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<document id="7CEE1BD8B5965B9F895AC27D2D34B874" ID-DOI="10.11646/zootaxa.3716.2.8" ID-GBIF-Dataset="703c8417-4446-4e68-9ff2-537f42339b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="221980" ID-ZooBank="D975C77F-83DB-48C6-A698-84A24665D838" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460681307907" checkinUser="plazi" docAuthor="Fikáček, Martin, Maruyama, Munetoshi, Vondráček, Dominik &amp; Short, Andrew E. Z." docDate="2013" docId="038087C28A3D1435FF76FCA1FD2EF878" docLanguage="en" docName="zt03716p288.pdf" docOrigin="Zootaxa 3716 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chimaerocyon Fikáček, Maruyama, Vondráček &amp; Short, 2013, gen. nov." docType="treatment" docVersion="8" lastPageNumber="280" masterDocId="FFB9FFBA8A3F1436FFE1FFBDFFBAFFC3" masterDocTitle="Chimaerocyon gen. nov., a morphologically aberrant myrmecophilous genus of water scavenger beetle (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="288" masterPageNumber="277" pageNumber="279" updateTime="1698350497756" updateUser="plazi">
<mods:mods id="947ADBA2E09A66E412978B8248F5FA21" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3A785B2A02D264691D9DF9BAE1F795B4">
<mods:title id="C8397112B472048CE0EF05C40F68E545">Chimaerocyon gen. nov., a morphologically aberrant myrmecophilous genus of water scavenger beetle (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
</mods:titleInfo>
<mods:name id="9E6B1ED812A654620C409C64C14FD1B4" type="personal">
<mods:role id="A8FF2A28F772DB0EC025B7DE1304C399">
<mods:roleTerm id="52A511B19C1497B76B8DFAC5E932C1FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB28700653418B792CF544FFC4608D42">Fikáček, Martin</mods:namePart>
</mods:name>
<mods:name id="6322ED4C99E6D2DB68826DBF163120FA" type="personal">
<mods:role id="06EC76B63631729D7F5A07B234E7EB95">
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</mods:role>
<mods:namePart id="A95041F027671C73F29B6890D49DB256">Maruyama, Munetoshi</mods:namePart>
</mods:name>
<mods:name id="519C390135CF4670770A152AB2EFC65E" type="personal">
<mods:role id="4F5599F9B2579E4A06C88357687267FD">
<mods:roleTerm id="3F5C04E4DBCBF214DEA7B3AF815B61A1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="305141AC669010B180AE7B608637A5FD">Vondráček, Dominik</mods:namePart>
</mods:name>
<mods:name id="2AE0416A34D50E0B71274C28ED501823" type="personal">
<mods:role id="5C86FEA7A8E1A22F8D42207E23125A0E">
<mods:roleTerm id="07E423B354A2116A0507655C44349EC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4E5449D52D0FBB3CD75C53E019D317F">Short, Andrew E. Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5A3BE89A074629EA52F1CA446B475FAF">text</mods:typeOfResource>
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<mods:titleInfo id="F2C48897DC8B01DCEB9C9C3ED980199C">
<mods:title id="B86FE278143CADD2BAD58CC0EACDEFA5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="120F714DF21C3E73BAB4A12EE96CF77B">
<mods:date id="D3C18DE88E17F14D32F772035A9774E9">2013</mods:date>
<mods:detail id="C5BFF5B2D97D4F413BA38E78B02949BB" type="volume">
<mods:number id="BB82C6016785E7D7B78472281F8E6767">3716</mods:number>
</mods:detail>
<mods:detail id="959565E736DF6805C55C51E5C6774A06" type="issue">
<mods:number id="E8D46271FD0DC5390525C1DC91D8B090">2</mods:number>
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<mods:start id="0D6D86CDA1963F853D2BE3421AA9679F">277</mods:start>
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<mods:classification id="57D59FDA8E2F6BE4D89871EE2B5B5EFE">journal article</mods:classification>
<mods:identifier id="4182A237276346E007725C4D20E183C8" type="DOI">10.11646/zootaxa.3716.2.8</mods:identifier>
<mods:identifier id="68C38C058824B533AB9628497C2723BD" type="GBIF-Dataset">703c8417-4446-4e68-9ff2-537f42339b5a</mods:identifier>
<mods:identifier id="56ABABB5E5197187C59FFEEAEA9E3A17" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AEBEFFEE7A67E554FE6A3F529F6BF800" type="Zenodo-Dep">221980</mods:identifier>
<mods:identifier id="A11E3D976C62285CF4E3CBE18A593AF2" type="ZooBank">D975C77F-83DB-48C6-A698-84A24665D838</mods:identifier>
</mods:mods>
<treatment id="038087C28A3D1435FF76FCA1FD2EF878" ID-DOI="http://doi.org/10.5281/zenodo.6149745" ID-GBIF-Taxon="119579971" ID-Zenodo-Dep="6149745" LSID="urn:lsid:plazi:treatment:038087C28A3D1435FF76FCA1FD2EF878" httpUri="http://treatment.plazi.org/id/038087C28A3D1435FF76FCA1FD2EF878" lastPageId="3" lastPageNumber="280" pageId="2" pageNumber="279">
<subSubSection id="C333655F8A3D1434FF76FCA1FEA5FC94" pageId="2" pageNumber="279" type="nomenclature">
<paragraph id="8B9636D48A3D1434FF76FCA1FE07FCF5" blockId="2.[151,445,796,855]" box="[151,445,796,822]" pageId="2" pageNumber="279">
<heading id="D0DE81B88A3D1434FF76FCA1FE07FCF5" bold="true" box="[151,445,796,822]" fontSize="11" level="1" pageId="2" pageNumber="279" reason="1">
<emphasis id="B95DEAC68A3D1434FF76FCA1FE07FCF5" bold="true" box="[151,445,796,822]" pageId="2" pageNumber="279">
<taxonomicName id="4C294D578A3D1434FF76FCA1FEF3FCF5" ID-CoL="8X775" box="[151,329,796,822]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95DEAC68A3D1434FF76FCA1FEF3FCF5" bold="true" box="[151,329,796,822]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57BD8A3D1434FEAEFCA1FE07FCF5" box="[335,445,796,822]" pageId="2" pageNumber="279" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9636D48A3D1434FF76FC83FEA5FC94" blockId="2.[151,445,796,855]" box="[151,287,830,855]" pageId="2" pageNumber="279">
(
<figureCitation id="13122A518A3D1434FF7EFC83FEADFC94" box="[159,279,830,855]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 14" captionStart-2="FIGURES 15 – 22" captionStart-3="FIGURES 23 – 30" captionStartId-0="4.[151,264,1928,1950]" captionStartId-1="5.[151,264,1470,1492]" captionStartId-2="6.[151,264,1567,1589]" captionStartId-3="7.[151,264,1910,1932]" captionTargetBox-0="[171,1417,240,1878]" captionTargetBox-1="[151,1436,205,1448]" captionTargetBox-2="[151,1436,193,1545]" captionTargetBox-3="[151,1436,482,1887]" captionTargetId-0="figure@4.[151,1436,195,1906]" captionTargetId-1="figure@5.[151,1436,193,1449]" captionTargetId-2="figure@6.[151,1436,193,1546]" captionTargetId-3="figure@7.[151,1436,481,1888]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." captionText-1="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." captionText-2="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." captionText-3="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri-0="https://zenodo.org/record/221981/files/figure.png" httpUri-1="https://zenodo.org/record/221982/files/figure.png" httpUri-2="https://zenodo.org/record/221983/files/figure.png" httpUri-3="https://zenodo.org/record/221984/files/figure.png" pageId="2" pageNumber="279">Figs. 1–30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333655F8A3D1434FF76FC3BFC72FC5C" box="[151,968,902,927]" pageId="2" pageNumber="279" type="materials_examined">
<paragraph id="8B9636D48A3D1434FF76FC3BFC72FC5C" blockId="2.[151,1436,902,2007]" box="[151,968,902,927]" pageId="2" pageNumber="279">
<emphasis id="B95DEAC68A3D1434FF76FC3BFE88FC5C" bold="true" box="[151,306,902,927]" pageId="2" pageNumber="279">
<typeStatus id="549288768A3D1434FF76FC3BFF68FC5C" box="[151,210,902,927]" pageId="2" pageNumber="279">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C294D578A3D1434FED8FC3BFDF1FC5D" box="[313,587,902,926]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="shimadai" status="sp. nov.">
<emphasis id="B95DEAC68A3D1434FED8FC3BFDF1FC5D" box="[313,587,902,926]" italics="true" pageId="2" pageNumber="279">Chimaerocyon shimadai</emphasis>
</taxonomicName>
<emphasis id="B95DEAC68A3D1434FDB0FC3BFD11FC5C" bold="true" box="[593,683,902,927]" pageId="2" pageNumber="279">
<taxonomicNameLabel id="A26E57BD8A3D1434FDB0FC3BFD11FC5C" box="[593,683,902,927]" pageId="2" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(by present designation).
</paragraph>
</subSubSection>
<subSubSection id="C333655F8A3D1434FF26FC17FD1BF9E4" pageId="2" pageNumber="279" type="diagnosis">
<paragraph id="8B9636D48A3D1434FF26FC17FD1BF9E4" blockId="2.[151,1436,902,2007]" pageId="2" pageNumber="279">
<emphasis id="B95DEAC68A3D1434FF26FC17FEFBFC00" bold="true" box="[199,321,938,963]" pageId="2" pageNumber="279">Diagnosis.</emphasis>
<taxonomicName id="4C294D578A3D1434FEADFC16FE4AFC00" box="[332,496,939,963]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FEADFC16FE4AFC00" box="[332,496,939,963]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
largely resembles the representatives of the sphaeridiine tribes
<taxonomicName id="4C294D578A3D1434FB3BFC17FA85FC00" box="[1242,1343,938,963]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Omicrini">Omicrini</taxonomicName>
(by the general thoracic and elytral morphology) and
<taxonomicName id="4C294D578A3D1434FD76FC73FCF8FC24" box="[663,834,974,999]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Coelostomatini">Coelostomatini</taxonomicName>
(by the head and elytral morphology).
<taxonomicName id="4C294D578A3D1434FB19FC73FA26FC25" box="[1272,1436,974,998]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95DEAC68A3D1434FB19FC73FA26FC25" box="[1272,1436,974,998]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57BD8A3D1434FF76FC4FFEB9FBC8" box="[151,259,1010,1035]" pageId="2" pageNumber="279" rank="genus">
<emphasis id="B95DEAC68A3D1434FF76FC4FFEB9FBC8" bold="true" box="[151,259,1010,1035]" pageId="2" pageNumber="279">gen. nov.</emphasis>
</taxonomicNameLabel>
is easily distinguishable from all known genera of
<taxonomicName id="4C294D578A3D1434FCB6FC4FFC4BFBC8" box="[855,1009,1010,1035]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="subFamily" subFamily="Sphaeridiinae">Sphaeridiinae</taxonomicName>
by the combination of the following characters: (1) elytral disc without striae or series of punctures including sutural stria, with remnants of 4–5 series preserved only subapically (
<figureCitation id="13122A518A3D1434FE39FB87FDFCFB90" box="[472,582,1082,1107]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="2" pageNumber="279">Figs. 1, 4</figureCitation>
,
<figureCitation id="13122A518A3D1434FDB5FB87FD24FB90" box="[596,670,1082,1107]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="2" pageNumber="279">26–27</figureCitation>
; nearly unique within
<taxonomicName id="4C294D578A3D1434FC7EFB87FBFEFB90" box="[927,1092,1082,1107]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
, only shared with
<taxonomicName id="4C294D578A3D1434FAF6FB86FA31FB91" box="[1303,1419,1083,1106]" class="Insecta" family="Hydrophilidae" genus="Litrosurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FAF6FB86FA31FB91" box="[1303,1419,1083,1106]" italics="true" pageId="2" pageNumber="279">Litrosurus</emphasis>
</taxonomicName>
); (2) abdominal ventrite 5 with sexually dimorphic emargination (
<figureCitation id="13122A518A3D1434FC54FBE3FBFDFBB4" box="[949,1095,1118,1143]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Figs. 12–14</figureCitation>
; similar sexually dimorphic emargination only present in the genus
<taxonomicName id="4C294D578A3D1434FDB5FB3EFD03FB58" box="[596,697,1155,1179]" class="Insecta" family="Hydrophilidae" genus="Psalitrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FDB5FB3EFD03FB58" box="[596,697,1155,1179]" italics="true" pageId="2" pageNumber="279">Psalitrus</emphasis>
</taxonomicName>
; a small emargination in some
<taxonomicName id="4C294D578A3D1434FBFDFB3FFB7DFB58" box="[1052,1223,1154,1179]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Coelostomatini">Coelostomatini</taxonomicName>
is fringed by stout setae absent in
<taxonomicName id="4C294D578A3D1434FEAFFB1BFE48FB7D" box="[334,498,1190,1214]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FEAFFB1BFE48FB7D" box="[334,498,1190,1214]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C294D578A3D1434FDD0FB1BFD2CFB7D" box="[561,662,1190,1214]" class="Insecta" family="Hydrophilidae" genus="Psalitrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FDD0FB1BFD2CFB7D" box="[561,662,1190,1214]" italics="true" pageId="2" pageNumber="279">Psalitrus</emphasis>
</taxonomicName>
). Additional characters distinguishing
<taxonomicName id="4C294D578A3D1434FBBDFB1BFABAFB7D" box="[1116,1280,1190,1214]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3D1434FBBDFB1BFABAFB7D" box="[1116,1280,1190,1214]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
from part of sphaeridiine genera include: (1) male maxilla with sucking discs (
<figureCitation id="13122A518A3D1434FC7FFB77FC48FB20" box="[926,1010,1226,1251]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Figs. 6</figureCitation>
,
<figureCitation id="13122A518A3D1434FBE3FB77FBF5FB20" box="[1026,1103,1226,1251]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">16, 19</figureCitation>
; absent in all
<taxonomicName id="4C294D578A3D1434FB1BFB77FA26FB20" box="[1274,1436,1226,1251]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
except
<taxonomicName id="4C294D578A3D1434FF07FB53FED5FAC4" box="[230,367,1262,1287]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Sphaeridiini">Sphaeridiini</taxonomicName>
and
<taxonomicName id="4C294D578A3D1434FE45FB53FD87FAC4" box="[420,573,1262,1287]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
); (2) prothoracic hypomeron without antennal grooves (
<figureCitation id="13122A518A3D1434FB4AFB53FB43FAC4" box="[1195,1273,1262,1287]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Figs. 6</figureCitation>
,
<figureCitation id="13122A518A3D1434FAE4FB52FA9EFAC4" box="[1285,1316,1263,1287]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">17</figureCitation>
; present in most
<taxonomicName id="4C294D578A3D1434FF37FAAFFED5FAE8" box="[214,367,1298,1323]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
); (3) prosternum extremely shortened in front of procoxae (
<figureCitation id="13122A518A3D1434FBF8FAAFFBD0FAE8" box="[1049,1130,1298,1323]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Figs. 6</figureCitation>
,
<figureCitation id="13122A518A3D1434FB99FAAEFB2DFAE8" box="[1144,1175,1299,1323]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">17</figureCitation>
; not shortened in most
<taxonomicName id="4C294D578A3D1434FF76FA8BFE83FA8C" box="[151,313,1334,1359]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
except many
<taxonomicName id="4C294D578A3D1434FE3EFA8BFDFDFA8C" box="[479,583,1334,1359]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Omicrini">Omicrini</taxonomicName>
, Amphiopini and
<taxonomicName id="4C294D578A3D1434FCFBFA8BFCC4FA8C" box="[794,894,1334,1359]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Berosini">Berosini</taxonomicName>
); (4) mesothorax with mesoventrite fused with mesanepisterna, but without grooves for reception of procoxae (
<figureCitation id="13122A518A3D1434FC94FAE7FBBBFAB0" box="[885,1025,1370,1395]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="2" pageNumber="279">Figs. 23–24</figureCitation>
; this combination is only present in
<taxonomicName id="4C294D578A3D1434FF76FAC3FF46FA54" box="[151,252,1406,1431]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Omicrini">Omicrini</taxonomicName>
and some
<taxonomicName id="4C294D578A3D1434FE6DFAC3FD87FA54" box="[396,573,1406,1431]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Coelostomatini">Coelostomatini</taxonomicName>
); (5) epipleuron wide throughout elytral length (
<figureCitation id="13122A518A3D1434FB70FAC3FB53FA54" box="[1169,1257,1406,1431]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Figs. 6</figureCitation>
,
<figureCitation id="13122A518A3D1434FB1CFAC2FAA6FA54" box="[1277,1308,1407,1431]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="2" pageNumber="279">23</figureCitation>
; absent in
<taxonomicName id="4C294D578A3D1434FF76FA1FFE91FA78" box="[151,299,1442,1467]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
and
<taxonomicName id="4C294D578A3D1434FE88FA1FFE4DFA78" box="[361,503,1442,1467]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Sphaeridiini">Sphaeridiini</taxonomicName>
); (6) median spur situated subapically of the M-Cu loop of hind wing (
<figureCitation id="13122A518A3D1434FAA0FA1FFA2CFA78" box="[1345,1430,1442,1467]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="2" pageNumber="279">Fig. 11</figureCitation>
; unique synapomorphy of the
<taxonomicName id="4C294D578A3D1434FE09FA7BFD3BFA1C" box="[488,641,1478,1503]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
); (7) antennal club compact (
<figureCitation id="13122A518A3D1434FC32FA7BFBE7FA1C" box="[979,1117,1478,1503]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Figs. 16–17</figureCitation>
; loosely segmented in most
<taxonomicName id="4C294D578A3D1434FF76FA57FE83F9C0" box="[151,313,1514,1539]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
except many
<taxonomicName id="4C294D578A3D1434FE3AFA57FDC0F9C0" box="[475,634,1514,1539]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="subFamily" subFamily="Sphaeridiinae">Sphaeridiinae</taxonomicName>
). Detailed discussion of the assignment of
<taxonomicName id="4C294D578A3D1434FB84FA56FAB3F9C0" box="[1125,1289,1515,1539]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95DEAC68A3D1434FB84FA56FAB3F9C0" box="[1125,1289,1515,1539]" italics="true" pageId="2" pageNumber="279">Chimaerocyon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57BD8A3D1434FAF3FA57FAC7F9C0" box="[1298,1405,1514,1539]" pageId="2" pageNumber="279" rank="genus">
<emphasis id="B95DEAC68A3D1434FAF3FA57FAC7F9C0" bold="true" box="[1298,1405,1514,1539]" pageId="2" pageNumber="279">gen. nov.</emphasis>
</taxonomicNameLabel>
to the
<taxonomicName id="4C294D578A3D1434FF21F9B3FEEEF9E4" box="[192,340,1550,1575]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
is provided in the discussion.
</paragraph>
</subSubSection>
<subSubSection id="C333655F8A3D1435FF26F98FFD8BF8E9" lastPageId="3" lastPageNumber="280" pageId="2" pageNumber="279" type="description">
<paragraph id="8B9636D48A3D1434FF26F98FFA26F988" blockId="2.[151,1436,902,2007]" box="[199,1436,1586,1611]" pageId="2" pageNumber="279">
<emphasis id="B95DEAC68A3D1434FF26F98FFEE3F988" bold="true" box="[199,345,1586,1611]" pageId="2" pageNumber="279">Description.</emphasis>
Body widely oval to subtrapezoid, highly convex, Coloration reddish brown to black (
<figureCitation id="13122A518A3D1434FAC2F98FFA34F988" box="[1315,1422,1586,1611]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="2" pageNumber="279">Figs. 1–5</figureCitation>
).
</paragraph>
<paragraph id="8B9636D48A3D1435FF26F9EBFD81FE6E" blockId="2.[151,1436,902,2007]" lastBlockId="3.[151,1436,151,1979]" lastPageId="3" lastPageNumber="280" pageId="2" pageNumber="279">
<emphasis id="B95DEAC68A3D1434FF26F9EBFEB1F9AD" box="[199,267,1622,1646]" italics="true" pageId="2" pageNumber="279">Head.</emphasis>
Anterior margin of clypeus subangular, median portion rather narrow, anterolateral portion explanate, covering antennal bases (
<figureCitation id="13122A518A3D1434FE5CF9C7FDF7F950" box="[445,589,1658,1683]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Figs. 15, 18</figureCitation>
); marginal bead wide and distinct; frontoclypeal suture distinct laterally, reduced medially; transverse ridges absent. Median portion of frons and clypeus not elevated above remaining surface. Whole dorsal surface with fine and dense punctation, punctures lacking setae. Eyes small, situated on lateral angular portions of head, with dorsal portion smaller than ventral one, divided by 7.5–10× the width of one eye in dorsal view. Labrum ca. 0.4× as wide as head, membranous, largely retracted under anterior margin of clypeus, weakly bisinuate on anterior margin, densely pubescent dorsally (
<figureCitation id="13122A518A3D1434FC05F893FB82F884" box="[996,1080,1838,1863]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Fig. 15</figureCitation>
). Maxilla of male with sucking disc on galea (
<figureCitation id="13122A518A3D1434FEDDF8EFFE72F8A8" box="[316,456,1874,1899]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Figs. 16, 19</figureCitation>
); maxillary palpus with basal palpomere minute, palpomere 2 large, widened apically, ca. as long as palpomere 4, palpomere 3 ca. half as long as palpomere 4, slightly widening apicad, palpomere 4 cylindrical, without digitiform sensilla (
<figureCitation id="13122A518A3D1434FDB9F827FD16F870" box="[600,684,1946,1971]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Fig. 16</figureCitation>
). Mentum (
<figureCitation id="13122A518A3D1434FCD1F827FC02F870" box="[816,952,1946,1971]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="2" pageNumber="279">Figs. 19, 22</figureCitation>
) transverse, ca. 2× wider than long, lateral margins without fringe of setae, anterior margin weakly bisinuate; premental lobes large, with long marginal setae; labial palps trimerous, basal palpomere very short, palpomeres 2 and 3 subequal in length, palpomere 2 with densely arranged setae. Antenna (
<figureCitation id="13122A518A3C1435FDFAFF01FDD5FF16" box="[539,623,188,213]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="3" pageNumber="280">Fig. 16</figureCitation>
) with 9 antennomeres; scape long, cylindrical, curved basally; pedicel short, bulbous basally; antennomeres 3–5 short, subequal in length, slightly widening distad; cupule slightly asymmetrical, slightly larger than antennomere 5; antennomeres 6–9 forming an elongate pubescent club, antennomere 7 shortest, antennomeres 8–9 subequal in length, antennomere slightly constricted subapically, blunt at apex; special sensorial antennal fields absent. Submentum nearly bare, gular sutures vaguely developed, widely separated from each other, tentorial pits minute. Genal ridge long, arising from inner margin of eye and mesally reaching the level of maxillary cardo.
</paragraph>
<paragraph id="8B9636D48A3C1435FF26FE05FDC5FD0E" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26FE05FEFBFE13" box="[199,321,440,464]" italics="true" pageId="3" pageNumber="280">Prothorax.</emphasis>
Pronotum transverse, highly convex, distinctly wider than bases of elytra combined; lateral margins narrowly beaded; punctation uniform, consisting of fine transverse punctures, punctures lacking setae except for those along lateral margins; series of larger punctures along posterior margin absent. Prosternum (
<figureCitation id="13122A518A3C1435FAF2FE42FADDFDDB" box="[1299,1383,511,536]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Figs. 6</figureCitation>
,
<figureCitation id="13122A518A3C1435FA97FDBDFA29FDDB" box="[1398,1427,512,536]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="3" pageNumber="280">17</figureCitation>
) narrow, as wide as ca. 0.3 of total width of prothorax, extremely short in front of procoxae; its median portion highly elevated into minute triangular plate (
<figureCitation id="13122A518A3C1435FD43FDFAFD43FDA3" box="[674,761,583,608]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="3" pageNumber="280">Fig. 21</figureCitation>
). Hypomeron extremely large, divided into inner sparsely pubescent portion and wide lateral bare portion; antennal groove absent (
<figureCitation id="13122A518A3C1435FC29FDD1FBA6FD47" box="[968,1052,620,645]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="3" pageNumber="280">Fig. 17</figureCitation>
). Transverse fold behind posterior margin of pronotum absent. Profurca rather massive, with crescent-like apical portion (
<figureCitation id="13122A518A3C1435FB90FD32FB7CFD6B" box="[1137,1222,655,680]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="3" pageNumber="280">Fig. 20</figureCitation>
). Procoxal cavities contiguous mesally, open medioposteriorly.
</paragraph>
<paragraph id="8B9636D48A3C1435FF26FD65FC10FBF6" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26FD65FEEEFD33" box="[199,340,728,752]" italics="true" pageId="3" pageNumber="280">Mesothorax.</emphasis>
Mesoventrite slightly longer than metaventrite, but inclined from the plane of metaventrite and hence appearing approximately as long as metaventrite (
<figureCitation id="13122A518A3C1435FCEBFD41FCE7FCD7" box="[778,861,764,789]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 23</figureCitation>
). Mesoventrite with a low median carina projecting to a short rhomboid elevation posteriorly; grooves for reception of procoxae absent; anapleural sutures absent, mesoventrite fused with mesanepisterna (
<figureCitation id="13122A518A3C1435FD93FCF9FD45FC9F" box="[626,767,836,861]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Figs. 23–24</figureCitation>
). Mesepimeron very narrow, widening laterad. Mesocoxal cavities very narrowly separated. Mesofurca not examined. Scutellar shield in shape of small equilateral triangle (
<figureCitation id="13122A518A3C1435FF7EFC31FEB1FC67" box="[159,267,908,933]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="3" pageNumber="280">Figs. 1, 4</figureCitation>
). Elytra (
<figureCitation id="13122A518A3C1435FE92FC31FE65FC67" box="[371,479,908,933]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="3" pageNumber="280">Figs. 1, 4</figureCitation>
,
<figureCitation id="13122A518A3C1435FE0AFC31FD88FC66" box="[491,562,908,933]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">26–27</figureCitation>
) narrowly to widely explanate laterally, punctural series absent on elytral disc as well as laterally, but distinctly developed, deeply impressed and hence forming a more or less deep depression mesally before elytral apex; epipleuron horizontal, very wide especially at junction of meso- and metathorax, gradually narrowing posteriad but still well developed at apex, indistinctly subdivided by a fine ridge into narrow lateral and wide mesal portions, its whole surface lacking pubescence.
</paragraph>
<paragraph id="8B9636D48A3C1435FF26FBFDFAA1FACE" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26FBFDFEEBFB9B" box="[199,337,1088,1112]" italics="true" pageId="3" pageNumber="280">Metathorax.</emphasis>
Metaventrite only indistinctly elevated mesally, anteromesally projecting into narrow but rather long metaventral process (
<figureCitation id="13122A518A3C1435FE21FBD9FDAFFBBE" box="[448,533,1124,1149]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 25</figureCitation>
), with a narrow bead behind mesocoxal cavities; femoral lines and anterolateral ridges absent. Metanepisternum ca. 5× as long as wide, without anterior oblique ridge, metepimeron with minute ventral portion. Metafurca not examined. Metathoracic wings (
<figureCitation id="13122A518A3C1435FC8CFB11FC7AFB07" box="[877,960,1196,1221]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Fig. 11</figureCitation>
) well developed, with transverse vein r4 arising from basal portion of radial cell,
<collectingCountry id="F33E76448A3C1435FD85FB6DFD3CFB24" box="[612,646,1232,1255]" name="Poland" pageId="3" pageNumber="280">RP</collectingCountry>
rather long, reaching ca. halfway to wing base, basal cubito-anal cell small, close, wedge cell absent, transverse vein mp-cua joining to MP3+4+CuA1+2; anal lobe not defined.
</paragraph>
<paragraph id="8B9636D48A3C1435FF26FAA6FB85F9EC" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26FAA6FF46FAF1" box="[199,252,1307,1330]" italics="true" pageId="3" pageNumber="280">Legs</emphasis>
generally rather short, with tips of femora not overlapping the outline of body (
<figureCitation id="13122A518A3C1435FB71FAA7FB45FAF0" box="[1168,1279,1306,1331]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="3" pageNumber="280">Figs. 1, 4</figureCitation>
,
<figureCitation id="13122A518A3C1435FAECFAA7FAA7FAF0" box="[1293,1309,1306,1331]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">6</figureCitation>
). Procoxae large, subglobular, slightly transverse, sparsely pubescent; meso- and metacoxae massive, transverse. Femora rather massive, with large tibial groove defined by high dorsal and ventral ridges (
<figureCitation id="13122A518A3C1435FBAAFADFFB1BFAB8" box="[1099,1185,1378,1403]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 23</figureCitation>
), lacking pubescence; profemur without impressed parts; metafemur extremely enlarged in both sexes (
<figureCitation id="13122A518A3C1435FBA4FA3BFB26FA5C" box="[1093,1180,1414,1439]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 23</figureCitation>
). Pro- and mesotibiae conical, with thin and short lateral spines and reduced mesal spines; metatibiae slightly depressed. Tarsi pentamerous, with extremely shortened but wide tarsomeres (
<figureCitation id="13122A518A3C1435FC8DFA73FC7EFA24" box="[876,964,1486,1511]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 28</figureCitation>
), each tarsomere with tuft of stiff setae ventrally and a few fine setae dorsally; metatarsomere 1 slightly longer than tarsomere 2, tarsomeres 2–4 subequal in length, tarsomere 5 longest. Claws simple, arcuate; empodium bisetose (
<figureCitation id="13122A518A3C1435FC3EF9ABFB89F9EC" box="[991,1075,1558,1583]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="3" pageNumber="280">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8B9636D48A3C1435FF26F986FD8BF8E9" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26F986FE8BF990" box="[199,305,1595,1619]" italics="true" pageId="3" pageNumber="280">Abdomen</emphasis>
with five ventrites. Ventrite 1 with low mesal carina developed only basally, hence seemingly ecarinate (
<figureCitation id="13122A518A3C1435FEEDF9E3FEEBF9B4" box="[268,337,1630,1655]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Fig. 6</figureCitation>
). Ventrites 2–5 ecarinate mesally. Apical portion of ventrite 5 narrowly emarginate in both sexes, shape of the emargination species-specific and sexually dimorphic (
<figureCitation id="13122A518A3C1435FC68F93FFBA8F958" box="[905,1042,1666,1691]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Figs. 12–14</figureCitation>
). Male sternite 8 crescent-like (
<figureCitation id="13122A518A3C1435FA8FF93FFF1DF97C" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Fig. 9</figureCitation>
), without median narrow projection on anterior margin, with two rows of setae on posterior margin not meeting mesally. Male sternite 9 tongue-like mesally (
<figureCitation id="13122A518A3C1435FD7BF977FD54F920" box="[666,750,1738,1763]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Fig. 10</figureCitation>
). Aedeagus (
<figureCitation id="13122A518A3C1435FC60F977FC7EF920" box="[897,964,1738,1763]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="3" pageNumber="280">Fig. 8</figureCitation>
) simple, median lobe with complex basal hooks, not protruding into phallobase, phallobase short, strongly asymmetrical; parameres simple. Female genitalia corresponding with those of
<taxonomicName id="4C294D578A3C1435FE36F8AEFD90F8E8" box="[471,554,1811,1835]" class="Insecta" family="Hydrophilidae" genus="Kanala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAC68A3C1435FE36F8AEFD90F8E8" box="[471,554,1811,1835]" italics="true" pageId="3" pageNumber="280">Kanala</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C333655F8A3C1435FF26F88BFD2EF878" pageId="3" pageNumber="280" type="etymology">
<paragraph id="8B9636D48A3C1435FF26F88BFD2EF878" blockId="3.[151,1436,151,1979]" pageId="3" pageNumber="280">
<emphasis id="B95DEAC68A3C1435FF26F88BFEF4F88C" bold="true" box="[199,334,1846,1871]" pageId="3" pageNumber="280">Etymology.</emphasis>
The generic name is derived from
<emphasis id="B95DEAC68A3C1435FD3DF88AFCFCF88C" box="[732,838,1847,1871]" italics="true" pageId="3" pageNumber="280">chimaera</emphasis>
(Greek, mythical animal with body parts taken from various animals) which reflects the fact that the genus combines the diagnostic characters of several sphaeridiine tribes, and the suffix
<emphasis id="B95DEAC68A3C1435FE65F83DFE78F854" box="[388,450,1920,1943]" italics="true" pageId="3" pageNumber="280">-cyon</emphasis>
, a typical suffix of generic names within the tribe
<taxonomicName id="4C294D578A3C1435FC19F8C3FB36F854" box="[1016,1164,1918,1943]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="280" phylum="Arthropoda" rank="tribe" tribe="Megasternini">Megasternini</taxonomicName>
to which the new genus belongs. The new generic name is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>
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