Hawaii (Big Island), SK710, host: Pipturus albidus, D Same species, Hawaii (Big Island), SK844 host: Pipturus sp., E P. urerana, paralectotype, SK672, F Philodoria sp. 1, SK835, G Same species, PHIL0021, H P. keahii, paratype, SK756, I P. funkae, holotype, SK761, J P. touchardiella, SK764, K P. keaensis, holotype, Hawaii (Big Island), SK777, L P. hauicola, Oahu, SK618." figureDoi="http://doi.org/10.5281/zenodo.4681937" httpUri="https://zenodo.org/record/4681937/files/figure.png" pageId="19" pageNumber="20">59J</figureCitation>
)..............................
<taxonomicName id="4C294D5EFFFD071CFB9C961CFB7CA31C" ID-CoL="76XQY" baseAuthorityName="Swezey" baseAuthorityYear="1913" box="[1150,1269,532,554]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hibiscella">
<emphasis id="B95DEACFFFFD071CFB9C961CFB7CA31C" bold="true" box="[1150,1269,532,554]" italics="true" pageId="19" pageNumber="20">P. hibiscella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFAE3961CFA15A31C" ID-CoL="4G8P2" baseAuthorityName="Swezey" baseAuthorityYear="1928" box="[1281,1436,532,554]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="touchardiella">
<emphasis id="B95DEACFFFFD071CFAE3961CFA15A31C" bold="true" box="[1281,1436,532,554]" italics="true" pageId="19" pageNumber="20">P. touchardiella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF75963AFA15A371" blockId="19.[151,1437,151,936]" box="[151,1436,562,584]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF75963AFA15A371" box="[151,1436,562,584]" pageId="19" pageNumber="20">
- Signa very small with a pair of slender minute spines (
<figureCitation id="13122A58FFFD071CFD0F963BFCCDA37E" box="[749,836,562,584]" captionStart="FIGURE 59" captionStartId="110.[151,250,1714,1738]" captionTargetBox="[166,1426,193,1682]" captionTargetId="figure-23@110.[151,1436,181,1690]" captionTargetPageId="110" captionText="FIGURE 59. Female genitalia of Philodoria. A P. neraudicola, paralectotype, Oahu, SK708, host: Neraudia melastomifalia, B Same species, Maui, SK838, host: Pipturus albidus, C P. obamaorum, paratype, Hawaii (Big Island), SK710, host: Pipturus albidus, D Same species, Hawaii (Big Island), SK844 host: Pipturus sp., E P. urerana, paralectotype, SK672, F Philodoria sp. 1, SK835, G Same species, PHIL0021, H P. keahii, paratype, SK756, I P. funkae, holotype, SK761, J P. touchardiella, SK764, K P. keaensis, holotype, Hawaii (Big Island), SK777, L P. hauicola, Oahu, SK618." figureDoi="http://doi.org/10.5281/zenodo.4681937" httpUri="https://zenodo.org/record/4681937/files/figure.png" pageId="19" pageNumber="20">Fig. 59K</figureCitation>
)...................................
<emphasis id="B95DEACFFFFD071CFB17963BFA15A371" bold="true" box="[1269,1436,562,584]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFB17963BFAE9A37E" ID-CoL="8VM3Z" box="[1269,1376,562,584]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="keaensis" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFB17963BFAE9A37E" bold="true" box="[1269,1376,562,584]" italics="true" pageId="19" pageNumber="20">P. keaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFA84963BFA15A371" box="[1382,1436,562,583]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF759659FA15A352" blockId="19.[151,1437,151,936]" box="[151,1436,591,613]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759659FA15A352" box="[151,1436,591,613]" pageId="19" pageNumber="20">
- Signum very small with a minute spine (
<figureCitation id="13122A58FFFD071CFD859646FD52A353" box="[615,731,591,613]" captionStart="FIGURE 60" captionStartId="111.[151,250,921,945]" captionTargetBox="[157,1433,191,891]" captionTargetId="figure-20@111.[151,1437,181,897]" captionTargetPageId="111" captionText="FIGURE 60. Female genitalia of Philodoria. A P. marginestrigata, SK665, B Same species, SK772, host: Abutilon menziesi, C Same species, SK787, host: Sida fallax, D P. sciallactis, paralectotype, SK713, E P. ureraella, paralectotype, SK677, F P. opuhe, paratype, SK675." figureDoi="http://doi.org/10.5281/zenodo.4681939" httpUri="https://zenodo.org/record/4681939/files/figure.png" pageId="19" pageNumber="20">Fig. 60AC</figureCitation>
)...........................................
<taxonomicName id="4C294D5EFFFD071CFB059659FA15A352" ID-CoL="4G8NJ" baseAuthorityName="Walsingham" baseAuthorityYear="1907" box="[1255,1436,591,613]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="marginestrigata">
<emphasis id="B95DEACFFFFD071CFB059659FA15A352" bold="true" box="[1255,1436,591,613]" italics="true" pageId="19" pageNumber="20">P. marginestrigata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E78FFFD071CFF759664FA15A38B" pageId="19" pageNumber="20">
<paragraph id="8B9636DDFFFD071CFF759664FA15A396" blockId="19.[151,1437,151,936]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF759664FA15A396" pageId="19" pageNumber="20">
8 Ductus bursae short, about
<quantity id="4CD19B38FFFD071CFE0A9664FDBBA3B4" box="[488,562,621,642]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="mm" value="300.0">300 µm</quantity>
in length; lamella antevaginalis inflexed on posterior margin (
<figureCitation id="13122A58FFFD071CFB6B9664FB75A3B5" box="[1161,1276,621,643]" captionStart="FIGURE 57" captionStartId="107.[151,250,1554,1578]" captionTargetBox="[181,1405,189,1520]" captionTargetId="figure-20@107.[176,1411,181,1530]" captionTargetPageId="107" captionText="FIGURE 57. Female genitalia of Pipturus-mining Philodoria. A P. costalis, paralectotype, SK653 B Same species, paratype, SK652 C P. pipturiella, SK722 D P. pipturicola, paralectotype, lateral view, SK717 E Same species, paralectotype, SK716 F P. haelaauensis, SK792 G P. floscula, SK819." figureDoi="http://doi.org/10.5281/zenodo.4681933" httpUri="https://zenodo.org/record/4681933/files/figure.png" pageId="19" pageNumber="20">Fig. 57AE</figureCitation>
).................................................................................
<taxonomicName id="4C294D5EFFFD071CFBC59683FB01A396" ID-CoL="4G8N8" box="[1063,1160,650,672]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B95DEACFFFFD071CFBC59683FB01A396" bold="true" box="[1063,1160,650,672]" italics="true" pageId="19" pageNumber="20">P. costalis</emphasis>
</taxonomicName>
,
<emphasis id="B95DEACFFFFD071CFB769683FA15A396" bold="true" box="[1172,1436,650,672]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFB769683FA9AA396" ID-CoL="76Y2Y" authorityName="Swezey" authorityYear="1923" box="[1172,1299,650,672]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pipturiella">P. pipturiella</taxonomicName>
<taxonomicName id="4C294D5EFFFD071CFAFB9683FA15A396" ID-CoL="4G8NT" authorityName="Det. By O. H. Swezey" authorityYear="1915" box="[1305,1436,650,672]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pipturicola">P. pipturicola</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636DDFFFD071CFF7596A1FA15A38B" blockId="19.[151,1437,151,936]" box="[151,1436,679,701]" pageId="19" pageNumber="20">
<keyLead id="30D895E8FFFD071CFF7596A1FA15A38B" box="[151,1436,679,701]" pageId="19" pageNumber="20">
- Ductus bursae short, about
<quantity id="4CD19B38FFFD071CFE0A96A1FDBBA38B" box="[488,562,680,701]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="mm" value="300.0">300 µm</quantity>
in length; lamella antevaginalis semicircular (
<figureCitation id="13122A58FFFD071CFC0D96AEFBCFA38B" box="[1007,1094,679,701]" captionStart="FIGURE 58" captionStartId="109.[151,250,1622,1646]" captionTargetBox="[164,1427,193,1591]" captionTargetId="figure-20@109.[152,1435,181,1598]" captionTargetPageId="109" captionText="FIGURE 58. Female genitalia of Philodoria. A P. wilkesiella, SK761, B P. epibathra, SK679, C P. dubauticola, SK645, D Same species, PHIL0008, E P. dubautiella, SK692, F P. alakaiensis, paratype, SK659, G Same species, lateral view, H P. hibiscella, SK723, I P. naenaeiella, SK703, J Same species, SK834, K P. knudseniiella, paratype, SK863, L P. hesperomanniella, paratype, SK832, M Same species, SK620." figureDoi="http://doi.org/10.5281/zenodo.4681935" httpUri="https://zenodo.org/record/4681935/files/figure.png" pageId="19" pageNumber="20">Fig. 58A</figureCitation>
)................
<taxonomicName id="4C294D5EFFFD071CFAFF96A1FA15A38B" ID-CoL="4G8P5" authorityName="Sweszey" authorityYear="1940" box="[1309,1436,679,701]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="wilkesiella">
<emphasis id="B95DEACFFFFD071CFAFF96A1FA15A38B" bold="true" box="[1309,1436,679,701]" italics="true" pageId="19" pageNumber="20">P. wilkesiella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8B9636DDFFFD071CFF7596CCFB7EA21E" blockId="19.[151,1437,151,936]" pageId="19" pageNumber="20">
- Ductus bursae long, 400~µm in length (
<figureCitation id="13122A58FFFD071CFD8296CCFC9DA3EC" box="[608,788,709,731]" captionStart="FIGURE 58" captionStartId="109.[151,250,1622,1646]" captionTargetBox="[164,1427,193,1591]" captionTargetId="figure-20@109.[152,1435,181,1598]" captionTargetPageId="109" captionText="FIGURE 58. Female genitalia of Philodoria. A P. wilkesiella, SK761, B P. epibathra, SK679, C P. dubauticola, SK645, D Same species, PHIL0008, E P. dubautiella, SK692, F P. alakaiensis, paratype, SK659, G Same species, lateral view, H P. hibiscella, SK723, I P. naenaeiella, SK703, J Same species, SK834, K P. knudseniiella, paratype, SK863, L P. hesperomanniella, paratype, SK832, M Same species, SK620." figureDoi="http://doi.org/10.5281/zenodo.4681935" httpUri="https://zenodo.org/record/4681935/files/figure.png" pageId="19" pageNumber="20">Figs. 58BG, IM</figureCitation>
,
<figureCitation id="13122A58FFFD071CFCFF96CCFCD4A3ED" box="[797,861,709,731]" captionStart="FIGURE 59" captionStartId="110.[151,250,1714,1738]" captionTargetBox="[166,1426,193,1682]" captionTargetId="figure-23@110.[151,1436,181,1690]" captionTargetPageId="110" captionText="FIGURE 59. Female genitalia of Philodoria. A P. neraudicola, paralectotype, Oahu, SK708, host: Neraudia melastomifalia, B Same species, Maui, SK838, host: Pipturus albidus, C P. obamaorum, paratype, Hawaii (Big Island), SK710, host: Pipturus albidus, D Same species, Hawaii (Big Island), SK844 host: Pipturus sp., E P. urerana, paralectotype, SK672, F Philodoria sp. 1, SK835, G Same species, PHIL0021, H P. keahii, paratype, SK756, I P. funkae, holotype, SK761, J P. touchardiella, SK764, K P. keaensis, holotype, Hawaii (Big Island), SK777, L P. hauicola, Oahu, SK618." figureDoi="http://doi.org/10.5281/zenodo.4681937" httpUri="https://zenodo.org/record/4681937/files/figure.png" pageId="19" pageNumber="20">59AI</figureCitation>
,
<figureCitation id="13122A58FFFD071CFC8596CCFC22A3ED" box="[871,939,709,731]" captionStart="FIGURE 60" captionStartId="111.[151,250,921,945]" captionTargetBox="[157,1433,191,891]" captionTargetId="figure-20@111.[151,1437,181,897]" captionTargetPageId="111" captionText="FIGURE 60. Female genitalia of Philodoria. A P. marginestrigata, SK665, B Same species, SK772, host: Abutilon menziesi, C Same species, SK787, host: Sida fallax, D P. sciallactis, paralectotype, SK713, E P. ureraella, paralectotype, SK677, F P. opuhe, paratype, SK675." figureDoi="http://doi.org/10.5281/zenodo.4681939" httpUri="https://zenodo.org/record/4681939/files/figure.png" pageId="19" pageNumber="20">60DF</figureCitation>
).....................................................................................
<emphasis id="B95DEACFFFFD071CFCEC96EBFC59A3C1" bold="true" box="[782,976,738,760]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFCEC96EBFC1DA3CE" ID-CoL="8VM3V" box="[782,916,738,760]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alakaiensis" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFCEC96EBFC1DA3CE" bold="true" box="[782,916,738,760]" italics="true" pageId="19" pageNumber="20">P. alakaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFC7896EBFC59A3C1" box="[922,976,738,759]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C294D5EFFFD071CFC3E96EBFBE1A3CE" ID-CoL="4G8N9" baseAuthorityName="Swezey" baseAuthorityYear="1940" box="[988,1128,738,760]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dubauticola">
<emphasis id="B95DEACFFFFD071CFC3E96EBFBE1A3CE" bold="true" box="[988,1128,738,760]" italics="true" pageId="19" pageNumber="20">P. dubauticola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFB9796EBFB75A3CE" ID-CoL="6V9QY" baseAuthorityName="Swezey" baseAuthorityYear="1913" box="[1141,1276,738,760]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dubautiella">
<emphasis id="B95DEACFFFFD071CFB9796EBFB75A3CE" bold="true" box="[1141,1276,738,760]" italics="true" pageId="19" pageNumber="20">P. dubautiella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFAEA96EBFAF6A3CE" ID-CoL="4G8NB" baseAuthorityName="Walsingham" baseAuthorityYear="1907" box="[1288,1407,738,760]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="epibathra">
<emphasis id="B95DEACFFFFD071CFAEA96EBFAF6A3CE" bold="true" box="[1288,1407,738,760]" italics="true" pageId="19" pageNumber="20">P. epibathra</emphasis>
</taxonomicName>
,
<emphasis id="B95DEACFFFFD071CFA6996EBFE1FA223" bold="true" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFA6996EBFED3A223" ID-CoL="8VM47" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obamaorum" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFA6996EBFED3A223" bold="true" italics="true" pageId="19" pageNumber="20">P. obamaorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFE829709FE1FA223" box="[352,406,768,789]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFD071CFE409709FCC5A223" bold="true" box="[418,844,767,789]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFE409709FDE3A223" ID-CoL="8VM3Y" box="[418,618,767,789]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hesperomanniella" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFE409709FDE3A223" bold="true" box="[418,618,767,789]" italics="true" pageId="19" pageNumber="20">P. hesperomanniella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFD939709FD21A223" box="[625,680,768,789]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName id="4C294D5EFFFD071CFD519709FC86A223" ID-CoL="8VM3W" box="[691,783,767,789]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="funkae" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFD519709FC86A223" bold="true" box="[691,783,767,789]" italics="true" pageId="19" pageNumber="20">P. funkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFCF49709FCC5A223" box="[790,844,768,789]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFD071CFCBA9709FC60A223" bold="true" box="[856,1001,767,789]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFCBA9709FC25A223" ID-CoL="8VM42" box="[856,940,767,789]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="keahii" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFCBA9709FC25A223" bold="true" box="[856,940,767,789]" italics="true" pageId="19" pageNumber="20">P. keahii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFC509709FC60A223" box="[946,1001,768,789]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFD071CFC169709FB47A223" bold="true" box="[1012,1230,767,789]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFC169709FB18A223" ID-CoL="8VM43" box="[1012,1169,767,789]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="knudseniiella" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFC169709FB18A223" bold="true" box="[1012,1169,767,789]" italics="true" pageId="19" pageNumber="20">P. knudseniiella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFB759709FB47A223" box="[1175,1230,768,789]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFD071CFB3B9709FEDCA204" bold="true" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFB3B9709FAF6A223" ID-CoL="4G8NG" baseAuthorityName="Swezey" baseAuthorityYear="1940" box="[1241,1407,767,789]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lipochaetaella">
<emphasis id="B95DEACFFFFD071CFB3B9709FB61A223" bold="true" box="[1241,1256,768,789]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="B95DEACFFFFD071CFB1696F6FAF6A223" bold="true" box="[1268,1407,767,789]" italics="true" pageId="19" pageNumber="20">lipochaetaella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFA699709FEDCA204" ID-CoL="4G8NN" baseAuthorityName="Swezey" baseAuthorityYear="1940" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="naenaeiella">
<emphasis id="B95DEACFFFFD071CFA699709FEDCA204" bold="true" italics="true" pageId="19" pageNumber="20">P. naenaeiella</emphasis>
</taxonomicName>
</emphasis>
,
<taxonomicName id="4C294D5EFFFD071CFE839714FE66A204" ID-CoL="4G8NP" baseAuthorityName="Swezey" baseAuthorityYear="1920" box="[353,495,796,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="neraudicola">
<emphasis id="B95DEACFFFFD071CFE839714FE66A204" bold="true" box="[353,495,796,818]" italics="true" pageId="19" pageNumber="20">P. neraudicola</emphasis>
</taxonomicName>
,
<emphasis id="B95DEACFFFFD071CFE199714FCADA204" bold="true" box="[507,804,796,818]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFE199714FD0FA204" ID-CoL="4G8NR" baseAuthorityName="Swezey" baseAuthorityYear="1946" box="[507,646,796,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrelloides">
<emphasis id="B95DEACFFFFD071CFE199714FD0FA204" bold="true" box="[507,646,796,818]" italics="true" pageId="19" pageNumber="20">P. nigrelloides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFD709714FD6EA204" ID-CoL="8VM48" box="[658,743,796,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="opuhe" status="sp. nov.">
<emphasis id="B95DEACFFFFD071CFD709714FD6EA204" bold="true" box="[658,743,796,818]" italics="true" pageId="19" pageNumber="20">P. opuhe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57B4FFFD071CFD0F9714FCADA204" box="[749,804,797,818]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B95DEACFFFFD071CFCCD9714FBAEA204" bold="true" box="[815,1063,796,818]" pageId="19" pageNumber="20">
<taxonomicName id="4C294D5EFFFD071CFCCD9714FC21A204" ID-CoL="4G8NW" baseAuthorityName="Meyrick" baseAuthorityYear="1928" box="[815,936,796,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="sciallactis">
<emphasis id="B95DEACFFFFD071CFCCD9714FC21A204" bold="true" box="[815,936,796,818]" italics="true" pageId="19" pageNumber="20">P. sciallactis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D5EFFFD071CFC569714FBAEA204" ID-CoL="4G8P3" baseAuthorityName="Swezey" baseAuthorityYear="1915" box="[948,1063,796,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ureraella">
<emphasis id="B95DEACFFFFD071CFC569714FBAEA204" bold="true" box="[948,1063,796,818]" italics="true" pageId="19" pageNumber="20">P. ureraella</emphasis>
</taxonomicName>
</emphasis>
,
<taxonomicName id="4C294D5EFFFD071CFBD09714FB13A204" ID-CoL="4G8P4" baseAuthorityName="Swezey" baseAuthorityYear="1915" box="[1074,1178,797,818]" class="Insecta" family="Gracillariidae" genus="Philodoria" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="urerana">
<emphasis id="B95DEACFFFFD071CFBD09714FB13A204" bold="true" box="[1074,1178,797,818]" italics="true" pageId="19" pageNumber="20">P. urerana</emphasis>
</taxonomicName>
,
<emphasis id="B95DEACFFFFD071CFB449714FB7EA21E" bold="true" box="[1190,1271,796,818]" pageId="19" pageNumber="20">
<emphasis id="B95DEACFFFFD071CFB449714FB3CA204" bold="true" box="[1190,1205,797,818]" italics="true" pageId="19" pageNumber="20">P</emphasis>
. sp. 1
<superScript id="7C5C9B95FFFD071CFB0D9715FB7EA21E" attach="left" box="[1263,1271,796,808]" fontSize="5" pageId="19" pageNumber="20"></superScript>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3336556FFFD071CFF759733FB14A29E" pageId="19" pageNumber="20" type="multiple">
<paragraph id="8B9636DDFFFD071CFF759733FB14A29E" blockId="19.[151,1437,151,936]" pageId="19" pageNumber="20">
<superScript id="7C5C9B95FFFD071CFF759733FF14A270" attach="left" box="[151,157,826,838]" fontSize="5" pageId="19" pageNumber="20"></superScript>
Females of the 18 species have the following similar characters: lamella antevaginalis weakly sclerotized, widening toward anterior margin of A7; Ductus bursae posterior extremity widening and membranous, and middle part weakly sclerotized, round and flat; terminus of ductus bursae tubular, biforked. Corpus bursae pyriform or oblong; signa absent; paired rows consisting of wrinkles running longitudinally. See also key to adults and male genitalia for species identification.
</paragraph>
</subSubSection>
</treatment>
</document>