291 lines
39 KiB
XML
291 lines
39 KiB
XML
<document id="9DE65F954A78E712D8094A9A9438E19D" ID-DOI="10.11646/zootaxa.3716.2.8" ID-GBIF-Dataset="703c8417-4446-4e68-9ff2-537f42339b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="221980" ID-ZooBank="D975C77F-83DB-48C6-A698-84A24665D838" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460681307907" checkinUser="plazi" docAuthor="Fikáček, Martin, Maruyama, Munetoshi, Vondráček, Dominik & Short, Andrew E. Z." docDate="2013" docId="038087C28A3A143EFF76F905FC40FC66" docLanguage="en" docName="zt03716p288.pdf" docOrigin="Zootaxa 3716 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chimaerocyon shimadai Fikáček, Maruyama, Vondráček & Short, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="285" masterDocId="FFB9FFBA8A3F1436FFE1FFBDFFBAFFC3" masterDocTitle="Chimaerocyon gen. nov., a morphologically aberrant myrmecophilous genus of water scavenger beetle (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="288" masterPageNumber="277" pageNumber="282" updateTime="1698350497756" updateUser="plazi">
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<mods:mods id="105DA98DAF02CE0514D7391032EB4EE2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AFBBE478AAAADB03BCBEFC6DA361DB47">
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<mods:title id="88F93C0F4B2DE2EFDDB6688CFD824AA7">Chimaerocyon gen. nov., a morphologically aberrant myrmecophilous genus of water scavenger beetle (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="8E3D28FFF31986A5026C003944DEEC18" type="personal">
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<mods:role id="6AC98D7166D842ABA977FEA45F1D3865">
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<mods:roleTerm id="9605ED68065D6051DAF257A92911DC11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FAF98F7412B3BDA1FC4E8185E555E566">Fikáček, Martin</mods:namePart>
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</mods:name>
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<mods:roleTerm id="FDB616CAC41AAD0C4EB43F5FA01ADC30">Author</mods:roleTerm>
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<mods:namePart id="0703B3BDEBB5A8828EEC33B184724776">Maruyama, Munetoshi</mods:namePart>
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</mods:name>
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<mods:name id="7ECAF47011B1621FF116BA2B938074D6" type="personal">
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<mods:roleTerm id="8535FCF809B0807570F895B2FFCC5F0A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5E970E31A7E289BB1756B4D5D302ED95">Vondráček, Dominik</mods:namePart>
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</mods:name>
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<mods:name id="0CA4B7C7329E6B5BE2BA24ED8E5A117E" type="personal">
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<mods:roleTerm id="B0D01B9FC8FE1E8C5407AFAE464AAA3E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3BF7897739993AAEB6B1DBAE400BA081">Short, Andrew E. Z.</mods:namePart>
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<mods:typeOfResource id="0ACC31AF2860C53C8BD8BAC083CCE3B0">text</mods:typeOfResource>
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<mods:title id="B9B0D758FF7C148D33A77BD8DCA4F8AD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6D07553E67E3F14F8249AFEE2BF96FAE">
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<mods:date id="89D78C6B7DC47D5DEB438B00EB1A97CB">2013</mods:date>
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<mods:detail id="81D4ABDA4B956D73F2F274350B2CC90F" type="volume">
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<mods:number id="B722BB81E50F2DEB5D925AABAC30A2EE">3716</mods:number>
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<mods:number id="01FE56384B3B55642483FF32458A456F">2</mods:number>
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<mods:classification id="2D6C0D8B2FC904A0C454880BC163AC62">journal article</mods:classification>
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<mods:identifier id="C92FBE11E23845F3FC8B6EA50866E646" type="DOI">10.11646/zootaxa.3716.2.8</mods:identifier>
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<mods:identifier id="BD1DAE5F144B74BBFF218EB59198EA30" type="GBIF-Dataset">703c8417-4446-4e68-9ff2-537f42339b5a</mods:identifier>
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<mods:identifier id="3C7420841BCA63604EA2686425395670" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7BC2D7EA82930D03D2F275AD3899F693" type="Zenodo-Dep">221980</mods:identifier>
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<mods:identifier id="54AE0CDBE08C4226113A26F11E0F7F08" type="ZooBank">D975C77F-83DB-48C6-A698-84A24665D838</mods:identifier>
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<treatment id="038087C28A3A143EFF76F905FC40FC66" ID-DOI="http://doi.org/10.5281/zenodo.6149747" ID-GBIF-Taxon="119579973" ID-Zenodo-Dep="6149747" LSID="urn:lsid:plazi:treatment:038087C28A3A143EFF76F905FC40FC66" httpUri="http://treatment.plazi.org/id/038087C28A3A143EFF76F905FC40FC66" lastPageId="8" lastPageNumber="285" pageId="5" pageNumber="282">
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<subSubSection id="C333655F8A3A1433FF76F905FDEEF937" pageId="5" pageNumber="282" type="nomenclature">
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<paragraph id="8B9636D48A3A1433FF76F905FD99F910" blockId="5.[151,596,1720,1780]" box="[151,547,1720,1747]" pageId="5" pageNumber="282">
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<heading id="D0DE81B88A3A1433FF76F905FD99F910" bold="true" box="[151,547,1720,1747]" fontSize="11" level="1" pageId="5" pageNumber="282" reason="1">
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<emphasis id="B95DEAC68A3A1433FF76F905FD99F910" bold="true" box="[151,547,1720,1747]" pageId="5" pageNumber="282">
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<taxonomicName id="4C294D578A3A1433FF76F905FE05F911" ID-CoL="5Y48M" box="[151,447,1720,1746]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="shimadai" status="sp. nov.">
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<emphasis id="B95DEAC68A3A1433FF76F905FE05F911" bold="true" box="[151,447,1720,1746]" italics="true" pageId="5" pageNumber="282">Chimaerocyon shimadai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26E57BD8A3A1433FE24F904FD99F910" box="[453,547,1721,1747]" pageId="5" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9636D48A3A1433FF76F966FDEEF937" blockId="5.[151,596,1720,1780]" box="[151,596,1755,1780]" pageId="5" pageNumber="282">
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||
(
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<figureCitation id="13122A518A3A1433FF7EF966FEB0F937" box="[159,266,1755,1780]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="5" pageNumber="282">Figs. 1–3</figureCitation>
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,
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<figureCitation id="13122A518A3A1433FEF7F966FEEBF937" box="[278,337,1755,1780]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="5" pageNumber="282">6–13</figureCitation>
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,
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<figureCitation id="13122A518A3A1433FEBCF966FE1EF937" box="[349,420,1755,1780]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="5" pageNumber="282">15–21</figureCitation>
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,
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<figureCitation id="13122A518A3A1433FE51F966FDF1F937" box="[432,587,1755,1780]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="5" pageNumber="282">23–26, 28–30</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C333655F8A3A1433FF76F89EFD15F80F" pageId="5" pageNumber="282" type="materials_examined">
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<paragraph id="8B9636D48A3A1433FF76F89EFBEEF8A3" blockId="5.[151,1437,1827,1996]" pageId="5" pageNumber="282">
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<emphasis id="B95DEAC68A3A1433FF76F89EFEFEF8FF" bold="true" box="[151,324,1827,1852]" pageId="5" pageNumber="282">
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<typeStatus id="549288768A3A1433FF76F89EFF68F8FF" box="[151,210,1827,1852]" pageId="5" pageNumber="282">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="549288768A3A1433FEAAF89EFE02F8FF" box="[331,440,1827,1852]" pageId="5" pageNumber="282" type="holotype">Holotype</typeStatus>
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: male (KUZC):
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<collectingCountry id="F33E76448A3A1433FD8DF899FCB8F8FF" box="[620,770,1828,1852]" name="Malaysia" pageId="5" pageNumber="282">MALAYSIA</collectingCountry>
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: Pahang / Bukit Fraser (
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<quantity id="4CD19B318A3A1433FBECF89EFBDDF8FF" box="[1037,1127,1827,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="282" unit="m" value="1250.0">1250 m</quantity>
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), /
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<date id="FF9710148A3A1433FB68F89EFABEF8FF" box="[1161,1284,1827,1852]" pageId="5" pageNumber="282" value="2011-04-25">25.IV.2011</date>
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/ SHIMADA Taku.
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<typeStatus id="549288768A3A1433FF3DF8F5FEF7F8A3" box="[220,333,1864,1888]" pageId="5" pageNumber="282" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D2FFD5D8A3A1433FEB9F8F5FE0BF8A3" box="[344,433,1863,1888]" pageId="5" pageNumber="282" type="male">2 males</specimenCount>
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,
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<specimenCount id="9D2FFD5D8A3A1433FE5DF8F5FDA7F8A3" box="[444,541,1863,1888]" pageId="5" pageNumber="282" type="female">1 female</specimenCount>
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(KUZC, NMPC): same label data as the
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<typeStatus id="549288768A3A1433FC0AF8FAFBEBF8A3" box="[1003,1105,1863,1888]" pageId="5" pageNumber="282" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8B9636D48A3A1433FF26F8D6FD15F80F" blockId="5.[151,1437,1827,1996]" pageId="5" pageNumber="282">
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<emphasis id="B95DEAC68A3A1433FF26F8D6FD52F847" bold="true" box="[199,744,1899,1924]" pageId="5" pageNumber="282">
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DNA isolate and sequences of the
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<typeStatus id="549288768A3A1433FD9AF8D6FD5EF847" box="[635,740,1899,1924]" pageId="5" pageNumber="282" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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The DNA isolate of the
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<typeStatus id="549288768A3A1433FBFCF8D6FBC5F847" box="[1053,1151,1899,1924]" pageId="5" pageNumber="282" type="holotype">holotype</typeStatus>
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is deposited in NMPC. Sequence data were submitted into GenBank under the following accession numbers:
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<emphasis id="B95DEAC68A3A1433FB93F82DFB1DF86B" box="[1138,1191,1936,1960]" italics="true" pageId="5" pageNumber="282">cox1</emphasis>
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—
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<accessionNumber id="947AAB378A3A1433FB25F832FA84F86B" box="[1220,1342,1935,1960]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF623548" pageId="5" pageNumber="282">KF623548</accessionNumber>
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;
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<emphasis id="B95DEAC68A3A1433FAAAF82DFAC5F86B" box="[1355,1407,1936,1960]" italics="true" pageId="5" pageNumber="282">cox2</emphasis>
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—
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<accessionNumber id="947AAB378A3A1433FF76F80EFEA9F80F" box="[151,275,1971,1996]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF623549" pageId="5" pageNumber="282">KF623549</accessionNumber>
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;
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<emphasis id="B95DEAC68A3A1433FEFCF809FEFDF80F" box="[285,327,1972,1996]" italics="true" pageId="5" pageNumber="282">18S</emphasis>
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—
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<accessionNumber id="947AAB378A3A1433FE84F80EFE65F80F" box="[357,479,1971,1996]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF623550" pageId="5" pageNumber="282">KF623550</accessionNumber>
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;
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<emphasis id="B95DEAC68A3A1433FE0BF809FDAEF80F" box="[490,532,1972,1996]" italics="true" pageId="5" pageNumber="282">28S</emphasis>
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—
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<accessionNumber id="947AAB378A3A1433FDD3F80EFD11F80F" box="[562,683,1971,1996]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF623551" pageId="5" pageNumber="282">KF623551</accessionNumber>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF56665C8A391430FF76F9A2FE61F952" httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="6" pageNumber="283" targetBox="[151,1436,193,1545]" targetPageId="6">
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<paragraph id="8B9636D48A391430FF76F9A2FE61F952" blockId="6.[151,1436,1567,1681]" pageId="6" pageNumber="283">
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<emphasis id="B95DEAC68A391430FF76F9A2FEEEF9F6" bold="true" box="[151,340,1567,1589]" pageId="6" pageNumber="283">FIGURES 15–22.</emphasis>
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Detailed morphology of
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<taxonomicName id="4C294D578A391430FDB3F9A2FCF3F9F6" box="[594,841,1567,1589]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="shimadai">
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<emphasis id="B95DEAC68A391430FDB3F9A2FCF3F9F6" box="[594,841,1567,1589]" italics="true" pageId="6" pageNumber="283">Chimaerocyon shimadai</emphasis>
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</taxonomicName>
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(15–21) and
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<taxonomicName id="4C294D578A391430FC31F99DFBD9F9F6" box="[976,1123,1568,1589]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="sumatranus">
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<emphasis id="B95DEAC68A391430FC31F99DFBD9F9F6" box="[976,1123,1568,1589]" italics="true" pageId="6" pageNumber="283">C. sumatranus</emphasis>
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</taxonomicName>
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(22). 15—head in dorsal view, 16—head in ventral view, 17—prothorax and head in ventral view, 18—clypeus in lateral view, 19, 22—mentum, 20— profurca, 21—elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32):
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<emphasis id="B95DEAC68A391430FAF1F9E1FA95F9B1" bold="true" box="[1296,1327,1628,1650]" pageId="6" pageNumber="283">cly</emphasis>
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—clypeus,
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<emphasis id="B95DEAC68A391430FF76F9C6FF75F952" bold="true" box="[151,207,1659,1681]" pageId="6" pageNumber="283">msuc</emphasis>
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—male sucker on maxilla.
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</paragraph>
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</caption>
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<subSubSection id="C333655F8A391431FF26F905FE9AFE6F" lastPageId="7" lastPageNumber="284" pageId="6" pageNumber="283" type="description">
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<paragraph id="8B9636D48A391431FF26F905FE9AFE6F" blockId="6.[151,1437,1720,2033]" lastBlockId="7.[151,1437,151,429]" lastPageId="7" lastPageNumber="284" pageId="6" pageNumber="283">
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<emphasis id="B95DEAC68A391430FF26F905FEE3F912" bold="true" box="[199,345,1720,1745]" pageId="6" pageNumber="283">Description.</emphasis>
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Body length 2.6–2.9 mm (2.7 mm in
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<typeStatus id="549288768A391430FCECF905FCC9F912" box="[781,883,1720,1745]" pageId="6" pageNumber="283" type="holotype">holotype</typeStatus>
|
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), body width 1.9–2.2 mm (2.0 mm in
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<typeStatus id="549288768A391430FAC8F905FA2AF912" box="[1321,1424,1720,1745]" pageId="6" pageNumber="283" type="holotype">holotype</typeStatus>
|
||
). Body widest at anterior margin of elytra, strongly narrowing posteriad; elytra slightly roof-like in anterior view. Coloration black with widely reddish margins of clypeus, elytra and lateral and sutural margins of elytra; ventral surface and appendages reddish brown. Head with dense punctation consisting of crescent-like to semicircular punctures, without microsculpture on interstices; punctation on pronotum and elytra as dense as that on head, consisting of slightly finer punctures; pronotal punctures transverse, scar-like, elytral punctures dot-like. Eyes divided by 7.5× the width of one eye in dorsal view. Mentum with sparse moderately strong punctation. Elytron with remnants of four or five striae posteromesally situated in deep depression; lateral portion of elytra widely explanate. Mesoventral plate rhomboid, slightly longer than wide, with slightly concave margins. Median portion of metaventrite with fine but dense punctation bearing enlarged setae posteromesally in male, uniformly pubescent in female. Abdominal ventrites with dense and rather coarse setiferous punctation, bearing longer and slightly denser pubescence around apical emargination in both sexes. Abdominal ventrite 5 widely emarginate apically with posterior margin of emargination irregularly convex in male, narrowly and simply emarginate in female. Metatarsus extremely short, with metatarsomere 1 ca. 1.5× as long as tarsomere 2. Male sternite 9 with numerous stout setae apically. Aedeagus 1.3 mm long, phallobase ca. 0.5× as long as paramere, strongly asymmetrical; paramere weakly narrowing from base to apical fourth, membranous, flat and slightly expanded apically, without apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF56665C8A381431FF76F8CBFA93F80A" httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="7" pageNumber="284" targetBox="[151,1436,482,1887]" targetPageId="7">
|
||
<paragraph id="8B9636D48A381431FF76F8CBFA93F80A" blockId="7.[151,1436,1910,1993]" pageId="7" pageNumber="284">
|
||
<emphasis id="B95DEAC68A381431FF76F8CBFEEFF84F" bold="true" box="[151,341,1910,1932]" pageId="7" pageNumber="284">FIGURES 23–30.</emphasis>
|
||
Detailed morphology of
|
||
<taxonomicName id="4C294D578A381431FDB8F8CBFCE8F84F" box="[601,850,1910,1932]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="shimadai">
|
||
<emphasis id="B95DEAC68A381431FDB8F8CBFCE8F84F" box="[601,850,1910,1932]" italics="true" pageId="7" pageNumber="284">Chimaerocyon shimadai</emphasis>
|
||
</taxonomicName>
|
||
(23–26, 28–30) and
|
||
<taxonomicName id="4C294D578A381431FBCAF8CAFB05F84F" box="[1067,1215,1911,1932]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A381431FBCAF8CAFB05F84F" box="[1067,1215,1911,1932]" italics="true" pageId="7" pageNumber="284">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
(27). 23—meso- and metathorax in ventral view, 24—mesoventrite in lateral view, 26—median portions of meso- and metaventrite in ventral view, 26–27—right elytral apex, 28—metatarsus, 29—metatarsal claws, 30—detail of apical portion of elytral series 1–2.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C333655F8A37143EFF26FF2AFCADFD0E" pageId="8" pageNumber="285" type="diagnosis">
|
||
<paragraph id="8B9636D48A37143EFF26FF2AFCADFD0E" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="285">
|
||
<emphasis id="B95DEAC68A37143EFF26FF2AFEFBFF73" bold="true" box="[199,321,151,176]" pageId="8" pageNumber="285">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C294D578A37143EFEAFFF25FDDCFF73" box="[334,614,152,176]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="shimadai" status="sp. nov.">
|
||
<emphasis id="B95DEAC68A37143EFEAFFF25FDDCFF73" box="[334,614,152,176]" italics="true" pageId="8" pageNumber="285">Chimaerocyon shimadai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26E57BD8A37143EFD92FF2AFD69FF73" box="[627,723,151,176]" pageId="8" pageNumber="285" rank="species">
|
||
<emphasis id="B95DEAC68A37143EFD92FF2AFD69FF73" bold="true" box="[627,723,151,176]" pageId="8" pageNumber="285">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
may be distinguished from
|
||
<taxonomicName id="4C294D578A37143EFBCDFF25FB6CFF73" box="[1068,1238,152,176]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
|
||
<emphasis id="B95DEAC68A37143EFBCDFF25FB6CFF73" box="[1068,1238,152,176]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95DEAC68A37143EFB02FF2AFAF9FF73" bold="true" box="[1251,1347,151,176]" pageId="8" pageNumber="285">
|
||
<taxonomicNameLabel id="A26E57BD8A37143EFB02FF2AFAF9FF73" box="[1251,1347,151,176]" pageId="8" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the combination of the following characters: (1) the dorsal surface is piceous with widely reddish margins of elytra, pronotum and clypeus (
|
||
<figureCitation id="13122A518A37143EFE48FF62FDACFF3B" box="[425,534,223,248]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="8" pageNumber="285">Figs. 1–3</figureCitation>
|
||
) (dorsal surface is uniformly reddish in
|
||
<taxonomicName id="4C294D578A37143EFC13FF5DFB23FF3B" box="[1010,1177,224,248]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFC13FF5DFB23FF3B" box="[1010,1177,224,248]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13122A518A37143EFB4BFF62FAA0FF3B" box="[1194,1306,223,248]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="8" pageNumber="285">Figs. 4–5</figureCitation>
|
||
); (2) elytra highly arched, forming a pointed arch in anterior view (
|
||
<figureCitation id="13122A518A37143EFCCCFEB9FCCEFEDF" box="[813,884,260,285]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="8" pageNumber="285">Fig. 3</figureCitation>
|
||
) (elytra widely rounded in anterior view in
|
||
<taxonomicName id="4C294D578A37143EFA63FEB9FEA0FE83" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFA63FEB9FEA0FE83" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13122A518A37143EFECEFE9AFEC0FE83" box="[303,378,295,320]" captionStart="FIGURES 1 – 5" captionStartId="4.[151,264,1928,1950]" captionTargetBox="[171,1417,240,1878]" captionTargetId="figure@4.[151,1436,195,1906]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Habitus of Chimaerocyon species. 1 – 3 — male paratype of Chimaerocyon shimadai (1 — dorsal view, 2 — lateral view, 3 — frontal view). 4 – 5 — holotype of C. sumatranus (4 — dorsal view, 5 — frontal view)." httpUri="https://zenodo.org/record/221981/files/figure.png" pageId="8" pageNumber="285">Fig. 5</figureCitation>
|
||
); (3) the emargination of abdominal ventrite 5 sexually dimorphic in shape, wide and subrectangular in male, narrowly semicircular in female (
|
||
<figureCitation id="13122A518A37143EFCB9FEF1FC52FEA7" box="[856,1000,332,357]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="8" pageNumber="285">Figs. 12–13</figureCitation>
|
||
) (widely bisinuate in female in
|
||
<taxonomicName id="4C294D578A37143EFA63FEF1FEA0FE4B" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFA63FEF1FEA0FE4B" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13122A518A37143EFECCFED2FE3EFE4B" box="[301,388,367,392]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="8" pageNumber="285">Fig. 14</figureCitation>
|
||
), male is unknown for the latter species); (4) both sexes with longer setae around apical emargination of abdominal ventrite 5 (
|
||
<figureCitation id="13122A518A37143EFDA4FE29FD71FE6F" box="[581,715,404,429]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="8" pageNumber="285">Figs. 12–13</figureCitation>
|
||
) (without such setae in
|
||
<taxonomicName id="4C294D578A37143EFC35FE29FBCDFE6F" box="[980,1143,404,428]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFC35FE29FBCDFE6F" box="[980,1143,404,428]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13122A518A37143EFB64FE29FB6DFE6F" box="[1157,1239,404,429]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="8" pageNumber="285">Fig. 14</figureCitation>
|
||
); (5) mesoventral plate as long as wide or only slightly longer, rhomboid in shape (ca. 1.4× longer than wide, with strongly concave sides in
|
||
<taxonomicName id="4C294D578A37143EFF1BFE61FE1BFE37" box="[250,417,476,500]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFF1BFE61FE1BFE37" box="[250,417,476,500]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
). Besides these characters, there are also slight differences in the number of impressed series at the elytral apex (4–5 series in
|
||
<taxonomicName id="4C294D578A37143EFDAAFDBDFD68FDDB" box="[587,722,512,536]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="shimadai">
|
||
<emphasis id="B95DEAC68A37143EFDAAFDBDFD68FDDB" box="[587,722,512,536]" italics="true" pageId="8" pageNumber="285">C. shimadai</emphasis>
|
||
</taxonomicName>
|
||
, 5 series in
|
||
<taxonomicName id="4C294D578A37143EFCBBFDBDFC47FDDB" box="[858,1021,512,536]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFCBBFDBDFC47FDDB" box="[858,1021,512,536]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13122A518A37143EFBEDFE42FB28FDDB" box="[1036,1170,511,536]" captionStart="FIGURES 23 – 30" captionStartId="7.[151,264,1910,1932]" captionTargetBox="[151,1436,482,1887]" captionTargetId="figure@7.[151,1436,481,1888]" captionTargetPageId="7" captionText="FIGURES 23 – 30. Detailed morphology of Chimaerocyon shimadai (23 – 26, 28 – 30) and C. sumatranus (27). 23 — meso- and metathorax in ventral view, 24 — mesoventrite in lateral view, 26 — median portions of meso- and metaventrite in ventral view, 26 – 27 — right elytral apex, 28 — metatarsus, 29 — metatarsal claws, 30 — detail of apical portion of elytral series 1 – 2." httpUri="https://zenodo.org/record/221984/files/figure.png" pageId="8" pageNumber="285">Figs. 26–27</figureCitation>
|
||
) and in the character of the punctation of the mentum (
|
||
<figureCitation id="13122A518A37143EFE12FD99FDC1FDFF" box="[499,635,548,573]" captionStart="FIGURES 15 – 22" captionStartId="6.[151,264,1567,1589]" captionTargetBox="[151,1436,193,1545]" captionTargetId="figure@6.[151,1436,193,1546]" captionTargetPageId="6" captionText="FIGURES 15 – 22. Detailed morphology of Chimaerocyon shimadai (15 – 21) and C. sumatranus (22). 15 — head in dorsal view, 16 — head in ventral view, 17 — prothorax and head in ventral view, 18 — clypeus in lateral view, 19, 22 — mentum, 20 — profurca, 21 — elevated mesal portion of prosternum. Abbreviations (numbers refer to those used in Fig. 32): cly — clypeus, msuc — male sucker on maxilla." httpUri="https://zenodo.org/record/221983/files/figure.png" pageId="8" pageNumber="285">Figs. 19, 22</figureCitation>
|
||
) and the metaventrite (moderately coarse in
|
||
<taxonomicName id="4C294D578A37143EFB95FD99FB47FDFF" box="[1140,1277,548,572]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="shimadai">
|
||
<emphasis id="B95DEAC68A37143EFB95FD99FB47FDFF" box="[1140,1277,548,572]" italics="true" pageId="8" pageNumber="285">C. shimadai</emphasis>
|
||
</taxonomicName>
|
||
, consisting of fine punctures in
|
||
<taxonomicName id="4C294D578A37143EFEBCFDF5FDB8FDA3" box="[349,514,584,608]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFEBCFDF5FDB8FDA3" box="[349,514,584,608]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
), but it is not clear at present to what extent these characters are intraspecifically variable due to the low number of specimens examined. We expect that both species also differ in the morphology of the aedeagus and the shape of the apical emargination of male abdominal ventrite 5, but this may be only confirmed once the male of
|
||
<taxonomicName id="4C294D578A37143EFE33FD09FDCFFD0F" box="[466,629,692,716]" class="Insecta" family="Hydrophilidae" genus="Chimaerocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="sumatranus">
|
||
<emphasis id="B95DEAC68A37143EFE33FD09FDCFFD0F" box="[466,629,692,716]" italics="true" pageId="8" pageNumber="285">C. sumatranus</emphasis>
|
||
</taxonomicName>
|
||
is discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C333655F8A37143EFF26FD6AFB07FD33" box="[199,1213,727,752]" pageId="8" pageNumber="285" type="etymology">
|
||
<paragraph id="8B9636D48A37143EFF26FD6AFB07FD33" blockId="8.[151,1437,151,933]" box="[199,1213,727,752]" pageId="8" pageNumber="285">
|
||
<emphasis id="B95DEAC68A37143EFF26FD6AFEF4FD33" bold="true" box="[199,334,727,752]" pageId="8" pageNumber="285">Etymology.</emphasis>
|
||
The species is dedicated to Mr. Taku Shimada, the collector of the
|
||
<typeStatus id="549288768A37143EFBDFFD65FBD4FD33" box="[1086,1134,728,752]" pageId="8" pageNumber="285">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C333655F8A37143EFF26FD41FB08FC43" pageId="8" pageNumber="285" type="biology_ecology">
|
||
<paragraph id="8B9636D48A37143EFF26FD41FA27FCF4" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="285">
|
||
<emphasis id="B95DEAC68A37143EFF26FD41FE9DFCD6" bold="true" box="[199,295,764,789]" pageId="8" pageNumber="285">Biology.</emphasis>
|
||
The beetles were collected from a brood cell of a nest of
|
||
<taxonomicName id="4C294D578A37143EFC2CFD41FA26FCD7" authority="Ozdikmen, 2010" authorityName="Ozdikmen" authorityYear="2010" box="[973,1436,764,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="singaporensis">
|
||
<emphasis id="B95DEAC68A37143EFC2CFD41FB6FFCD7" box="[973,1237,764,788]" italics="true" pageId="8" pageNumber="285">Pheidole singaporensis</emphasis>
|
||
Özdikmen, 2010
|
||
</taxonomicName>
|
||
(replacement name for
|
||
<taxonomicName id="4C294D578A37143EFE7AFC9DFCA9FCFB" authority="Smith, 1857" authorityName="Smith" authorityYear="1857" box="[411,787,799,824]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B95DEAC68A37143EFE7AFC9DFDDEFCFB" box="[411,612,800,824]" italics="true" pageId="8" pageNumber="285">Pheidole longipes</emphasis>
|
||
(Smith, 1857))
|
||
</taxonomicName>
|
||
. The nest was in a cliff along a forest trail in Bukit Fraser.
|
||
</paragraph>
|
||
<paragraph id="8B9636D48A37143EFF26FCF9FB08FC43" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="285">
|
||
The only female available had a single egg (
|
||
<figureCitation id="13122A518A37143EFD01FCF9FC93FC9F" box="[736,809,836,861]" captionStart="FIGURES 6 – 14" captionStartId="5.[151,264,1470,1492]" captionTargetBox="[151,1436,205,1448]" captionTargetId="figure@5.[151,1436,193,1449]" captionTargetPageId="5" captionText="FIGURES 6 – 14. Morphological details of Chimaerocyon shimadai (6 – 13) and C. sumatranus (14). 6 — ventral view, 7 — egg drawn to scale with Fig. 6, 8 — aedeagus of the holotype, 9 — male sternite 8 of the holotype, 10 — male sternite 9, 11 — hind wing of the holotype, 12 – 14 — abdominal apex (12 — male holotype of C. shimadai, 13 — female paratype of C. shimadai, 14 — female holotype of C. sumatranus). Abbreviations (numbers refer to characters in Fig. 32): msuc — male suckers on maxilla, elpr — elevated median portion of prosternum, wepi — wide epipleuron posteriorly, shtars — shortened tarsus, abdem — abdominal emargination." httpUri="https://zenodo.org/record/221982/files/figure.png" pageId="8" pageNumber="285">Fig. 7</figureCitation>
|
||
) with completely developed chorion in its abdomen, measuring 0.85×0.35 mm and hence occupying a large part of the internal abdominal volume.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C333655F8A37143EFF26FC31FC40FC66" box="[199,1018,908,933]" pageId="8" pageNumber="285" type="distribution">
|
||
<paragraph id="8B9636D48A37143EFF26FC31FC40FC66" blockId="8.[151,1437,151,933]" box="[199,1018,908,933]" pageId="8" pageNumber="285">
|
||
<emphasis id="B95DEAC68A37143EFF26FC31FEDAFC66" bold="true" box="[199,352,908,933]" pageId="8" pageNumber="285">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="549288768A37143EFD81FC30FD2AFC66" box="[608,656,909,933]" pageId="8" pageNumber="285">type</typeStatus>
|
||
locality in Peninsular
|
||
<collectingCountry id="F33E76448A37143EFC6DFC31FC4CFC66" box="[908,1014,908,933]" name="Malaysia" pageId="8" pageNumber="285">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |