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<document id="1C31C50B3F2DB09D17D0661BF4555B21" ID-DOI="10.5281/zenodo.12996779" ID-Zenodo-Dep="12996779" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="luisschmitz" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1721870935690" checkinUser="luisschmitz" docAuthor="Gerry Cassis &amp; Loren Vanags" docDate="2006" docId="0380878FFF86FFA6FDD0FAFEC67CFE58" docLanguage="en" docName="Denisia.19.275-398.pdf.imf" docOrigin="Denisia 19" docTitle="Scutellerinae" docType="treatment" docVersion="11" lastPageNumber="334" masterDocId="FFB9FFF7FFBFFF9DFFC4FF8CC239FFED" masterDocTitle="Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)" masterLastPageNumber="398" masterPageNumber="275" pageNumber="332" updateTime="1725646846613" updateUser="ExternalLinkService">
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<mods:title id="849C09AEF2EC9487E48EDD05F803ED4F">Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)</mods:title>
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<mods:namePart id="4745162D25E9AB6FA33308CEB6278EB0">Gerry Cassis</mods:namePart>
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<mods:namePart id="6A705C42575C36C77A8A5D3466EA4A3B">Loren Vanags</mods:namePart>
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<mods:title id="D620EEF5C058C21E38937CD6B150E800">Denisia</mods:title>
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<mods:date id="0F1710517D8A84CD90446D277B33AE1E">2006</mods:date>
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<treatment id="0380878FFF86FFA6FDD0FAFEC67CFE58" LSID="urn:lsid:plazi:treatment:0380878FFF86FFA6FDD0FAFEC67CFE58" httpUri="http://treatment.plazi.org/id/0380878FFF86FFA6FDD0FAFEC67CFE58" lastPageId="59" lastPageNumber="334" pageId="57" pageNumber="332">
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<paragraph id="8B963699FF86FFA4FDD0FAFEC124FA67" blockId="57.[532,966,624,1917]" box="[532,797,1394,1418]" pageId="57" pageNumber="332">
Subfamily
<taxonomicName id="4C294D1AFF86FFA4FD4FFAFEC124FA67" box="[651,797,1394,1418]" class="Insecta" family="Scutelleridae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="subFamily" subFamily="Scutellerinae">Scutellerinae</taxonomicName>
</paragraph>
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<subSubSection id="C3336512FF86FFA4FDD0FA10C0C8F8E5" pageId="57" pageNumber="332" type="reference_group">
<paragraph id="8B963699FF86FFA4FDD0FA10C0C8F8E5" blockId="57.[532,966,624,1917]" pageId="57" pageNumber="332">
<taxonomicName id="4C294D1AFF86FFA4FDD0FA10C0A9FA5C" baseAuthorityName="DAI &amp; ZHENG" baseAuthorityYear="2004" box="[532,656,1436,1457]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
, Scutellerina, Scutelleraria
<bibRefCitation id="EFB84B68FF86FFA4FC5CFA10C058FA3D" author="STAL C." journalOrPublisher="Akad. Forh." pageId="57" pageNumber="332" pagination="31 - 40" part="29" refId="ref89017" refString="STAL C. (1872): Genera Pentatomidarum Europae disposuit. Ofvers. Kongl. Svenska Vetensk. - Akad. Forh. 29: 31 - 40." title="Genera Pentatomidarum Europae disposuit. Ofvers. Kongl. Svenska Vetensk" type="journal article" year="1872">
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1872: 32
</bibRefCitation>
(new suprageneric taxon)
<taxonomicName id="4C294D1AFF86FFA4FDD0FA56C0A9FA02" baseAuthorityName="DAI &amp; ZHENG" baseAuthorityYear="2004" box="[532,656,1498,1519]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
, Scutellerinae, Scutelleraria:
<bibRefCitation id="EFB84B68FF86FFA4FDD0FA75C0E3F9E3" author="SCHOUTEDEN H." bookContentInfo="5 pls." box="[532,730,1529,1550]" journalOrPublisher="Genera Insectorum" pageId="57" pageNumber="332" pagination="1 - 98" part="24" refId="ref87964" refString="SCHOUTEDEN H. (1904): Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae. - Genera Insectorum 24: 1 - 98, 5 pls." title="Heteroptera. Fam. Pentatomidae. Subfam. Scutellerinae" type="book chapter" year="1904">
<smallCapsWord id="8D70A045FF86FFA4FDD0FA75C0BEF9E1" baselines="1545,1544" box="[532,647,1529,1550]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Schouteden" pageId="57" pageNumber="332">SCHOUTEDEN</smallCapsWord>
1904: 13
</bibRefCitation>
(description)
<taxonomicName id="4C294D1AFF86FFA4FDD0F995C0A9F9C3" baseAuthorityName="DAI &amp; ZHENG" baseAuthorityYear="2004" box="[532,656,1561,1582]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
, Scutellerinae, Scutellerini:
<bibRefCitation id="EFB84B68FF86FFA4FDD0F9B4C0FEF9A0" author="KIRKALDY G. W." box="[532,711,1592,1613]" journalOrPublisher="Felix Dames, Berlin" pageId="57" pageNumber="332" pagination="1 - 392" refId="ref86002" refString="KIRKALDY G. W. (1909): Catalogue of the Hemiptera (Heteroptera) Vol. 1. Cimicidae. - Felix Dames, Berlin: xl, 1 - 392." title="Catalogue of the Hemiptera (Heteroptera) Vol. 1. Cimicidae" type="book chapter" year="1909">
<smallCapsWord id="8D70A045FF86FFA4FDD0F9B4C050F9A6" baselines="1608,1607" box="[532,617,1592,1613]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kirkaldy" pageId="57" pageNumber="332">KIRKALDY</smallCapsWord>
1909: 289
</bibRefCitation>
(catalogue)
<taxonomicName id="4C294D1AFF86FFA4FDD0F9DBC0A9F981" baseAuthorityName="DAI &amp; ZHENG" baseAuthorityYear="2004" box="[532,656,1623,1644]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
, Scutellerinae, Scutellerini:
<bibRefCitation id="EFB84B68FF86FFA4FC6DF9DBC0A6F966" author="LESTON D." journalOrPublisher="Publ. Cult. Ca. Diamante Angola" pageId="57" pageNumber="332" pagination="9 - 26" part="16" refId="ref86505" refString="LESTON D. (1952 d): Notes on the Ethiopian Pentatomoidea (Hemiptera): VIII. Scutelleridae LEACH of Angola, with remarks upon the male genitalia and classification of the subfamily. - Publ. Cult. Ca. Diamante Angola 16: 9 - 26." title="Notes on the Ethiopian Pentatomoidea (Hemiptera): VIII. Scutelleridae LEACH of Angola, with remarks upon the male genitalia and classification of the subfamily" type="journal article" year="1952">
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1952d: 13
</bibRefCitation>
(new suprageneric classification)
<taxonomicName id="4C294D1AFF86FFA4FDD0F91AC0B0F946" authorityName="Leach" authorityYear="1815" box="[532,649,1686,1707]" class="Insecta" family="Scutelleridae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Scutelleridae</taxonomicName>
, Scutellerinae:
<bibRefCitation id="EFB84B68FF86FFA4FCDAF91AC015F927" author="MCDONALD F. J. D." journalOrPublisher="Quaest. Entomol." pageId="57" pageNumber="332" pagination="7 - 150" part="2" refId="ref87143" refString="MCDONALD F. J. D. (1966): The genitalia of North American Pentatomoidea (Hemiptera: Heteroptera). - Quaest. Entomol. 2: 7 - 150." title="The genitalia of North American Pentatomoidea (Hemiptera: Heteroptera)" type="journal article" year="1966">
<smallCapsWord id="8D70A045FF86FFA4FCDAF91AC107F944" baselines="1702,1701" box="[798,830,1686,1707]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Mc" pageId="57" pageNumber="332">
<smallCapsWord id="8D70A045FF86FFA4FCF9F91AC1B1F944" baselines="1702,1701" box="[829,904,1686,1707]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Donald" pageId="57" pageNumber="332">MCDONALD</smallCapsWord>
</smallCapsWord>
1966: 67
</bibRefCitation>
, 68 (new suprageneric classification)
<taxonomicName id="4C294D1AFF86FFA4FDD0F958C0B0F904" authorityName="Leach" authorityYear="1815" box="[532,649,1748,1769]" class="Insecta" family="Scutelleridae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="family">Scutelleridae</taxonomicName>
, Scutellerinae:
<bibRefCitation id="EFB84B68FF86FFA4FCD5F958C0B0F8E5" author="MCDONALD F. J. D. &amp; G. CASSIS" journalOrPublisher="Aust. J. Zool." pageId="57" pageNumber="332" pagination="537 - 572" part="32" refId="ref87212" refString="MCDONALD F. J. D. &amp; G. CASSIS (1984): Revision of the Australian Scutelleridae LEACH (Hemiptera). - Aust. J. Zool. 32: 537 - 572." title="Revision of the Australian Scutelleridae LEACH (Hemiptera)" type="journal article" year="1984">
<smallCapsWord id="8D70A045FF86FFA4FCD5F958C108F90A" baselines="1764,1763" box="[785,817,1748,1769]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Mc" pageId="57" pageNumber="332">
<smallCapsWord id="8D70A045FF86FFA4FCF4F958C142F90A" baselines="1764,1763" box="[816,891,1748,1769]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Donald" pageId="57" pageNumber="332">MCDONALD</smallCapsWord>
</smallCapsWord>
&amp;
<smallCapsWord id="8D70A045FF86FFA4FC5CF958C015F8EB" baselines="1764,1763,1764,1794" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Cassis" pageId="57" pageNumber="332">CASSIS</smallCapsWord>
1984: 550
</bibRefCitation>
(Australia)
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF86FFA4FDFEF8AEC75EFB19" pageId="57" pageNumber="332" type="diagnosis">
<paragraph id="8B963699FF86FFA4FDFEF8AEC75EFB19" blockId="57.[532,966,624,1917]" lastBlockId="57.[1000,1435,110,1917]" pageId="57" pageNumber="332">
Diagnosis: The
<taxonomicName id="4C294D1AFF86FFA4FD1CF8AEC163F8D4" box="[728,858,1826,1849]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="384" phylum="Arthropoda" rank="subFamily" subFamily="Scutellerinae">Scutellerinae</taxonomicName>
are recog-nised by the following combination charac-ters: small (e.g.,
<figureCitation id="13122A1CFF86FFA4FD7FF8EAC129F890" box="[699,784,1894,1917]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="57" pageNumber="332">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FCD8F8EAC10FF890" box="[796,822,1894,1917]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="57" pageNumber="332">37</figureCitation>
) to very large body (e.g.,
<figureCitation id="13122A1CFF86FFA4FB92FFE2C6A0FF68" box="[1110,1177,110,133]" captionStart="Fig" captionStartId="59.[60,96,1451,1470]" captionTargetBox="[63,967,110,1427]" captionTargetId="figure-485@59.[62,968,105,1435]" captionTargetPageId="59" captionText="Fig. 19: Habitus of Australian Calliphara species (a) Calliphara dimidiata cruenta, Oi (b) Calliphara imperialis, Oi (c) Calliphara nobilis, Oi (d) Calliphara regalis, Oi. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714011" httpUri="https://zenodo.org/record/13714011/files/figure.png" pageId="57" pageNumber="332">Figs 19</figureCitation>
a-d, 28a); ovoid (e.g.,
<figureCitation id="13122A1CFF86FFA4FAB0FFE2C636FF4B" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="57" pageNumber="332">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FBDBFF03C600FF4B" box="[1055,1081,143,166]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="57" pageNumber="332">37</figureCitation>
) to elongate-ovoid (
<figureCitation id="13122A1CFF86FFA4FAD7FF03C7B4FF4B" box="[1299,1421,143,166]" captionStart="Fig" captionStartId="74.[1089,1125,1601,1620]" captionTargetBox="[237,1506,114,1579]" captionTargetId="figure-155@74.[205,1509,105,1585]" captionTargetPageId="74" captionText="Fig. 28: Habitus of Australian Scutellerinae (a) Cantao parentum, Oi (b) Scutiphora pedicellata, ♀ (c) Lampromicra aerea, Oi (d) Lampromicra regia, Oi (e) Lampromicra senator, Oi. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13714035" httpUri="https://zenodo.org/record/13714035/files/figure.png" pageId="57" pageNumber="332">Figs 28a, b</figureCitation>
), sometimes strongly tapered towards terminalia; body weakly (e.g.,
<figureCitation id="13122A1CFF86FFA4FB25FF5EC708FF04" box="[1249,1329,210,233]" captionStart="Fig" captionStartId="74.[1089,1125,1601,1620]" captionTargetBox="[237,1506,114,1579]" captionTargetId="figure-155@74.[205,1509,105,1585]" captionTargetPageId="74" captionText="Fig. 28: Habitus of Australian Scutellerinae (a) Cantao parentum, Oi (b) Scutiphora pedicellata, ♀ (c) Lampromicra aerea, Oi (d) Lampromicra regia, Oi (e) Lampromicra senator, Oi. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13714035" httpUri="https://zenodo.org/record/13714035/files/figure.png" pageId="57" pageNumber="332">Fig. 28a</figureCitation>
) to heavily punctate (
<figureCitation id="13122A1CFF86FFA4FBA9FF7FC6FFFEE7" box="[1133,1222,243,266]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="57" pageNumber="332">Figs 24a</figureCitation>
, c-f); body brilliant, aposematically coloured, often with iridescent blue, green, orange colouration, often with contrasting markings (e.g.,
<figureCitation id="13122A1CFF86FFA4FAEAFEDBC74AFE83" box="[1326,1395,343,366]" captionStart="Fig" captionStartId="59.[60,96,1451,1470]" captionTargetBox="[63,967,110,1427]" captionTargetId="figure-485@59.[62,968,105,1435]" captionTargetPageId="59" captionText="Fig. 19: Habitus of Australian Calliphara species (a) Calliphara dimidiata cruenta, Oi (b) Calliphara imperialis, Oi (c) Calliphara nobilis, Oi (d) Calliphara regalis, Oi. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714011" httpUri="https://zenodo.org/record/13714011/files/figure.png" pageId="57" pageNumber="332">Figs 19</figureCitation>
a-d, 24a, c-f, 28a-e); head subtriangular to suboval (e.g.,
<figureCitation id="13122A1CFF86FFA4FB80FE16C6ACFE5C" box="[1092,1173,410,433]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="57" pageNumber="332">Fig. 31a</figureCitation>
); thoracic sterna flat (e.g.,
<figureCitation id="13122A1CFF86FFA4FC2CFE37C67DFE3F" box="[1000,1092,443,466]" captionStart="Fig" captionStartId="60.[1190,1226,107,126]" captionTargetBox="[151,1174,106,1322]" captionTargetId="figure-274@60.[151,1176,105,1323]" captionTargetPageId="60" captionText="Fig. 20: Scanning electron micrographs of key characters of Calliphara imperialis (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714015" httpUri="https://zenodo.org/record/13714015/files/figure.png" pageId="57" pageNumber="332">Figs 20c</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB9CFE37C644FE3F" box="[1112,1149,443,466]" captionStart="Fig" captionStartId="76.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1432]" captionTargetId="figure-280@76.[151,1277,105,1432]" captionTargetPageId="76" captionText="Fig. 29: Scanning electron micrographs of key characters of Cantao parentum (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; P = pronotum; P(al) = anterolateral margin of pronotum; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714043" httpUri="https://zenodo.org/record/13714043/files/figure.png" pageId="57" pageNumber="332">29c</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB55FE37C68EFE3F" box="[1169,1207,443,466]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="57" pageNumber="332">31c</figureCitation>
); anterior margin of proepisternum weakly explanate (e.g.,
<figureCitation id="13122A1CFF86FFA4FAB0FE51C637FDF8" captionStart="Fig" captionStartId="60.[1190,1226,107,126]" captionTargetBox="[151,1174,106,1322]" captionTargetId="figure-274@60.[151,1176,105,1323]" captionTargetPageId="60" captionText="Fig. 20: Scanning electron micrographs of key characters of Calliphara imperialis (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714015" httpUri="https://zenodo.org/record/13714015/files/figure.png" pageId="57" pageNumber="332">Figs 20c</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FBD8FE72C67BFDF8" box="[1052,1090,510,533]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="57" pageNumber="332">31c</figureCitation>
); efferent system of metathoracic glands well-developed (e.g.,
<figureCitation id="13122A1CFF86FFA4FACFFDACC758FDDA" box="[1291,1377,544,567]" captionStart="Fig" captionStartId="60.[1190,1226,107,126]" captionTargetBox="[151,1174,106,1322]" captionTargetId="figure-274@60.[151,1176,105,1323]" captionTargetPageId="60" captionText="Fig. 20: Scanning electron micrographs of key characters of Calliphara imperialis (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714015" httpUri="https://zenodo.org/record/13714015/files/figure.png" pageId="57" pageNumber="332">Figs 20d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FAABFDACC7AFFDDA" box="[1391,1430,544,567]" captionStart="Fig" captionStartId="71.[60,96,1556,1575]" captionTargetBox="[60,1436,106,1538]" captionTargetId="figure-240@71.[60,1436,105,1538]" captionTargetPageId="71" captionText="Fig. 25: Scanning electron micrographs of key characters of Calliscyta stalii (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; P = pronotum; P(al) = anterolateral margin of pronotum; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger; EES = external efferent system." figureDoi="http://doi.org/10.5281/zenodo.13714027" httpUri="https://zenodo.org/record/13714027/files/figure.png" pageId="57" pageNumber="332">25d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FC2CFDCDC629FDB5" box="[1000,1040,577,600]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="57" pageNumber="332">31d</figureCitation>
); peritreme mostly subreniform (e.g.,
<figureCitation id="13122A1CFF86FFA4FC2CFDEEC605FD94" box="[1000,1084,610,633]" captionStart="Fig" captionStartId="60.[1190,1226,107,126]" captionTargetBox="[151,1174,106,1322]" captionTargetId="figure-274@60.[151,1176,105,1323]" captionTargetPageId="60" captionText="Fig. 20: Scanning electron micrographs of key characters of Calliphara imperialis (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714015" httpUri="https://zenodo.org/record/13714015/files/figure.png" pageId="57" pageNumber="332">Figs 20d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB82FDEEC655FD94" box="[1094,1132,610,633]" captionStart="Fig" captionStartId="97.[126,162,107,126]" captionTargetBox="[317,1436,106,1423]" captionTargetId="figure-236@97.[317,1437,106,1424]" captionTargetPageId="97" captionText="Fig. 39: Scanning electron micrographs of key characters of Lampromicra senator (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714071" httpUri="https://zenodo.org/record/13714071/files/figure.png" pageId="57" pageNumber="332">39d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FBB3FDEEC6A6FD94" box="[1143,1183,610,633]" captionStart="Fig" captionStartId="106.[1294,1330,107,126]" captionTargetBox="[151,1278,106,1431]" captionTargetId="figure-283@106.[151,1278,105,1431]" captionTargetPageId="106" captionText="Fig. 42: Scanning electron micrographs of key characters of Scutiphora pedicelllata (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Ea = evaporative areas; GO = genital opening; Pe = peritreme; P(r) = right paramere. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714079" httpUri="https://zenodo.org/record/13714079/files/figure.png" pageId="57" pageNumber="332">42d</figureCitation>
), sometimes large, subtriangular (e.g.,
<figureCitation id="13122A1CFF86FFA4FBB9FD08C6EEFD76" box="[1149,1239,644,667]" captionStart="Fig" captionStartId="71.[60,96,1556,1575]" captionTargetBox="[60,1436,106,1538]" captionTargetId="figure-240@71.[60,1436,105,1538]" captionTargetPageId="71" captionText="Fig. 25: Scanning electron micrographs of key characters of Calliscyta stalii (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; P = pronotum; P(al) = anterolateral margin of pronotum; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger; EES = external efferent system." figureDoi="http://doi.org/10.5281/zenodo.13714027" httpUri="https://zenodo.org/record/13714027/files/figure.png" pageId="57" pageNumber="332">Figs 25d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB2CFD08C729FD76" box="[1256,1296,644,667]" captionStart="Fig" captionStartId="76.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1432]" captionTargetId="figure-280@76.[151,1277,105,1432]" captionTargetPageId="76" captionText="Fig. 29: Scanning electron micrographs of key characters of Cantao parentum (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; P = pronotum; P(al) = anterolateral margin of pronotum; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714043" httpUri="https://zenodo.org/record/13714043/files/figure.png" pageId="57" pageNumber="332">29d</figureCitation>
), or obovate (e.g.,
<figureCitation id="13122A1CFF86FFA4FBE4FD29C64DFD51" box="[1056,1140,677,700]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="57" pageNumber="332">Figs 31d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FBBAFD29C69CFD51" box="[1150,1189,677,700]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="57" pageNumber="332">38c</figureCitation>
); males without abdominal androconial or setose sternal glands; pregenital abdomen in both sexes without stridulatory vittae; ventral surface of pygophore caudally (e.g.,
<figureCitation id="13122A1CFF86FFA4FB12FCA7C727FCAF" box="[1238,1310,811,834]" captionStart="Fig" captionStartId="83.[60,96,1816,1835]" captionTargetBox="[72,958,106,1790]" captionTargetId="figure-443@83.[67,962,105,1800]" captionTargetPageId="83" captionText="Fig. 32: Colouration of Abdominal Venter of Choerocoris species (a-b) Choerocoris grossi nov.sp. (a) ventral view (b) lateral view (c-d) Choerocoris lattini nov.sp. (c) ventral view (d) lateral view (e-f) Choerocoris paganus (e) ventral view (f) lateral view (g-h) Choerocoris variegatus (g) ventral view (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13714053" httpUri="https://zenodo.org/record/13714053/files/figure.png" pageId="57" pageNumber="332">Figs 32</figureCitation>
a-h) or ventrally oriented (e.g.,
<figureCitation id="13122A1CFF86FFA4FB07FCC0C720FC8E" box="[1219,1305,844,867]" captionStart="Fig" captionStartId="76.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1432]" captionTargetId="figure-280@76.[151,1277,105,1432]" captionTargetPageId="76" captionText="Fig. 29: Scanning electron micrographs of key characters of Cantao parentum (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: Ea = evaporative area; Go = genital opening; P = pronotum; P(al) = anterolateral margin of pronotum; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger; EES = external efferent system. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714043" httpUri="https://zenodo.org/record/13714043/files/figure.png" pageId="57" pageNumber="332">Figs 29e</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FAEDFCC0C768FC8E" box="[1321,1361,844,867]" captionStart="Fig" captionStartId="77.[60,96,107,126]" captionTargetBox="[60,1436,107,1916]" captionTargetId="figure-15@77.[189,1437,105,1921]" captionTargetPageId="77" captionText="Fig. 30: Male genitalia of Cantao parentum (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII = second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714047" httpUri="https://zenodo.org/record/13714047/files/figure.png" pageId="57" pageNumber="332">30e</figureCitation>
); parameres hook-shaped (e.g.,
<figureCitation id="13122A1CFF86FFA4FB3EFCE1C76DFC69" box="[1274,1364,877,900]" captionStart="Fig" captionStartId="63.[60,96,1708,1727]" captionTargetBox="[60,1412,108,1807]" captionTargetId="figure-13@63.[89,1414,105,1812]" captionTargetPageId="63" captionText="Fig. 22: Male genitalia of Calliphara regalis (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; Vp = distal process of vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714019" httpUri="https://zenodo.org/record/13714019/files/figure.png" pageId="57" pageNumber="332">Figs 22b</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FAA2FCE1C7B7FC69" box="[1382,1422,877,900]" captionStart="Fig" captionStartId="98.[1114,1150,107,126]" captionTargetBox="[157,1523,106,1912]" captionTargetId="figure-9@98.[151,1461,105,1921]" captionTargetPageId="98" captionText="Fig. 40: Male genitalia of Lampromicra aerea (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; Pt(P) = process of phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patch; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714075" httpUri="https://zenodo.org/record/13714075/files/figure.png" pageId="57" pageNumber="332">40b</figureCitation>
); ejaculatory apparatus usually well-developed, with convoluted ventral conducting canal and oval to subelliptoid ejaculatory reservoir (e.g.,
<figureCitation id="13122A1CFF86FFA4FB45FC7FC6C8FBE7" box="[1153,1265,1011,1034]" captionStart="Fig" captionStartId="77.[60,96,107,126]" captionTargetBox="[60,1436,107,1916]" captionTargetId="figure-15@77.[189,1437,105,1921]" captionTargetPageId="77" captionText="Fig. 30: Male genitalia of Cantao parentum (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII = second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714047" httpUri="https://zenodo.org/record/13714047/files/figure.png" pageId="57" pageNumber="332">Figs 30c, d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB3AFC7FC707FBE7" box="[1278,1342,1011,1034]" captionStart="Fig" captionStartId="89.[585,621,1689,1708]" captionTargetBox="[59,1436,113,1921]" captionTargetId="figure-15@89.[58,1437,107,1920]" captionTargetPageId="89" captionText="Fig. 36: Male genitalia of Choerocoris variegatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; S = stem of paramere; Sg = secondary gonopore; V = vesica; Vp = process of vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714063" httpUri="https://zenodo.org/record/13714063/files/figure.png" pageId="57" pageNumber="332">36c, d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FA88FC7FC7B4FBE7" box="[1356,1421,1011,1034]" captionStart="Fig" captionStartId="98.[1114,1150,107,126]" captionTargetBox="[157,1523,106,1912]" captionTargetId="figure-9@98.[151,1461,105,1921]" captionTargetPageId="98" captionText="Fig. 40: Male genitalia of Lampromicra aerea (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; Pt(P) = process of phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patch; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714075" httpUri="https://zenodo.org/record/13714075/files/figure.png" pageId="57" pageNumber="332">40c, d</figureCitation>
); CAI usually absent, rarely present (e.g.,
<figureCitation id="13122A1CFF86FFA4FAB0FB98C613FBA0" captionStart="Fig" captionStartId="63.[60,96,1708,1727]" captionTargetBox="[60,1412,108,1807]" captionTargetId="figure-13@63.[89,1414,105,1812]" captionTargetPageId="63" captionText="Fig. 22: Male genitalia of Calliphara regalis (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; Vp = distal process of vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714019" httpUri="https://zenodo.org/record/13714019/files/figure.png" pageId="57" pageNumber="332">Figs 22c, d</figureCitation>
); CAII usually bifurcate, with lobal sclerites (e.g.,
<figureCitation id="13122A1CFF86FFA4FBB8FBDBC6D4FB83" box="[1148,1261,1111,1134]" captionStart="Fig" captionStartId="63.[60,96,1708,1727]" captionTargetBox="[60,1412,108,1807]" captionTargetId="figure-13@63.[89,1414,105,1812]" captionTargetPageId="63" captionText="Fig. 22: Male genitalia of Calliphara regalis (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; Vp = distal process of vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714019" httpUri="https://zenodo.org/record/13714019/files/figure.png" pageId="57" pageNumber="332">Figs 22c, d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB3FFBDBC704FB83" box="[1275,1341,1111,1134]" captionStart="Fig" captionStartId="89.[585,621,1689,1708]" captionTargetBox="[59,1436,113,1921]" captionTargetId="figure-15@89.[58,1437,107,1920]" captionTargetPageId="89" captionText="Fig. 36: Male genitalia of Choerocoris variegatus (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; S = stem of paramere; Sg = secondary gonopore; V = vesica; Vp = process of vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714063" httpUri="https://zenodo.org/record/13714063/files/figure.png" pageId="57" pageNumber="332">36c, d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FA8FFBDBC7B4FB83" box="[1355,1421,1111,1134]" captionStart="Fig" captionStartId="98.[1114,1150,107,126]" captionTargetBox="[157,1523,106,1912]" captionTargetId="figure-9@98.[151,1461,105,1921]" captionTargetPageId="98" captionText="Fig. 40: Male genitalia of Lampromicra aerea (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; Pt(P) = process of phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patch; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714075" httpUri="https://zenodo.org/record/13714075/files/figure.png" pageId="57" pageNumber="332">40c, d</figureCitation>
), CAIII undivided, often U-shaped (e.g.,
<figureCitation id="13122A1CFF86FFA4FAB0FBF5C636FB5C" captionStart="Fig" captionStartId="72.[151,187,107,126]" captionTargetBox="[151,1527,107,1920]" captionTargetId="figure-14@72.[247,1527,86,1921]" captionTargetPageId="72" captionText="Fig. 26: Male genitalia of Calliscyta stalii (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714029" httpUri="https://zenodo.org/record/13714029/files/figure.png" pageId="57" pageNumber="332">Figs 26d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FBDEFB16C679FB5C" box="[1050,1088,1178,1201]" captionStart="Fig" captionStartId="77.[60,96,107,126]" captionTargetBox="[60,1436,107,1916]" captionTargetId="figure-15@77.[189,1437,105,1921]" captionTargetPageId="77" captionText="Fig. 30: Male genitalia of Cantao parentum (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII = second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714047" httpUri="https://zenodo.org/record/13714047/files/figure.png" pageId="57" pageNumber="332">30d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FB8FFB16C648FB5C" box="[1099,1137,1178,1201]" captionStart="Fig" captionStartId="107.[60,96,107,126]" captionTargetBox="[60,1436,106,1917]" captionTargetId="figure-15@107.[186,1438,105,1921]" captionTargetPageId="107" captionText="Fig. 43: Male genitalia of Scutiphora pedicellata (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patch; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714083" httpUri="https://zenodo.org/record/13714083/files/figure.png" pageId="57" pageNumber="332">43d</figureCitation>
) or divided (e.g.,
<figureCitation id="13122A1CFF86FFA4FAE6FB16C7B4FB5C" box="[1314,1421,1178,1201]" captionStart="Fig" captionStartId="63.[60,96,1708,1727]" captionTargetBox="[60,1412,108,1807]" captionTargetId="figure-13@63.[89,1414,105,1812]" captionTargetPageId="63" captionText="Fig. 22: Male genitalia of Calliphara regalis (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; Vp = distal process of vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714019" httpUri="https://zenodo.org/record/13714019/files/figure.png" pageId="57" pageNumber="332">Figs 22c, d</figureCitation>
); spermathecal fecundation canal short or long; and, spermathecal reservoir oval.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF86FFA6FBCAFA81C67CFE58" lastPageId="59" lastPageNumber="334" pageId="57" pageNumber="332" type="discussion">
<paragraph id="8B963699FF86FFA6FBCAFA81C102F932" blockId="57.[1000,1435,110,1917]" lastBlockId="59.[532,965,1451,1917]" lastPageId="59" lastPageNumber="334" pageId="57" pageNumber="332">
Remarks: There is no unique character state defining the
<taxonomicName id="4C294D1AFF86FFA4FB5EFAA3C724FAAB" box="[1178,1309,1327,1350]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="384" phylum="Arthropoda" rank="subFamily" subFamily="Scutellerinae">Scutellerinae</taxonomicName>
, and most of the above-mentioned features are also true for the family. They are the most brilliantly coloured of all the scutellerids, but numerous
<taxonomicName id="4C294D1AFF86FFA4FC2CFA38C652FA26" box="[1000,1131,1460,1483]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="384" phylum="Arthropoda" rank="tribe" tribe="Pachycorinae">Pachycorinae</taxonomicName>
also exhibit colouration consistent with models of warning colouration. The male and female genitalia are also similar to taxa in other scutellerid subfamilies, such as the
<taxonomicName id="4C294D1AFF86FFA4FB99F9B6C6F0F9BC" box="[1117,1225,1594,1617]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
and Sphaerocorinae, although the development of the ventral conducting canal of the ejaculatory apparatus is putatively greatest in the scutellerines, such as in
<taxonomicName id="4C294D1AFF86FFA4FB9AF933C73DF93B" box="[1118,1284,1727,1750]" class="Insecta" family="Scutelleridae" genus="Cantao" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="species" species="parentum">
<emphasis id="B95DEA8BFF86FFA4FB9AF933C73DF93B" box="[1118,1284,1727,1750]" italics="true" pageId="57" pageNumber="332">Cantao parentum</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1CFF86FFA4FAD2F933C7B4F93B" box="[1302,1421,1727,1750]" captionStart="Fig" captionStartId="77.[60,96,107,126]" captionTargetBox="[60,1436,107,1916]" captionTargetId="figure-15@77.[189,1437,105,1921]" captionTargetPageId="77" captionText="Fig. 30: Male genitalia of Cantao parentum (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII = second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714047" httpUri="https://zenodo.org/record/13714047/files/figure.png" pageId="57" pageNumber="332">Figs 30c, d</figureCitation>
). Many species have a U-shaped, and basally fused CAIII (e.g.,
<figureCitation id="13122A1CFF86FFA4FB61F88EC6C2F8F4" box="[1189,1275,1794,1817]" captionStart="Fig" captionStartId="72.[151,187,107,126]" captionTargetBox="[151,1527,107,1920]" captionTargetId="figure-14@72.[247,1527,86,1921]" captionTargetPageId="72" captionText="Fig. 26: Male genitalia of Calliscyta stalii (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714029" httpUri="https://zenodo.org/record/13714029/files/figure.png" pageId="57" pageNumber="332">Figs 26d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FACCF88EC716F8F4" box="[1288,1327,1794,1817]" captionStart="Fig" captionStartId="77.[60,96,107,126]" captionTargetBox="[60,1436,107,1916]" captionTargetId="figure-15@77.[189,1437,105,1921]" captionTargetPageId="77" captionText="Fig. 30: Male genitalia of Cantao parentum (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAII = second conjunctival appendage; CAIII = third conjunctival appendage; DCC = dorsal conducting canal; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; VCC = ventral conducting canal; V = vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714047" httpUri="https://zenodo.org/record/13714047/files/figure.png" pageId="57" pageNumber="332">30d</figureCitation>
,
<figureCitation id="13122A1CFF86FFA4FAF8F88EC75DF8F4" box="[1340,1380,1794,1817]" captionStart="Fig" captionStartId="107.[60,96,107,126]" captionTargetBox="[60,1436,106,1917]" captionTargetId="figure-15@107.[186,1438,105,1921]" captionTargetPageId="107" captionText="Fig. 43: Male genitalia of Scutiphora pedicellata (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) aedeagus, ventral view. Abbreviations: C = crown of right paramere; CAII(L) = second conjunctival appendage, lateral branch; CAII(M) = second conjunctival appendage, medial branch; CAIII = third conjunctival appendage; DS(D) = ductus seminis dorsalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patch; V = vesica; VCC = ventral conducting canal of ejaculatory apparatus; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714083" httpUri="https://zenodo.org/record/13714083/files/figure.png" pageId="57" pageNumber="332">43d</figureCitation>
), but there are significant variations, particularly in
<taxonomicName id="4C294D1AFF86FFA4FBC7F8C9C65BF8B1" authorityName="GERMAR" authorityYear="1839" box="[1027,1122,1861,1884]" class="Insecta" family="Scutelleridae" genus="Calliphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFF86FFA4FBC7F8C9C65BF8B1" box="[1027,1122,1861,1884]" italics="true" pageId="57" pageNumber="332">Calliphara</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1CFF86FFA4FBABF8C9C6FDF8B1" box="[1135,1220,1861,1884]" captionStart="Fig" captionStartId="63.[60,96,1708,1727]" captionTargetBox="[60,1412,108,1807]" captionTargetId="figure-13@63.[89,1414,105,1812]" captionTargetPageId="63" captionText="Fig. 22: Male genitalia of Calliphara regalis (a) pygophore, dorsal view (b) paramere, lateral view (c) aedeagus, lateral view (d) apex of aedeagus, ventral view. Abbreviations: C = crown of paramere; CAI(V) = ventral branch of first conjunctival appendage; CAI(D) = dorsal branch of first conjunctival appendage; CAII(L) = lateral branch of second conjunctival appendage; CAII(M) = medial branch of second conjunctival appendage; CAIII = third conjunctival appendage; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; SP = setose patches of male genital opening; VCC = ventral conducting canal; Vp = distal process of vesica; VM = ventral margin of genital opening." figureDoi="http://doi.org/10.5281/zenodo.13714019" httpUri="https://zenodo.org/record/13714019/files/figure.png" pageId="57" pageNumber="332">Fig. 22d</figureCitation>
). Scutellerines can be separated from the
<taxonomicName id="4C294D1AFF86FFA4FB65F8EAC71FF890" box="[1185,1318,1894,1917]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="384" phylum="Arthropoda" rank="tribe" tribe="Pachycorinae">Pachycorinae</taxonomicName>
, and Odon-totarsinae +
<taxonomicName id="4C294D1AFF84FFA6FD55FA27C12BFA2F" box="[657,786,1451,1474]" pageId="59" pageNumber="380" rank="subFamily" subFamily="Tectocorinae">Tectocorinae</taxonomicName>
, in lacking stridulatory vittae and androconial/setose sternal glands respectively. They can be separated from the
<taxonomicName id="4C294D1AFF84FFA6FDB8F99AC0D2F9C0" box="[636,747,1558,1581]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
,
<taxonomicName id="4C294D1AFF84FFA6FD3FF99AC1AAF9C0" box="[763,915,1558,1581]" pageId="59" pageNumber="320" rank="subFamily" subFamily="Odontotarsinae">Odontotarsinae</taxonomicName>
and Sphaerocorinae superficially by their aposematic colouration (cf. dull colouration), although even in the latter subfamily some taxa exhibit moderately striking colouration, as in
<taxonomicName id="4C294D1AFF84FFA6FD8CF944C102F932" authority="BURMEISTER." authorityName="BURMEISTER." box="[584,827,1736,1759]" class="Insecta" family="Scutelleridae" genus="Sphaerocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFF84FFA6FD8CF944C083F932" box="[584,698,1736,1759]" italics="true" pageId="59" pageNumber="334">Sphaerocoris</emphasis>
<smallCapsWord id="8D70A045FF84FFA6FD04F944C10CF933" baselines="1753,1754" box="[704,821,1736,1759]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Burmeister" pageId="59" pageNumber="334">BURMEISTER</smallCapsWord>
.
</taxonomicName>
</paragraph>
<caption id="DF566611FF85FFA7FD3FFA56C187F94A" ID-DOI="http://doi.org/10.5281/zenodo.13714007" ID-Zenodo-Dep="13714007" httpUri="https://zenodo.org/record/13714007/files/figure.png" pageId="58" pageNumber="333" startId="58.[763,799,1498,1517]" targetBox="[171,1456,106,1920]" targetPageId="58" targetType="figure">
<paragraph id="8B963699FF85FFA7FD3FFA56C187F94A" blockId="58.[763,1456,1498,1703]" pageId="58" pageNumber="333">
<emphasis id="B95DEA8BFF85FFA7FD3FFA56C17DFA00" bold="true" box="[763,836,1498,1517]" pageId="58" pageNumber="333">Fig. 18</emphasis>
: Male genitalia of
<taxonomicName id="4C294D1AFF85FFA7FBC7FA56C6C9FA00" box="[1027,1264,1498,1517]" class="Insecta" family="Dolichopodidae" genus="Agonosoma" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="333" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="B95DEA8BFF85FFA7FBC7FA56C6C9FA00" box="[1027,1264,1498,1517]" italics="true" pageId="58" pageNumber="333">Agonosoma trilineatum</emphasis>
</taxonomicName>
(
<emphasis id="B95DEA8BFF85FFA7FB3AFA56C732FA00" bold="true" box="[1278,1291,1498,1517]" pageId="58" pageNumber="333">a</emphasis>
) pygophore, dorsal view (
<emphasis id="B95DEA8BFF85FFA7FCBEFA78C1B1F9EA" bold="true" box="[890,904,1524,1543]" pageId="58" pageNumber="333">b</emphasis>
) paramere, lateral view (
<emphasis id="B95DEA8BFF85FFA7FB41FA78C6A9F9EA" bold="true" box="[1157,1168,1524,1543]" pageId="58" pageNumber="333">c</emphasis>
) aedeagus, lateral view (
<emphasis id="B95DEA8BFF85FFA7FA4FFA78C7A0F9EA" bold="true" box="[1419,1433,1524,1543]" pageId="58" pageNumber="333">d</emphasis>
) apex of aedeagus, ventral view. Abbreviations: C = crown of right paramere; DS(D) = ductus seminis distalis; DS(P) = ductus seminis proximalis; Er = ejaculatory reservoir; F = flange at base of crown of paramere; GO = genital opening; Pt = phallotheca; S = stem of paramere; Sg = secondary gonopore; V = vesica; VM = ventral margin of genital opening.
</paragraph>
</caption>
<caption id="DF566611FF84FFA6FFF8FA27C2D6F9C5" ID-DOI="http://doi.org/10.5281/zenodo.13714011" ID-Zenodo-Dep="13714011" httpUri="https://zenodo.org/record/13714011/files/figure.png" pageId="59" pageNumber="334" startId="59.[60,96,1451,1470]" targetBox="[63,967,110,1427]" targetPageId="59" targetType="figure">
<paragraph id="8B963699FF84FFA6FFF8FA27C2D6F9C5" blockId="59.[60,481,1451,1576]" pageId="59" pageNumber="334">
<emphasis id="B95DEA8BFF84FFA6FFF8FA27C2BCFA53" bold="true" box="[60,133,1451,1470]" pageId="59" pageNumber="334">Fig. 19</emphasis>
: Habitus of Australian
<taxonomicName id="4C294D1AFF84FFA6FEA9FA27C2B8FA35" pageId="59" pageNumber="334">
<emphasis id="B95DEA8BFF84FFA6FEA9FA27C3EBFA53" box="[365,466,1451,1470]" italics="true" pageId="59" pageNumber="334">Calliphara</emphasis>
species
</taxonomicName>
(
<emphasis id="B95DEA8BFF84FFA6FF4BFA49C2A5FA35" bold="true" box="[143,156,1477,1496]" pageId="59" pageNumber="334">a</emphasis>
)
<taxonomicName id="4C294D1AFF84FFA6FF6DFA49C3D8FA34" authority=", Oi" authorityName="Oi" box="[169,481,1477,1497]" class="Insecta" family="Scutelleridae" genus="Calliphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="334" phylum="Arthropoda" rank="subSpecies" species="dimidiata" subSpecies="cruenta">
<emphasis id="B95DEA8BFF84FFA6FF6DFA49C3FFFA35" box="[169,454,1477,1496]" italics="true" pageId="59" pageNumber="334">Calliphara dimidiata cruenta</emphasis>
, ♂
</taxonomicName>
(
<emphasis id="B95DEA8BFF84FFA6FF80FA6CC26BFA1E" bold="true" box="[68,82,1504,1523]" pageId="59" pageNumber="334">b</emphasis>
)
<taxonomicName id="4C294D1AFF84FFA6FF9BFA6CC37FFA19" authority=", Oi" authorityName="Oi" box="[95,326,1504,1524]" class="Insecta" family="Scutelleridae" genus="Calliphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="334" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B95DEA8BFF84FFA6FF9BFA6CC312FA1E" box="[95,299,1504,1523]" italics="true" pageId="59" pageNumber="334">Calliphara imperialis</emphasis>
, ♂
</taxonomicName>
(
<emphasis id="B95DEA8BFF84FFA6FE90FA6CC366FA1E" bold="true" box="[340,351,1504,1523]" pageId="59" pageNumber="334">c</emphasis>
)
<taxonomicName id="4C294D1AFF84FFA6FEA8FA6CC2A0F9E2" authority=", Oi" authorityName="Oi" class="Insecta" family="Scutelleridae" genus="Calliphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="334" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="B95DEA8BFF84FFA6FEA8FA6CC244F9E3" italics="true" pageId="59" pageNumber="334">Calliphara nobilis</emphasis>
, ♂
</taxonomicName>
(
<emphasis id="B95DEA8BFF84FFA6FF63FA77C28CF9E3" bold="true" box="[167,181,1531,1550]" pageId="59" pageNumber="334">d</emphasis>
)
<taxonomicName id="4C294D1AFF84FFA6FF07FA77C257F9C5" authority=", Oi. Scale" authorityName="Oi. Scale" class="Insecta" family="Scutelleridae" genus="Calliphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="334" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="B95DEA8BFF84FFA6FF07FA77C356F9E3" box="[195,367,1531,1550]" italics="true" pageId="59" pageNumber="334">Calliphara regalis</emphasis>
, ♂. Scale
</taxonomicName>
bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8B963699FF84FFA6FDFEF977C67CFE58" blockId="59.[532,965,1451,1917]" lastBlockId="59.[1000,1435,110,1917]" pageId="59" pageNumber="334">
<bibRefCitation id="EFB84B68FF84FFA6FDFEF977C106F8FF" author="SCHUH R. T. &amp; J. A. SLATER" box="[570,831,1787,1810]" journalOrPublisher="Cornell University Press, Ithaca" pageId="59" pageNumber="334" pagination="1 - 336" refId="ref88208" refString="SCHUH R. T. &amp; J. A. SLATER (1995): True Bugs of the World (Hemiptera: Heteroptera). Classification and Natural History. - Cornell University Press, Ithaca: xii + 1 - 336." title="True Bugs of the World (Hemiptera: Heteroptera). Classification and Natural History" type="book chapter" year="1995">
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&amp;
<smallCapsWord id="8D70A045FF84FFA6FD6EF977C0C8F8FC" baselines="1804,1805" box="[682,753,1787,1810]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Slater" pageId="59" pageNumber="334">SLATER</smallCapsWord>
(1995)
</bibRefCitation>
adopted
<smallCapsWord id="8D70A045FF84FFA6FC61F977C073F8DB" baselines="1804,1805,1804,1841,1840" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Leston'" pageId="59" pageNumber="334">LESTON</smallCapsWord>
s (1952d) definition of the
<taxonomicName id="4C294D1AFF84FFA6FC9AF893C00EF8B7" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="384" phylum="Arthropoda" rank="subFamily" subFamily="Scutellerinae">Scutellerinae</taxonomicName>
(including the
<taxonomicName id="4C294D1AFF84FFA6FD0BF8CFC105F8B7" box="[719,828,1859,1882]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="59" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
and Sphaerocorinae), but on the basis of the emarginate abdominal sterna (recurved anteriorly) and the presence of two intervannal veins in the hindwing. Neither of these characters is considered exclusive to this conception of the taxon. We have retained the scutellerines in the narrow sense, despite it being a convenience group, as it helps in distinguishing the colourful Australian scutellerid species, pending a more in-depth review of the issue.
</paragraph>
</subSubSection>
</treatment>
</document>