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<mods:title id="496DC684278A97A298885443355766FF">Revision of Indian wolf spiders: I. Genus Arctosa C. L. Koch, 1847 (Araneae Lycosidae, Tricassinae)</mods:title>
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<mods:affiliation id="318B287FED83426D6053C44ABBF67D63">Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India. caleb 87 woodgate @ gmail. com; https: // orcid. org / 0000 - 0002 - 9471 - 9467</mods:affiliation>
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<taxonomicName id="4C294D1CFFCBFF90FF21FF1F40AFDAFD" ID-CoL="GBFC" authority="Tikader &amp; Malhotra, 1980" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[151,665,151,178]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFCBFF90FF21FF1F40AFDAFD" bold="true" box="[151,665,151,178]" pageId="6" pageNumber="495">
<emphasis id="B95DEA8DFFCBFF90FF21FF1F4370DAFE" bold="true" box="[151,326,151,178]" italics="true" pageId="6" pageNumber="495">Arctosa indica</emphasis>
<bibRefCitation id="EFB84B6EFFCBFF90FEFBFF1F40AFDAFD" author="Tikader, B. K. &amp; Malhotra, M. S." box="[333,665,151,177]" pageId="6" pageNumber="495" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra, 1980</bibRefCitation>
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<figureCitation id="13122A1AFFCBFF90FF21FF3342CFDA9A" box="[151,249,187,214]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="6.[151,250,1965,1989]" captionStartId-1="7.[151,250,1966,1990]" captionTargetBox-0="[181,1404,546,1928]" captionTargetBox-1="[158,1430,785,1933]" captionTargetId-0="figure-146@6.[173,1414,540,1941]" captionTargetId-1="figure-291@7.[151,1437,777,1943]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 5. Arctosa indica Tikader &amp; Malhotra, 1980. A female holotype habitus, dorsal. B male allotype habitus, dorsal. CD labels from type tubes. Scale bars: AB, 2 mm." captionText-1="FIGURE 6. Arctosa indica Tikader &amp; Malhotra, 1980. AC male allotype, left pedipalp: A ventral, B ventral, enlarged; C retrolateral, enlarged. DE female holotype, genitalia: D epigynum, ventral, E vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4447147" figureDoi-1="http://doi.org/10.5281/zenodo.4447151" httpUri-0="https://zenodo.org/record/4447147/files/figure.png" httpUri-1="https://zenodo.org/record/4447151/files/figure.png" pageId="6" pageNumber="495">Figs 56</figureCitation>
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<treatmentCitation id="0A88108EFFCBFF90FF21FE8840B4DB54" author="Tikader, B. K. &amp; Malhotra, M. S." box="[151,642,256,280]" page="371" pageId="6" pageNumber="495" year="1980">
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<emphasis id="B95DEA8DFFCBFF90FF21FE884300DB54" box="[151,310,256,280]" italics="true" pageId="6" pageNumber="495">Arctosa indicus</emphasis>
<bibRefCitation id="EFB84B6EFFCBFF90FE8BFE8840B4DB54" author="Tikader, B. K. &amp; Malhotra, M. S." box="[317,642,256,280]" pageId="6" pageNumber="495" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra, 1980: 371</bibRefCitation>
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, figs 242246.
<treatmentCitation id="0A88108EFFCBFF90FC97FE884609DB5B" author="Tikader, B. K. &amp; Biswas, B." box="[801,1087,255,280]" page="58" pageId="6" pageNumber="495" year="1981">
<bibRefCitation id="EFB84B6EFFCBFF90FC97FE884609DB5B" author="Tikader, B. K. &amp; Biswas, B." box="[801,1087,255,280]" pageId="6" pageNumber="495" pagination="1 - 149" refId="ref6603" refString="Tikader, B. K. &amp; Biswas, B. (1981) Spider fauna of Calcutta and vicinity: Part-I. Records of the Zoological Survey of India, Occasional Paper, 30, 1 - 149." type="journal article" year="1981">Tikader &amp; Biswas 1981: 58</bibRefCitation>
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, plate VIII, figs 9495.
<treatmentCitation id="0A88108EFFCBFF90FA8EFE884349DB7A" author="Biswas, B. &amp; Biswas, K." page="438" pageId="6" pageNumber="495" year="1992">
<bibRefCitation id="EFB84B6EFFCBFF90FA8EFE884349DB7A" author="Biswas, B. &amp; Biswas, K." pageId="6" pageNumber="495" pagination="357 - 500" refId="ref5575" refString="Biswas, B. &amp; Biswas, K. (1992) Araneae: Spiders. State Fauna Series 3. Fauna of West Bengal, 3, 357 - 500." type="journal article" year="1992">Biswas &amp; Biswas 1992: 438</bibRefCitation>
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.
<treatmentCitation id="0A88108EFFCBFF90FE31FE964075DB7A" author="Yin, C. M. &amp; Peng, X. J. &amp; Xie, L. P. &amp; Bao, Y. H. &amp; Wang, J. F." box="[391,579,286,310]" page="84" pageId="6" pageNumber="495" year="1997">
<bibRefCitation id="EFB84B6EFFCBFF90FE31FE964075DB7A" author="Yin, C. M. &amp; Peng, X. J. &amp; Xie, L. P. &amp; Bao, Y. H. &amp; Wang, J. F." box="[391,579,286,310]" pageId="6" pageNumber="495" refId="ref6828" refString="Yin, C. M., Peng, X. J., Xie, L. P., Bao, Y. H. &amp; Wang, J. F. (1997) Lycosids in China (Arachnida: Araneae). Hunan Normal University Press, Changsha, Hunan, 317 pp." type="book" year="1997">
Yin
<emphasis id="B95DEA8DFFCBFF90FE07FE9743D2DB7A" box="[433,484,286,310]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
1997: 84
</bibRefCitation>
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, fig. 37af.
<treatmentCitation id="0A88108EFFCBFF90FD0AFE9641A3DB7A" author="Song, D. X. &amp; Zhu, M. S. &amp; Chen, J." box="[700,917,286,310]" page="319" pageId="6" pageNumber="495" year="1999">
<bibRefCitation id="EFB84B6EFFCBFF90FD0AFE9641A3DB7A" author="Song, D. X. &amp; Zhu, M. S. &amp; Chen, J." box="[700,917,286,310]" pageId="6" pageNumber="495" refId="ref6514" refString="Song, D. X., Zhu, M. S. &amp; Chen, J. (1999) The spiders of China. Hebei University of Science and Technology Publishing House, Shijiazhuang, 640 pp." type="book" year="1999">
Song
<emphasis id="B95DEA8DFFCBFF90FD43FE97411EDB7A" box="[757,808,286,310]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
1999: 319
</bibRefCitation>
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, figs 188P, 189A.
<treatmentCitation id="0A88108EFFCBFF90FBFAFE9646DCDB7A" author="Gajbe, U. A." box="[1100,1258,286,310]" page="31" pageId="6" pageNumber="495" year="2004">
<bibRefCitation id="EFB84B6EFFCBFF90FBFAFE9646DCDB7A" author="Gajbe, U. A." box="[1100,1258,286,310]" pageId="6" pageNumber="495" pagination="1 - 40" refId="ref5945" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40." type="journal article" year="2004">Gajbe 2004: 31</bibRefCitation>
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.
<treatmentCitation id="0A88108EFFCBFF90FB45FE9642D5DB19" author="Majumder, S. C." page="26" pageId="6" pageNumber="495" year="2005">
<bibRefCitation id="EFB84B6EFFCBFF90FB45FE9642D5DB19" author="Majumder, S. C." pageId="6" pageNumber="495" pagination="1 - 57" refId="ref6250" refString="Majumder, S. C. (2005) Studies on some spiders from eastern coastal region of India. Memoirs of the Zoological Survey of India, 20, 1 - 57." type="journal article" year="2005">Majumder 2005: 26</bibRefCitation>
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.
<treatmentCitation id="0A88108EFFCBFF90FF5BFEB543AEDB19" author="Gajbe, U. A." box="[237,408,317,341]" page="505" pageId="6" pageNumber="495" year="2007">
<bibRefCitation id="EFB84B6EFFCBFF90FF5BFEB543AEDB19" author="Gajbe, U. A." box="[237,408,317,341]" pageId="6" pageNumber="495" pagination="419 - 540" refId="ref5990" refString="Gajbe, U. A. (2007) Araneae: Arachnida. In: Fauna of Madhya Pradesh (including Chhattisgarh), State Fauna Series. Zoological Survey of India, Kolkata, 15, 419 - 540." type="journal article" year="2007">Gajbe 2007: 505</bibRefCitation>
</treatmentCitation>
, figs 252256.
<treatmentCitation id="0A88108EFFCBFF90FD82FEB54114DB19" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[564,802,317,341]" page="1200" pageId="6" pageNumber="495" year="2012">
<bibRefCitation id="EFB84B6EFFCBFF90FD82FEB54114DB19" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[564,802,317,341]" pageId="6" pageNumber="495" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFCBFF90FDC5FEB64090DB19" box="[627,678,317,341]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
2012: 1200
</bibRefCitation>
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.
<treatmentCitation id="0A88108EFFCBFF90FC9CFEB541C0DB19" author="Yin, C. M. &amp; Peng, X. J. &amp; Yan, H. M. &amp; Bao, Y. H. &amp; Xu, X. &amp; Tang, G. &amp; Zhou, Q. S. &amp; Liu, P." box="[810,1014,317,341]" page="796" pageId="6" pageNumber="495" year="2012">
<bibRefCitation id="EFB84B6EFFCBFF90FC9CFEB541C0DB19" author="Yin, C. M. &amp; Peng, X. J. &amp; Yan, H. M. &amp; Bao, Y. H. &amp; Xu, X. &amp; Tang, G. &amp; Zhou, Q. S. &amp; Liu, P." box="[810,1014,317,341]" pageId="6" pageNumber="495" refId="ref6886" refString="Yin, C. M., Peng, X. J., Yan, H. M., Bao, Y. H., Xu, X., Tang, G., Zhou, Q. S. &amp; Liu, P. (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">
Yin
<emphasis id="B95DEA8DFFCBFF90FCE3FEB641BFDB19" box="[853,905,317,341]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
2012: 796
</bibRefCitation>
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, fig. 397af.
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</paragraph>
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<taxonomicName id="4C294D1CFFCBFF90FF21FED44377DB38" ID-CoL="GBFC" authority="Tikader &amp; Malhotra, 1980: 371" authorityName="Tikader &amp; Malhotra" authorityPageNumber="371" authorityYear="1980" box="[151,321,348,372]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFCBFF90FF21FED44377DB38" box="[151,321,348,372]" italics="true" pageId="6" pageNumber="495">Arctosa indica—</emphasis>
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<treatmentCitation id="0A88108EFFCBFF90FEF7FED443C5DB38" author="Yin, C. M. &amp; Wang, J. F. &amp; Xie, L. P. &amp; Peng, X. J." box="[321,499,348,372]" page="9" pageId="6" pageNumber="495" year="1993">
<bibRefCitation id="EFB84B6EFFCBFF90FEF7FED443C5DB38" author="Yin, C. M. &amp; Wang, J. F. &amp; Xie, L. P. &amp; Peng, X. J." box="[321,499,348,372]" pageId="6" pageNumber="495" pagination="9 - 18" refId="ref6772" refString="Yin, C. M., Wang, J. F., Xie, L. P. &amp; Peng, X. J. (1993) Some species of the genus Arctosa from the southern China (Araneae: Lycosidae). Acta Arachnologica Sinica, 2, 9 - 18." type="journal article" year="1993">
Yin
<emphasis id="B95DEA8DFFCBFF90FEDAFED443A9DB38" box="[364,415,348,372]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
1993: 9
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, figs 16.
<treatmentCitation id="0A88108EFFCBFF90FDE8FED44107DB38" author="Dhali, D. C. &amp; Saha, S. &amp; Raychaudhuri, D." box="[606,817,348,372]" page="78" pageId="6" pageNumber="495" year="2017">
<bibRefCitation id="EFB84B6EFFCBFF90FDE8FED44107DB38" author="Dhali, D. C. &amp; Saha, S. &amp; Raychaudhuri, D." box="[606,817,348,372]" pageId="6" pageNumber="495" pagination="1 - 242" refId="ref5648" refString="Dhali, D. C., Saha, S. &amp; Raychaudhuri, D. (2017) Litter and ground dwelling spiders (Araneae: Arachnida) of reserve forests of Dooars, West Bengal. World Scientific News, 63, 1 - 242." type="journal article" year="2017">
Dhali
<emphasis id="B95DEA8DFFCBFF90FD2AFED440F9DB38" box="[668,719,348,372]" italics="true" pageId="6" pageNumber="495">et al.</emphasis>
2017: 78
</bibRefCitation>
</treatmentCitation>
, plate XXIII, figs 399403.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3336514FFCBFF90FF21FE2A4616D848" pageId="6" pageNumber="495" type="materials_examined">
<paragraph id="8B96369FFFCBFF90FF21FE2A4616D848" blockId="6.[151,1437,417,516]" pageId="6" pageNumber="495">
<emphasis id="B95DEA8DFFCBFF90FF21FE2A437DDBF7" bold="true" box="[151,331,417,444]" pageId="6" pageNumber="495">Type material.</emphasis>
<materialsCitation id="3B413CC2FFCBFF90FEEEFE2A462AD848" ID-GBIF-Occurrence="3023039303" collectingDate="1975-02-24" collectorName="M. S. Malhotra &amp; Male" country="India" county="Female" elevation="565" latitude="18.53144" location="Shivajinagar" longLatPrecision="1" longitude="73.84457" municipality="Pune" pageId="6" pageNumber="495" specimenCount="2" stateProvince="Maharashtra" typeStatus="holotype">
<collectingCounty id="62F74E13FFCBFF90FEEEFE2A439CDBF0" box="[344,426,418,444]" pageId="6" pageNumber="495">Female</collectingCounty>
<typeStatus id="5492883DFFCBFF90FE01FE2A402FDBF0" box="[439,537,418,444]" pageId="6" pageNumber="495" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F33E760FFFCBFF90FDDCFE2A4088DBF0" box="[618,702,418,444]" name="India" pageId="6" pageNumber="495">INDIA</collectingCountry>
:
<collectingRegion id="49EDF87DFFCBFF90FD78FE2A4168DBF0" box="[718,862,418,444]" country="India" name="Maharashtra" pageId="6" pageNumber="495">Maharashtra</collectingRegion>
:
<collectingMunicipality id="6BF2ACE5FFCBFF90FCD9FE2A4191DBF0" box="[879,935,418,444]" pageId="6" pageNumber="495">Pune</collectingMunicipality>
(=Poona):
<location id="8EF66044FFCBFF90FB87FE2A4689DBF0" LSID="urn:lsid:plazi:treatment:03808789FFCBFF9EFF21FF1F408BDBC5:8EF66044FFCBFF90FB87FE2A4689DBF0" box="[1073,1215,418,444]" country="India" county="Female" latitude="18.53144" longLatPrecision="1" longitude="73.84457" municipality="Pune" name="Shivajinagar" pageId="6" pageNumber="495" stateProvince="Maharashtra">Shivajinagar</location>
(= Shivaji Nagar) (
<geoCoordinate id="EE1D5058FFCBFF90FF16FE4E4365DBAD" box="[160,339,454,481]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="6" pageNumber="495" precision="1" seconds="53.19" value="18.53144">18°3153.19N</geoCoordinate>
,
<geoCoordinate id="EE1D5058FFCBFF90FEEAFE4E403BDBAD" box="[348,525,454,481]" degrees="73" direction="east" minutes="50" orientation="longitude" pageId="6" pageNumber="495" precision="1" seconds="40.42" value="73.84457">73°5040.42E</geoCoordinate>
),
<quantity id="4CD19B7AFFCBFF90FDA8FE4E4055DBAC" box="[542,611,454,481]" metricMagnitude="2" metricUnit="m" metricValue="5.65" pageId="6" pageNumber="495" unit="m" value="565.0">
<elevation id="0004D1ACFFCBFF90FDA8FE4E4055DBAC" box="[542,611,454,481]" metricMagnitude="2" metricUnit="m" metricValue="5.65" pageId="6" pageNumber="495" unit="m" value="565.0">565 m</elevation>
</quantity>
alt.;
<collectorName id="26DC5349FFCBFF90FD21FE4E410FDBAC" box="[663,825,454,480]" pageId="6" pageNumber="495">M.S. Malhotra</collectorName>
leg.;
<date id="FF97105FFFCBFF90FCC4FE4E460EDBAD" box="[882,1080,454,481]" pageId="6" pageNumber="495" value="1975-02-24">
<collectingDate id="EFD3E9B7FFCBFF90FCC4FE4E460EDBAD" box="[882,1080,454,481]" pageId="6" pageNumber="495" value="1975-02-24">24 February 1975</collectingDate>
</date>
; repository NZC-ZSI (4671/18), examined.
<collectorName id="26DC5349FFCBFF90FEA4FE62437AD848" box="[274,332,490,516]" pageId="6" pageNumber="495">Male</collectorName>
<typeStatus id="5492883DFFCBFF90FEE5FE62439BD848" box="[339,429,490,516]" pageId="6" pageNumber="495" type="allotype">allotype</typeStatus>
with same data as holotype, except 4672/18, examined
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF566617FFCBFF90FF21F8254040DDA4" ID-DOI="http://doi.org/10.5281/zenodo.4447147" ID-Zenodo-Dep="4447147" httpUri="https://zenodo.org/record/4447147/files/figure.png" pageId="6" pageNumber="495" startId="6.[151,250,1965,1989]" targetBox="[181,1404,546,1928]" targetPageId="6">
<paragraph id="8B96369FFFCBFF90FF21F8254040DDA4" blockId="6.[151,1436,1965,2025]" pageId="6" pageNumber="495">
<emphasis id="B95DEA8DFFCBFF90FF21F8254324DD89" bold="true" box="[151,274,1965,1989]" pageId="6" pageNumber="495">FIGURE 5.</emphasis>
<taxonomicName id="4C294D1CFFCBFF90FEAEF825408BDD89" authority="Tikader &amp; Malhotra, 1980" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[280,701,1965,1989]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="495" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFCBFF90FEAEF825439DDD89" box="[280,427,1965,1989]" italics="true" pageId="6" pageNumber="495">Arctosa indica</emphasis>
<bibRefCitation id="EFB84B6EFFCBFF90FE07F825408BDD89" author="Tikader, B. K. &amp; Malhotra, M. S." box="[433,701,1965,1989]" pageId="6" pageNumber="495" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra, 1980</bibRefCitation>
</taxonomicName>
.
<emphasis id="B95DEA8DFFCBFF90FD70F82540EEDD89" bold="true" box="[710,728,1965,1989]" pageId="6" pageNumber="495">A</emphasis>
female holotype habitus, dorsal.
<emphasis id="B95DEA8DFFCBFF90FB90F8254601DD89" bold="true" box="[1062,1079,1965,1989]" pageId="6" pageNumber="495">B</emphasis>
male allotype habitus, dorsal.
<emphasis id="B95DEA8DFFCBFF90FADDF82547AADD89" bold="true" box="[1387,1436,1965,1989]" pageId="6" pageNumber="495">CD</emphasis>
labels from type tubes. Scale bars: AB, 2 mm.
</paragraph>
</caption>
<subSubSection id="C3336514FFCAFF91FF71FF1E472BD8BD" pageId="7" pageNumber="496" type="diagnosis">
<paragraph id="8B96369FFFCAFF91FF71FF1E4117DB0E" blockId="7.[151,1437,150,754]" pageId="7" pageNumber="496">
<emphasis id="B95DEA8DFFCAFF91FF71FF1E4374DAFC" bold="true" box="[199,322,150,176]" pageId="7" pageNumber="496">Diagnosis.</emphasis>
<taxonomicName id="4C294D1CFFCAFF91FEF1FF1F43DFDAFD" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[327,489,151,177]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFCAFF91FEF1FF1F43DFDAFD" box="[327,489,151,177]" italics="true" pageId="7" pageNumber="496">Arctosa indica</emphasis>
</taxonomicName>
seem closely related to
<taxonomicName id="4C294D1CFFCAFF91FD44FF1F46ADDAFE" authority="(Schenkel, 1963)" baseAuthorityName="Schenkel" baseAuthorityYear="1963" box="[754,1179,151,178]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="kiangsiensis">
<emphasis id="B95DEA8DFFCAFF91FD44FF1F41E1DAFD" box="[754,983,151,177]" italics="true" pageId="7" pageNumber="496">Arctosa kiangsiensis</emphasis>
(Schenkel, 1963)
</taxonomicName>
as both share a median apophysis with narrow distal part and oval spermathecae with slender, twisted stalks. It can be distinguished from the latter by median apophysis with a small, retrolateral protrusion, which is absent in
<taxonomicName id="4C294D1CFFCAFF91FBE2FF5746CDDAB5" baseAuthorityName="Schenkel" baseAuthorityYear="1963" box="[1108,1275,223,249]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="kiangsiensis">
<emphasis id="B95DEA8DFFCAFF91FBE2FF5746CDDAB5" box="[1108,1275,223,249]" italics="true" pageId="7" pageNumber="496">A. kiangsiensis</emphasis>
</taxonomicName>
, and a median septum with short anterior and broad posterior parts, while
<taxonomicName id="4C294D1CFFCAFF91FC8FFE8B41D4DB51" baseAuthorityName="Schenkel" baseAuthorityYear="1963" box="[825,994,259,285]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="kiangsiensis">
<emphasis id="B95DEA8DFFCAFF91FC8FFE8B41D4DB51" box="[825,994,259,285]" italics="true" pageId="7" pageNumber="496">A. kiangsiensis</emphasis>
</taxonomicName>
has long anterior and narrow posterior parts (compare
<figureCitation id="13122A1AFFCAFF91FEF3FEAF43B0DB0D" box="[325,390,295,321]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1990]" captionTargetBox="[158,1430,785,1933]" captionTargetId="figure-291@7.[151,1437,777,1943]" captionTargetPageId="7" captionText="FIGURE 6. Arctosa indica Tikader &amp; Malhotra, 1980. AC male allotype, left pedipalp: A ventral, B ventral, enlarged; C retrolateral, enlarged. DE female holotype, genitalia: D epigynum, ventral, E vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447151" httpUri="https://zenodo.org/record/4447151/files/figure.png" pageId="7" pageNumber="496">Fig. 6</figureCitation>
BE to
<bibRefCitation id="EFB84B6EFFCAFF91FE6DFEAF40B0DB0D" author="Yin, C. M. &amp; Peng, X. J. &amp; Yan, H. M. &amp; Bao, Y. H. &amp; Xu, X. &amp; Tang, G. &amp; Zhou, Q. S. &amp; Liu, P." box="[475,646,295,321]" pageId="7" pageNumber="496" refId="ref6886" refString="Yin, C. M., Peng, X. J., Yan, H. M., Bao, Y. H., Xu, X., Tang, G., Zhou, Q. S. &amp; Liu, P. (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">
Yin
<emphasis id="B95DEA8DFFCAFF91FDBDFEAF4075DB0D" box="[523,579,295,321]" italics="true" pageId="7" pageNumber="496">et al.</emphasis>
2012
</bibRefCitation>
: fig. 398be).
</paragraph>
<paragraph id="8B96369FFFCAFF91FF71FEC24366D871" blockId="7.[151,1437,150,754]" pageId="7" pageNumber="496">
<emphasis id="B95DEA8DFFCAFF91FF71FEC24023DB28" bold="true" box="[199,533,330,356]" pageId="7" pageNumber="496">Supplementary description.</emphasis>
<emphasis id="B95DEA8DFFCAFF91FDA9FEC3406FDB29" box="[543,601,331,357]" italics="true" pageId="7" pageNumber="496">Male</emphasis>
(
<typeStatus id="5492883DFFCAFF91FDDCFEC340FEDB29" box="[618,712,331,357]" pageId="7" pageNumber="496" type="allotype">allotype</typeStatus>
,
<figureCitation id="13122A1AFFCAFF91FD60FEC34106DB29" box="[726,816,330,357]" captionStart="FIGURE 5" captionStartId="6.[151,250,1965,1989]" captionTargetBox="[181,1404,546,1928]" captionTargetId="figure-146@6.[173,1414,540,1941]" captionTargetPageId="6" captionText="FIGURE 5. Arctosa indica Tikader &amp; Malhotra, 1980. A female holotype habitus, dorsal. B male allotype habitus, dorsal. CD labels from type tubes. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447147" httpUri="https://zenodo.org/record/4447147/files/figure.png" pageId="7" pageNumber="496">Fig. 5B</figureCitation>
). Fovea vertical, dark. Cheliceral promargin with two teeth, retromargin with three. Body length 7.34. Prosoma length 4.15, width 2.99. Opisthosoma length 3.19, width 2.13. Eye diameters: ALE 0.11. AME 0.14. PLE 0.19. PME 0.23. Eye interdistances: ALEAME 0.04. AMEAME 0.06. PLEPLE 0.51. PMEPLE 0.24. PMEPME 0.16. Chelicerae length 2.05. Clypeus height at ALEs 0.11, at AMEs 0.06. Measurements of pedipalp and legs. Pedipalp (right) 3.94 [1.66, 0.74, 0.63, 0.91], I (right) 9.65 [2.86, 1.51, 1.99, 2.13, 1.16], II -----, III 7.85 [2.24, 1.14, 1.19, 2.17, 1.11], IV 11.54 [3.21, 1.46, 2.28, 3.17, 1.42].
<emphasis id="B95DEA8DFFCAFF91FA81FE7747AAD855" box="[1335,1436,511,537]" italics="true" pageId="7" pageNumber="496">Pedipalp</emphasis>
as in
<figureCitation id="13122A1AFFCAFF91FF64FDAB4325D871" box="[210,275,547,573]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1990]" captionTargetBox="[158,1430,785,1933]" captionTargetId="figure-291@7.[151,1437,777,1943]" captionTargetPageId="7" captionText="FIGURE 6. Arctosa indica Tikader &amp; Malhotra, 1980. AC male allotype, left pedipalp: A ventral, B ventral, enlarged; C retrolateral, enlarged. DE female holotype, genitalia: D epigynum, ventral, E vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447151" httpUri="https://zenodo.org/record/4447151/files/figure.png" pageId="7" pageNumber="496">Fig. 6</figureCitation>
AC.
</paragraph>
<paragraph id="8B96369FFFCAFF91FF71FDCF472BD8BD" blockId="7.[151,1437,150,754]" pageId="7" pageNumber="496">
<emphasis id="B95DEA8DFFCAFF91FF71FDCF432DD82D" box="[199,283,583,609]" italics="true" pageId="7" pageNumber="496">Female</emphasis>
(
<typeStatus id="5492883DFFCAFF91FE9CFDCF43A6D82D" box="[298,400,583,609]" pageId="7" pageNumber="496" type="holotype">holotype</typeStatus>
,
<figureCitation id="13122A1AFFCAFF91FE2DFDCF43C2D82D" box="[411,500,582,609]" captionStart="FIGURE 5" captionStartId="6.[151,250,1965,1989]" captionTargetBox="[181,1404,546,1928]" captionTargetId="figure-146@6.[173,1414,540,1941]" captionTargetPageId="6" captionText="FIGURE 5. Arctosa indica Tikader &amp; Malhotra, 1980. A female holotype habitus, dorsal. B male allotype habitus, dorsal. CD labels from type tubes. Scale bars: AB, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447147" httpUri="https://zenodo.org/record/4447147/files/figure.png" pageId="7" pageNumber="496">Fig. 5A</figureCitation>
). Same as male except the following: body length 8.13. Prosoma length 4.28, width 3.20. Opisthosoma length 3.85, width 2.77. Eye diameters: ALE 0.11. AME 0.12. PLE 0.20. PME 0.24. Eye interdistances: ALEAME 0.06. AMEAME 0.12. PLEPLE 0.67. PMEPLE 0.34. PMEPME 0.25. Chelicerae length 2.06. Clypeus height at ALEs 0.11, at AMEs 0.08. Measurements of palp and legs. Palp (right) 4.24 [1.47, 0.87, 0.83, 1.07], I (right) 10.59 [3.17, 1.81, 2.15, 2.10, 1.36], II----, III----, IV----.
<emphasis id="B95DEA8DFFCAFF91FC42FD5F4668D8BD" box="[1012,1118,727,753]" italics="true" pageId="7" pageNumber="496">Genitalia</emphasis>
as in
<figureCitation id="13122A1AFFCAFF91FB16FD5F46D7D8BD" box="[1184,1249,727,753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1990]" captionTargetBox="[158,1430,785,1933]" captionTargetId="figure-291@7.[151,1437,777,1943]" captionTargetPageId="7" captionText="FIGURE 6. Arctosa indica Tikader &amp; Malhotra, 1980. AC male allotype, left pedipalp: A ventral, B ventral, enlarged; C retrolateral, enlarged. DE female holotype, genitalia: D epigynum, ventral, E vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447151" httpUri="https://zenodo.org/record/4447151/files/figure.png" pageId="7" pageNumber="496">Fig. 6</figureCitation>
DE.
</paragraph>
</subSubSection>
<caption id="DF566617FFCAFF91FF21F82647A4DDA6" ID-DOI="http://doi.org/10.5281/zenodo.4447151" ID-Zenodo-Dep="4447151" httpUri="https://zenodo.org/record/4447151/files/figure.png" pageId="7" pageNumber="496" startId="7.[151,250,1966,1990]" targetBox="[158,1430,785,1933]" targetPageId="7">
<paragraph id="8B96369FFFCAFF91FF21F82647A4DDA6" blockId="7.[151,1436,1966,2027]" pageId="7" pageNumber="496">
<emphasis id="B95DEA8DFFCAFF91FF21F8264324DD8A" bold="true" box="[151,274,1966,1990]" pageId="7" pageNumber="496">FIGURE 6.</emphasis>
<taxonomicName id="4C294D1CFFCAFF91FEA1F827408CDD8A" authority="Tikader &amp; Malhotra, 1980" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[279,698,1966,1990]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="496" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFCAFF91FEA1F827439CDD8A" box="[279,426,1966,1990]" italics="true" pageId="7" pageNumber="496">Arctosa indica</emphasis>
<bibRefCitation id="EFB84B6EFFCAFF91FE19F826408CDD8A" author="Tikader, B. K. &amp; Malhotra, M. S." box="[431,698,1966,1990]" pageId="7" pageNumber="496" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra, 1980</bibRefCitation>
</taxonomicName>
.
<emphasis id="B95DEA8DFFCAFF91FD74F82640C5DD8A" bold="true" box="[706,755,1966,1990]" pageId="7" pageNumber="496">AC</emphasis>
male allotype, left pedipalp:
<emphasis id="B95DEA8DFFCAFF91FBA1F826461FDD8A" bold="true" box="[1047,1065,1966,1990]" pageId="7" pageNumber="496">A</emphasis>
ventral,
<emphasis id="B95DEA8DFFCAFF91FB37F82646A4DD8A" bold="true" box="[1153,1170,1966,1990]" pageId="7" pageNumber="496">B</emphasis>
ventral, enlarged;
<emphasis id="B95DEA8DFFCAFF91FAFAF8264768DD8A" bold="true" box="[1356,1374,1966,1990]" pageId="7" pageNumber="496">C</emphasis>
retrolateral, enlarged.
<emphasis id="B95DEA8DFFCAFF91FEF1F85A4341DDA6" bold="true" box="[327,375,2002,2026]" pageId="7" pageNumber="496">DE</emphasis>
female holotype, genitalia:
<emphasis id="B95DEA8DFFCAFF91FD24F85A4092DDA6" bold="true" box="[658,676,2002,2026]" pageId="7" pageNumber="496">D</emphasis>
epigynum, ventral,
<emphasis id="B95DEA8DFFCAFF91FCC6F85A41B7DDA6" bold="true" box="[880,897,2002,2026]" pageId="7" pageNumber="496">E</emphasis>
vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3336514FFC5FF9EFF71FF1F408BDBC5" pageId="8" pageNumber="497" type="discussion">
<paragraph id="8B96369FFFC5FF9EFF71FF1F408BDBC5" blockId="8.[151,1437,151,393]" pageId="8" pageNumber="497">
<emphasis id="B95DEA8DFFC5FF9EFF71FF1F430DDAFD" bold="true" box="[199,315,151,177]" pageId="8" pageNumber="497">Remarks.</emphasis>
The NZC-ZSI collection has two glass tubes for this species. A first tube, labelled holotype (4671/18), contains one female specimen in fairly good condition, with broken legs. The same tube has a small glass vial that contains the dissected genitalia. A second tube, labelled allotype (4672/18), contains one male specimen in fairly good condition, with broken legs. The same tube has a small glass vial that contains the dissected left pedipalp. In contrast to
<bibRefCitation id="EFB84B6EFFC5FF9EFEC5FEAF409CDB0D" author="Tikader, B. K. &amp; Malhotra, M. S." box="[371,682,295,321]" pageId="8" pageNumber="497" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra (1980)</bibRefCitation>
, who mentioned the collection locality of both male and female as Shivajinagar, the
<typeStatus id="5492883DFFC5FF9EFEDBFEC343F1DB29" box="[365,455,331,357]" pageId="8" pageNumber="497" type="allotype">allotype</typeStatus>
label, which lacks collector name and date of collection, mentions the collection locality as Gamatia in Birbhum district of West Bengal.
</paragraph>
</subSubSection>
</treatment>
</document>