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<document id="E5959C813CCFC0E68DA5BB31ABED58CD" ID-CLB-Dataset="8713" ID-DOI="10.11646/zootaxa.4908.4.3" ID-GBIF-Dataset="b1fca0fe-4044-4293-bc35-4a316bd84b17" ID-ISSN="1175-5326" ID-Zenodo-Dep="4447127" ID-ZooBank="2F90F982-E4B7-46D6-A7CA-C666496B5ACE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610977825505" checkinUser="plazi" docAuthor="Sankaran, Pradeep M., Caleb, John T. D. &amp; Sebastian, Pothalil A." docDate="2021" docId="03808789FFC5FF9CFF21F9E74165DD20" docLanguage="en" docName="zootaxa4908.4.3.pdf" docOrigin="Zootaxa 4908 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arctosa lesserti Reimoser 1934" docType="treatment" docVersion="7" lastPageNumber="499" masterDocId="FFB9FFF1FFCDFF96FFB6FF884236DA4C" masterDocTitle="Revision of Indian wolf spiders: I. Genus Arctosa C. L. Koch, 1847 (Araneae Lycosidae, Tricassinae)" masterLastPageNumber="504" masterPageNumber="489" pageNumber="497" updateTime="1698924574895" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7A87E4A42922CAB592546A2FB05BE8BC">Revision of Indian wolf spiders: I. Genus Arctosa C. L. Koch, 1847 (Araneae Lycosidae, Tricassinae)</mods:title>
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<mods:namePart id="ECF489C800AD5B3DA7764CE6EE9104E0">Sankaran, Pradeep M.</mods:namePart>
<mods:affiliation id="FA452F03AD4DA37A22BD4D6648FA5A77">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
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<mods:namePart id="B97FED5A968B7E15BD54A3EAB5FABF9E">Caleb, John T. D.</mods:namePart>
<mods:nameIdentifier id="9931A5B2FF8A897538C24555AC6C06E9" type="ORCID">0000-0002-9471-9467</mods:nameIdentifier>
<mods:affiliation id="95D8EE8208EA7E1520FBDA3570A60CA5">Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India. caleb 87 woodgate @ gmail. com; https: // orcid. org / 0000 - 0002 - 9471 - 9467</mods:affiliation>
<mods:nameIdentifier id="3544E008F1E29D9C06644548CC832F58" type="email">caleb87woodgate@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B9D3EE5772C2A765E2F6708647A1526D">Sebastian, Pothalil A.</mods:namePart>
<mods:nameIdentifier id="A533A1F12AF89FF759C5F64D0327BFCD" type="ORCID">0000-0002-4936-4310</mods:nameIdentifier>
<mods:affiliation id="ACD28EBFD187E79D18EA11A369AB7E85">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India. &amp; drpothalil @ rediffmail. com; https: // orcid. org / 0000 - 0002 - 4936 - 4310</mods:affiliation>
<mods:nameIdentifier id="2ACA8DDB079061CC2C38AD15127625AB" type="email">drpothalil@rediffmail.com</mods:nameIdentifier>
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<mods:date id="B22E9427C6746AE60CCC9B425C399D92">2021</mods:date>
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<treatment id="03808789FFC5FF9CFF21F9E74165DD20" ID-DOI="http://doi.org/10.5281/zenodo.4447141" ID-GBIF-Taxon="177013997" ID-Zenodo-Dep="4447141" LSID="urn:lsid:plazi:treatment:03808789FFC5FF9CFF21F9E74165DD20" httpUri="http://treatment.plazi.org/id/03808789FFC5FF9CFF21F9E74165DD20" lastPageId="10" lastPageNumber="499" pageId="8" pageNumber="497">
<subSubSection id="C3336514FFC5FF9EFF21F9E74023DCC4" box="[151,533,1646,1673]" pageId="8" pageNumber="497" type="nomenclature">
<paragraph id="8B96369FFFC5FF9EFF21F9E74023DCC4" blockId="8.[151,533,1646,1709]" box="[151,533,1646,1673]" pageId="8" pageNumber="497">
<heading id="D0DE81F3FFC5FF9EFF21F9E74023DCC4" bold="true" box="[151,533,1646,1673]" fontSize="11" level="1" pageId="8" pageNumber="497" reason="1">
<taxonomicName id="4C294D1CFFC5FF9EFF21F9E74023DCC4" ID-CoL="5W3S9" authority="Reimoser, 1934" authorityName="Reimoser" authorityYear="1934" box="[151,533,1646,1673]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC5FF9EFF21F9E74023DCC4" bold="true" box="[151,533,1646,1673]" pageId="8" pageNumber="497">
<emphasis id="B95DEA8DFFC5FF9EFF21F9E74366DCC5" bold="true" box="[151,336,1646,1673]" italics="true" pageId="8" pageNumber="497">Arctosa lesserti</emphasis>
<bibRefCitation id="EFB84B6EFFC5FF9EFEE1F9E64023DCC4" author="Reimoser, E." box="[343,533,1646,1672]" pageId="8" pageNumber="497" pagination="465 - 511" refId="ref6327" refString="Reimoser, E. (1934) Araneae aus Sud-Indien. Revue Suisse de Zoologie, 41, 465 - 511. https: // doi. org / 10.5962 / bhl. part. 155628" type="journal article" year="1934">Reimoser, 1934</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C3336514FFC5FF9EFF21F91B4330DCE1" box="[151,262,1683,1709]" pageId="8" pageNumber="497" type="description">
<paragraph id="8B96369FFFC5FF9EFF21F91B4330DCE1" blockId="8.[151,533,1646,1709]" box="[151,262,1683,1709]" pageId="8" pageNumber="497">
<figureCitation id="13122A1AFFC5FF9EFF21F91B4330DCE1" box="[151,262,1683,1709]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="11.[151,250,1477,1501]" captionStartId-1="12.[151,250,1220,1244]" captionTargetBox-0="[166,1424,189,1432]" captionTargetBox-1="[158,1427,246,1190]" captionTargetId-0="figure-20@11.[151,1436,181,1453]" captionTargetId-1="figure-50@12.[151,1436,237,1197]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 9. AB Arctosa lesserti Reimoser, 1934. A female lectotype habitus, dorsal. B female paralectotype habitus, dorsal. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, habitus, dorsal. D label from type tube. Scale bars: AC, 1 mm. © Christoph Hörweg, NHM, Vienna." captionText-1="FIGURE 10. AB Arctosa lesserti Reimoser, 1934. A female lectotype epigynum, ventral. B female paralectotype epigynum, ventral. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, epigynum, ventral. Scale bars: 0.1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi-0="http://doi.org/10.5281/zenodo.4447157" figureDoi-1="http://doi.org/10.5281/zenodo.4447159" httpUri-0="https://zenodo.org/record/4447157/files/figure.png" httpUri-1="https://zenodo.org/record/4447159/files/figure.png" pageId="8" pageNumber="497">Figs 910</figureCitation>
</paragraph>
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<paragraph id="8B96369FFFC5FF9EFF21F95F4042DCA3" blockId="8.[151,1437,1751,1837]" box="[151,628,1751,1775]" pageId="8" pageNumber="497">
<treatmentCitationGroup id="AB3911B1FFC5FF9EFF21F95F4042DCA3" box="[151,628,1751,1775]" pageId="8" pageNumber="497">
<treatmentCitation id="0A88108EFFC5FF9EFF21F95F4026DCA3" author="Reimoser, E." box="[151,528,1751,1775]" page="470" pageId="8" pageNumber="497" year="1934">
<taxonomicName id="4C294D1CFFC5FF9EFF21F95F4026DCA3" ID-CoL="5W3S9" authority="Reimoser, 1934: 470" authorityName="Reimoser" authorityPageNumber="470" authorityYear="1934" box="[151,528,1751,1775]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC5FF9EFF21F95F4302DCA3" box="[151,308,1751,1775]" italics="true" pageId="8" pageNumber="497">Arctosa lesserti</emphasis>
<bibRefCitation id="EFB84B6EFFC5FF9EFE8DF95F4026DCA3" author="Reimoser, E." box="[315,528,1751,1775]" pageId="8" pageNumber="497" pagination="465 - 511" refId="ref6327" refString="Reimoser, E. (1934) Araneae aus Sud-Indien. Revue Suisse de Zoologie, 41, 465 - 511. https: // doi. org / 10.5962 / bhl. part. 155628" type="journal article" year="1934">Reimoser, 1934: 470</bibRefCitation>
</taxonomicName>
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, figs 23.
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</paragraph>
<paragraph id="8B96369FFFC5FF9EFF21F97E4778DD60" blockId="8.[151,1437,1751,1837]" pageId="8" pageNumber="497">
<treatmentCitationGroup id="AB3911B1FFC5FF9EFF21F97E4778DD60" pageId="8" pageNumber="497">
<taxonomicName id="4C294D1CFFC5FF9EFF21F97E4307DD42" ID-CoL="GBGY" baseAuthorityName="Dyal" baseAuthorityYear="1935" box="[151,305,1782,1806]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="497" phylum="Arthropoda" rank="species" species="mulani">
<emphasis id="B95DEA8DFFC5FF9EFF21F97E4307DD42" box="[151,305,1782,1806]" italics="true" pageId="8" pageNumber="497">Arctosa mulani</emphasis>
</taxonomicName>
<treatmentCitation id="0A88108EFFC5FF9EFEFCF97E40B4DD42" author="Tikader, B. K. &amp; Malhotra, M. S." box="[330,642,1782,1806]" page="373" pageId="8" pageNumber="497" year="1980">
<bibRefCitation id="EFB84B6EFFC5FF9EFEFCF97E40B4DD42" author="Tikader, B. K. &amp; Malhotra, M. S." box="[330,642,1782,1806]" pageId="8" pageNumber="497" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra 1980: 373</bibRefCitation>
</treatmentCitation>
, figs 247251 (
<typeStatus id="5492883DFFC5FF9EFCABF97E4159DD42" box="[797,879,1782,1806]" pageId="8" pageNumber="497" type="neotype">neotype</typeStatus>
designation; transfer from
<taxonomicName id="4C294D1CFFC5FF9EFB36F97E46E3DD42" ID-CoL="6HK8" authorityName="C. L. Koch" authorityYear="1847" box="[1152,1237,1782,1806]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8DFFC5FF9EFB36F97E46E3DD42" box="[1152,1237,1782,1806]" italics="true" pageId="8" pageNumber="497">Pardosa</emphasis>
</taxonomicName>
; misidentification).
<treatmentCitation id="0A88108EFFC5FF9EFF71F89C43A6DD60" author="Majumder, S. C." box="[199,400,1812,1836]" page="26" pageId="8" pageNumber="497" year="2005">
<bibRefCitation id="EFB84B6EFFC5FF9EFF71F89C43A6DD60" author="Majumder, S. C." box="[199,400,1812,1836]" pageId="8" pageNumber="497" pagination="1 - 57" refId="ref6250" refString="Majumder, S. C. (2005) Studies on some spiders from eastern coastal region of India. Memoirs of the Zoological Survey of India, 20, 1 - 57." type="journal article" year="2005">Majumder 2005: 26</bibRefCitation>
</treatmentCitation>
(misidentification).
<treatmentCitation id="0A88108EFFC5FF9EFDD6F89D4115DD61" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[608,803,1812,1837]" page="130" pageId="8" pageNumber="497" year="2016">
<bibRefCitation id="EFB84B6EFFC5FF9EFDD6F89D4115DD61" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[608,803,1812,1837]" pageId="8" pageNumber="497" pagination="130 - 142" refId="ref6107" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B95DEA8DFFC5FF9EFD34F89D4080DD60" box="[642,694,1812,1836]" italics="true" pageId="8" pageNumber="497">et al.</emphasis>
2016: 130
</bibRefCitation>
</treatmentCitation>
, figs 1AH, 2AD, 3AD, 12AB (misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3336514FFC5FF9EFF21F8D34693DDF1" pageId="8" pageNumber="497" type="materials_examined">
<paragraph id="8B96369FFFC5FF9EFF21F8D34693DDF1" blockId="8.[151,1437,1882,1982]" pageId="8" pageNumber="497">
<emphasis id="B95DEA8DFFC5FF9EFF21F8D34373DD38" bold="true" box="[151,325,1882,1909]" pageId="8" pageNumber="497">Type material.</emphasis>
<materialsCitation id="3B413CC2FFC5FF9EFEFAF8D34697DDF1" ID-GBIF-Occurrence="3023039311" collectingDate="1934-10-15" collectorName="Reimoser" country="India" county="Female" elevation="942" latitude="11.572178" location="Masinagudi" longLatPrecision="1" longitude="76.642715" municipality="Nilgiris" pageId="8" pageNumber="497" specimenCount="2" stateProvince="Tamil Nadu" typeStatus="lectotype">
<collectingCounty id="62F74E13FFC5FF9EFEFAF8D343A8DD39" box="[332,414,1883,1909]" pageId="8" pageNumber="497">Female</collectingCounty>
<typeStatus id="5492883DFFC5FF9EFE10F8D3403BDD39" box="[422,525,1883,1909]" pageId="8" pageNumber="497" type="lectotype">lectotype</typeStatus>
(here designated) and male and female
<typeStatus id="5492883DFFC5FF9EFC7AF8D34658DD39" box="[972,1134,1883,1909]" pageId="8" pageNumber="497" type="paralectotype">paralectotypes</typeStatus>
from
<collectingCountry id="F33E760FFFC5FF9EFB04F8D34730DD39" box="[1202,1286,1883,1909]" name="India" pageId="8" pageNumber="497">INDIA</collectingCountry>
:
<collectingRegion id="49EDF87DFFC5FF9EFAA6F8D347AFDD39" box="[1296,1433,1883,1909]" country="India" name="Tamil Nadu" pageId="8" pageNumber="497">Tamil Nadu</collectingRegion>
:
<collectingMunicipality id="6BF2ACE5FFC5FF9EFF21F8F742C7DDD5" box="[151,241,1919,1945]" pageId="8" pageNumber="497">Nilgiris</collectingMunicipality>
:
<location id="8EF66044FFC5FF9EFF4DF8F74349DDD5" LSID="urn:lsid:plazi:treatment:03808789FFC5FF9CFF21F9E74165DD20:8EF66044FFC5FF9EFF4DF8F74349DDD5" box="[251,383,1919,1945]" country="India" county="Female" latitude="11.572178" longLatPrecision="1" longitude="76.642715" municipality="Nilgiris" name="Masinagudi" pageId="8" pageNumber="497" stateProvince="Tamil Nadu">Masinagudi</location>
(=Masnigudi) (
<geoCoordinate id="EE1D5058FFC5FF9EFD99F8F740D7DDD5" box="[559,737,1919,1945]" degrees="11" direction="north" minutes="34" orientation="latitude" pageId="8" pageNumber="497" precision="1" seconds="19.84" value="11.572178">11°3419.84N</geoCoordinate>
,
<geoCoordinate id="EE1D5058FFC5FF9EFD5BF8F741ABDDD6" box="[749,925,1919,1946]" degrees="76" direction="east" minutes="38" orientation="longitude" pageId="8" pageNumber="497" precision="1" seconds="33.77" value="76.642715">76°3833.77E</geoCoordinate>
),
<quantity id="4CD19B7AFFC5FF9EFC07F8F741CEDDD5" box="[945,1016,1919,1945]" metricMagnitude="2" metricUnit="m" metricValue="9.42" pageId="8" pageNumber="497" unit="m" value="942.0">
<elevation id="0004D1ACFFC5FF9EFC07F8F741CEDDD5" box="[945,1016,1919,1945]" metricMagnitude="2" metricUnit="m" metricValue="9.42" pageId="8" pageNumber="497" unit="m" value="942.0">942 m</elevation>
</quantity>
alt.;
<collectorName id="26DC5349FFC5FF9EFB86F8F746ADDDD5" box="[1072,1179,1919,1945]" pageId="8" pageNumber="497">Reimoser</collectorName>
leg.;
<date id="FF97105FFFC5FF9EFB6FF8F647A1DDD5" box="[1241,1431,1918,1945]" pageId="8" pageNumber="497" value="1934-10-15">
<collectingDate id="EFD3E9B7FFC5FF9EFB6FF8F647A1DDD5" box="[1241,1431,1918,1945]" pageId="8" pageNumber="497" value="1934-10-15">15 October 1934</collectingDate>
</date>
; repository NHM (no register number specified), examined (only females) from photographs
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF566617FFC4FF9FFF21F80B401DDDF3" ID-DOI="http://doi.org/10.5281/zenodo.4447153" ID-Zenodo-Dep="4447153" httpUri="https://zenodo.org/record/4447153/files/figure.png" pageId="9" pageNumber="498" startId="9.[151,250,1923,1947]" targetBox="[158,1428,190,1891]" targetPageId="9">
<paragraph id="8B96369FFFC4FF9FFF21F80B401DDDF3" blockId="9.[151,1436,1923,1984]" pageId="9" pageNumber="498">
<emphasis id="B95DEA8DFFC4FF9FFF21F80B4322DDD7" bold="true" box="[151,276,1923,1947]" pageId="9" pageNumber="498">FIGURE 7.</emphasis>
<taxonomicName id="4C294D1CFFC4FF9FFEADF80B4046DDD7" authority="(Sinha, 1951)" baseAuthorityName="Sinha" baseAuthorityYear="1951" box="[283,624,1923,1947]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="498" phylum="Arthropoda" rank="species" species="khudiensis">
<emphasis id="B95DEA8DFFC4FF9FFEADF80B43EDDDD7" box="[283,475,1923,1947]" italics="true" pageId="9" pageNumber="498">Arctosa khudiensis</emphasis>
(
<bibRefCitation id="EFB84B6EFFC4FF9FFE5CF80B405EDDD7" author="Sinha, T. B." box="[490,616,1923,1947]" pageId="9" pageNumber="498" pagination="9 - 52" refId="ref6478" refString="Sinha, T. B. (1951) On the collection of lycosid spiders in the Zoological Survey of India. Records of the Indian Museum, Calcutta, 48, 9 - 52." type="journal article" year="1951">Sinha, 1951</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B95DEA8DFFC4FF9FFDCDF80B40BBDDD7" bold="true" box="[635,653,1923,1947]" pageId="9" pageNumber="498">A</emphasis>
female holotype habitus, dorsal. B female paratype habitus, dorsal.
<emphasis id="B95DEA8DFFC4FF9FFAFEF80B476CDDD7" bold="true" box="[1352,1370,1923,1947]" pageId="9" pageNumber="498">C</emphasis>
labels from type tube. Scale bars: AB, 2 mm.
</paragraph>
</caption>
<caption id="DF566617FFC7FF9CFF21FB4340A0DF4B" ID-DOI="http://doi.org/10.5281/zenodo.4447155" ID-Zenodo-Dep="4447155" httpUri="https://zenodo.org/record/4447155/files/figure.png" pageId="10" pageNumber="499" startId="10.[151,250,1227,1251]" targetBox="[158,1430,191,1198]" targetPageId="10">
<paragraph id="8B96369FFFC7FF9CFF21FB4340A0DF4B" blockId="10.[151,1436,1227,1287]" pageId="10" pageNumber="499">
<emphasis id="B95DEA8DFFC7FF9CFF21FB434324DEAF" bold="true" box="[151,274,1227,1251]" pageId="10" pageNumber="499">FIGURE 8.</emphasis>
<taxonomicName id="4C294D1CFFC7FF9CFEAEFB44405FDEAF" authority="(Sinha, 1951)" baseAuthorityName="Sinha" baseAuthorityYear="1951" box="[280,617,1227,1251]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="khudiensis">
<emphasis id="B95DEA8DFFC7FF9CFEAEFB4443E1DEAF" box="[280,471,1227,1251]" italics="true" pageId="10" pageNumber="499">Arctosa khudiensis</emphasis>
(
<bibRefCitation id="EFB84B6EFFC7FF9CFE52FB434057DEAF" author="Sinha, T. B." box="[484,609,1227,1251]" pageId="10" pageNumber="499" pagination="9 - 52" refId="ref6478" refString="Sinha, T. B. (1951) On the collection of lycosid spiders in the Zoological Survey of India. Records of the Indian Museum, Calcutta, 48, 9 - 52." type="journal article" year="1951">Sinha, 1951</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B95DEA8DFFC7FF9CFDC4FB4340B2DEAF" bold="true" box="[626,644,1227,1251]" pageId="10" pageNumber="499">A</emphasis>
female holotype, epigynum, ventral.
<emphasis id="B95DEA8DFFC7FF9CFC4BFB43461BDEAF" bold="true" box="[1021,1069,1227,1251]" pageId="10" pageNumber="499">BC</emphasis>
female paratype, genitalia:
<emphasis id="B95DEA8DFFC7FF9CFAF3FB434760DEAF" bold="true" box="[1349,1366,1227,1251]" pageId="10" pageNumber="499">B</emphasis>
epigynum, ventral,
<emphasis id="B95DEA8DFFC7FF9CFE92FB674300DF4B" bold="true" box="[292,310,1263,1287]" pageId="10" pageNumber="499">C</emphasis>
vulva, dorsal. Scale bars: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3336514FFC7FF9CFF71FABD405CDC64" pageId="10" pageNumber="499" type="diagnosis">
<paragraph id="8B96369FFFC7FF9CFF71FABD405CDC64" blockId="10.[151,1437,1333,1900]" pageId="10" pageNumber="499">
<emphasis id="B95DEA8DFFC7FF9CFF71FABD4374DF03" bold="true" box="[199,322,1333,1359]" pageId="10" pageNumber="499">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C294D1CFFC7FF9CFE0FFABE401DDF03" authorityName="Reimoser" authorityYear="1934" box="[441,555,1333,1360]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC7FF9CFE0FFABE401DDF03" box="[441,555,1333,1360]" italics="true" pageId="10" pageNumber="499">A. lesserti</emphasis>
</taxonomicName>
seem closely related to the males of
<taxonomicName id="4C294D1CFFC7FF9CFC6CFABE4777DF03" authority="Li, 2016" authorityName="Li" authorityYear="2016" box="[986,1345,1333,1360]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="zhaojingzhaoi">
<emphasis id="B95DEA8DFFC7FF9CFC6CFABE46E0DF03" box="[986,1238,1333,1360]" italics="true" pageId="10" pageNumber="499">Arctosa zhaojingzhaoi</emphasis>
Li, 2016
</taxonomicName>
as both share a downwardly directed median apophysis, but can be distinguished from the latter by uniformly thick median apophysis, which is narrow distally in
<taxonomicName id="4C294D1CFFC7FF9CFDFEFAF64133DFDB" authorityName="Li" authorityYear="2016" box="[584,773,1405,1432]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="zhaojingzhaoi">
<emphasis id="B95DEA8DFFC7FF9CFDFEFAF64133DFDB" box="[584,773,1405,1432]" italics="true" pageId="10" pageNumber="499">A. zhaojingzhaoi</emphasis>
</taxonomicName>
(compare
<bibRefCitation id="EFB84B6EFFC7FF9CFCCBFAF6461CDFDB" author="Pan, F. &amp; Zheng, G. &amp; Li, S. Q." box="[893,1066,1405,1432]" pageId="10" pageNumber="499" pagination="404 - 414" refId="ref6284" refString="Pan, F., Zheng, G. &amp; Li, S. Q. (2016) Wolf spiders (Araneae: Lycosidae) from Xishuangbanna Tropical Botanical Garden, China. Zoological Systematics, 41, 404 - 414." type="journal article" year="2016">
Pan
<emphasis id="B95DEA8DFFC7FF9CFC18FAF641D1DFDB" box="[942,999,1405,1431]" italics="true" pageId="10" pageNumber="499">et al.</emphasis>
2016
</bibRefCitation>
: fig. 2AB to
<bibRefCitation id="EFB84B6EFFC7FF9CFB7FFAF6475DDFDB" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[1225,1387,1405,1432]" pageId="10" pageNumber="499" pagination="130 - 142" refId="ref6107" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B95DEA8DFFC7FF9CFB46FAF6471EDFDB" box="[1264,1320,1405,1431]" italics="true" pageId="10" pageNumber="499">et al.</emphasis>
2016
</bibRefCitation>
: fig. 1CF). Females of
<taxonomicName id="4C294D1CFFC7FF9CFEDDFA2A43ECDFF7" authorityName="Reimoser" authorityYear="1934" box="[363,474,1441,1468]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC7FF9CFEDDFA2A43ECDFF7" box="[363,474,1441,1468]" italics="true" pageId="10" pageNumber="499">A. lesserti</emphasis>
</taxonomicName>
closely resemble the females of
<taxonomicName id="4C294D1CFFC7FF9CFCF7FA2A4193DFF7" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[833,933,1441,1468]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFC7FF9CFCF7FA2A4193DFF7" box="[833,933,1441,1468]" italics="true" pageId="10" pageNumber="499">A. indica</emphasis>
</taxonomicName>
as both share oval spermathecae with slender stalks, but can be distinguished from the latter by distally narrowing anterior part of median septum, which is uniform in thickness in
<taxonomicName id="4C294D1CFFC7FF9CFECAFA6243D6DC4F" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[380,480,1513,1540]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFC7FF9CFECAFA6243D6DC4F" box="[380,480,1513,1540]" italics="true" pageId="10" pageNumber="499">A. indica</emphasis>
</taxonomicName>
, and pear-shaped epigynal atria, which are circular in
<taxonomicName id="4C294D1CFFC7FF9CFB8BFA624694DC4F" authorityName="Tikader &amp; Malhotra" authorityYear="1980" box="[1085,1186,1513,1540]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95DEA8DFFC7FF9CFB8BFA624694DC4F" box="[1085,1186,1513,1540]" italics="true" pageId="10" pageNumber="499">A. indica</emphasis>
</taxonomicName>
(compare
<figureCitation id="13122A1AFFC7FF9CFAAEFA614753DC48" box="[1304,1381,1513,1540]" captionStart="FIGURE 10" captionStartId="12.[151,250,1220,1244]" captionTargetBox="[158,1427,246,1190]" captionTargetId="figure-50@12.[151,1436,237,1197]" captionTargetPageId="12" captionText="FIGURE 10. AB Arctosa lesserti Reimoser, 1934. A female lectotype epigynum, ventral. B female paralectotype epigynum, ventral. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, epigynum, ventral. Scale bars: 0.1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447159" httpUri="https://zenodo.org/record/4447159/files/figure.png" pageId="10" pageNumber="499">Fig. 10</figureCitation>
AB to
<figureCitation id="13122A1AFFC7FF9CFF02F98542C2DC6B" box="[180,244,1549,1575]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1990]" captionTargetBox="[158,1430,785,1933]" captionTargetId="figure-291@7.[151,1437,777,1943]" captionTargetPageId="7" captionText="FIGURE 6. Arctosa indica Tikader &amp; Malhotra, 1980. AC male allotype, left pedipalp: A ventral, B ventral, enlarged; C retrolateral, enlarged. DE female holotype, genitalia: D epigynum, ventral, E vulva, dorsal. Scale bars: A, 0.5 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447151" httpUri="https://zenodo.org/record/4447151/files/figure.png" pageId="10" pageNumber="499">Fig. 6</figureCitation>
DE and
<bibRefCitation id="EFB84B6EFFC7FF9CFEE9F9864036DC6B" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[351,512,1549,1576]" pageId="10" pageNumber="499" pagination="130 - 142" refId="ref6107" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B95DEA8DFFC7FF9CFE33F986438BDC6B" box="[389,445,1549,1575]" italics="true" pageId="10" pageNumber="499">et al.</emphasis>
2016
</bibRefCitation>
: fig. 1G).
</paragraph>
</subSubSection>
<subSubSection id="C3336514FFC7FF9CFF71F9B94065DC00" box="[199,595,1585,1612]" pageId="10" pageNumber="499" type="description">
<paragraph id="8B96369FFFC7FF9CFF71F9B94065DC00" blockId="10.[151,1437,1333,1900]" box="[199,595,1585,1612]" pageId="10" pageNumber="499">
<emphasis id="B95DEA8DFFC7FF9CFF71F9B9436FDC07" bold="true" box="[199,345,1585,1611]" pageId="10" pageNumber="499">Description.</emphasis>
See
<bibRefCitation id="EFB84B6EFFC7FF9CFE39F9B94079DC00" author="Reimoser, E." box="[399,591,1585,1612]" pageId="10" pageNumber="499" pagination="465 - 511" refId="ref6327" refString="Reimoser, E. (1934) Araneae aus Sud-Indien. Revue Suisse de Zoologie, 41, 465 - 511. https: // doi. org / 10.5962 / bhl. part. 155628" type="journal article" year="1934">Reimoser (1934)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3336514FFC7FF9CFF71F9DE4165DD20" pageId="10" pageNumber="499" type="discussion">
<paragraph id="8B96369FFFC7FF9CFF71F9DE46F7DC97" blockId="10.[151,1437,1333,1900]" pageId="10" pageNumber="499">
<emphasis id="B95DEA8DFFC7FF9CFF71F9DE430DDC3C" bold="true" box="[199,315,1622,1648]" pageId="10" pageNumber="499">Remarks.</emphasis>
The NHM collection has three female specimens in good condition under the name
<taxonomicName id="4C294D1CFFC7FF9CFA97F9DE47A3DC23" authorityName="Reimoser" authorityYear="1934" box="[1313,1429,1621,1648]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC7FF9CFA97F9DE4704DC3C" box="[1313,1330,1622,1648]" italics="true" pageId="10" pageNumber="499">A</emphasis>
.
<emphasis id="B95DEA8DFFC7FF9CFAF2F9DD47A3DC23" box="[1348,1429,1621,1647]" italics="true" pageId="10" pageNumber="499">lesserti</emphasis>
</taxonomicName>
, without any male specimen (Hörweg, pers. comm.). Of the three female specimens, two belong to
<taxonomicName id="4C294D1CFFC7FF9CFB5DF9F2476ADCDF" authorityName="Reimoser" authorityYear="1934" box="[1259,1372,1657,1684]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC7FF9CFB5DF9F246CADCD8" box="[1259,1276,1658,1684]" italics="true" pageId="10" pageNumber="499">A</emphasis>
.
<emphasis id="B95DEA8DFFC7FF9CFABDF9F1476ADCDF" box="[1291,1372,1657,1683]" italics="true" pageId="10" pageNumber="499">lesserti</emphasis>
</taxonomicName>
(
<figureCitation id="13122A1AFFC7FF9CFADCF9F14292DCF4" captionStart="FIGURE 9" captionStartId="11.[151,250,1477,1501]" captionTargetBox="[166,1424,189,1432]" captionTargetId="figure-20@11.[151,1436,181,1453]" captionTargetPageId="11" captionText="FIGURE 9. AB Arctosa lesserti Reimoser, 1934. A female lectotype habitus, dorsal. B female paralectotype habitus, dorsal. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, habitus, dorsal. D label from type tube. Scale bars: AC, 1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447157" httpUri="https://zenodo.org/record/4447157/files/figure.png" pageId="10" pageNumber="499">Figs 9</figureCitation>
AB, 10AB) while the third is of
<taxonomicName id="4C294D1CFFC7FF9CFD8FF91541F8DCF4" authority="Kronestedt, 2017" authorityName="Kronestedt" authorityYear="2017" box="[569,974,1693,1720]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="ghatica">
<emphasis id="B95DEA8DFFC7FF9CFD8FF9154134DCFB" box="[569,770,1693,1719]" italics="true" pageId="10" pageNumber="499">Wadicosa ghatica</emphasis>
<bibRefCitation id="EFB84B6EFFC7FF9CFCBDF91541F8DCF4" author="Kronestedt, T." box="[779,974,1693,1720]" pageId="10" pageNumber="499" pagination="295 - 300" refId="ref6031" refString="Kronestedt, T. (2017) Species of Wadicosa (Araneae, Lycosidae): a new species close to W. quadrifera (Gravely) from the Western Ghats, India. Zootaxa, 4300 (2), 295 - 300. https: // doi. org / 10.11646 / zootaxa. 4300.2.11" type="journal article" year="2017">Kronestedt, 2017</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13122A1AFFC7FF9CFC69F915460ADCF4" box="[991,1084,1693,1720]" captionStart="FIGURE 9" captionStartId="11.[151,250,1477,1501]" captionTargetBox="[166,1424,189,1432]" captionTargetId="figure-20@11.[151,1436,181,1453]" captionTargetPageId="11" captionText="FIGURE 9. AB Arctosa lesserti Reimoser, 1934. A female lectotype habitus, dorsal. B female paralectotype habitus, dorsal. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, habitus, dorsal. D label from type tube. Scale bars: AC, 1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447157" httpUri="https://zenodo.org/record/4447157/files/figure.png" pageId="10" pageNumber="499">Figs 9C</figureCitation>
,
<figureCitation id="13122A1AFFC7FF9CFBFCF916464CDCF4" box="[1098,1146,1694,1720]" captionStart="FIGURE 10" captionStartId="12.[151,250,1220,1244]" captionTargetBox="[158,1427,246,1190]" captionTargetId="figure-50@12.[151,1436,237,1197]" captionTargetPageId="12" captionText="FIGURE 10. AB Arctosa lesserti Reimoser, 1934. A female lectotype epigynum, ventral. B female paralectotype epigynum, ventral. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, epigynum, ventral. Scale bars: 0.1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447159" httpUri="https://zenodo.org/record/4447159/files/figure.png" pageId="10" pageNumber="499">10C</figureCitation>
) (compare
<figureCitation id="13122A1AFFC7FF9CFB48F9154757DCF4" box="[1278,1377,1693,1720]" captionStart="FIGURE 10" captionStartId="12.[151,250,1220,1244]" captionTargetBox="[158,1427,246,1190]" captionTargetId="figure-50@12.[151,1436,237,1197]" captionTargetPageId="12" captionText="FIGURE 10. AB Arctosa lesserti Reimoser, 1934. A female lectotype epigynum, ventral. B female paralectotype epigynum, ventral. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, epigynum, ventral. Scale bars: 0.1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447159" httpUri="https://zenodo.org/record/4447159/files/figure.png" pageId="10" pageNumber="499">Fig. 10C</figureCitation>
with
<bibRefCitation id="EFB84B6EFFC7FF9CFF21F9494363DC90" author="Kronestedt, T." box="[151,341,1729,1756]" pageId="10" pageNumber="499" pagination="295 - 300" refId="ref6031" refString="Kronestedt, T. (2017) Species of Wadicosa (Araneae, Lycosidae): a new species close to W. quadrifera (Gravely) from the Western Ghats, India. Zootaxa, 4300 (2), 295 - 300. https: // doi. org / 10.11646 / zootaxa. 4300.2.11" type="journal article" year="2017">Kronestedt 2017</bibRefCitation>
: fig. 15). The male specimen might be lost, so we could not redescribe it here.
</paragraph>
<paragraph id="8B96369FFFC7FF9CFF71F96D4165DD20" blockId="10.[151,1437,1333,1900]" pageId="10" pageNumber="499">
The illustrations and colour images of the epigynum of
<taxonomicName id="4C294D1CFFC7FF9CFC8FF96E4191DCB3" baseAuthorityName="Dyal" baseAuthorityYear="1935" box="[825,935,1765,1792]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="mulani">
<emphasis id="B95DEA8DFFC7FF9CFC8FF96E4191DCB3" box="[825,935,1765,1792]" italics="true" pageId="10" pageNumber="499">A. mulani</emphasis>
</taxonomicName>
presented in
<bibRefCitation id="EFB84B6EFFC7FF9CFB8BF96D475DDD4C" author="Tikader, B. K. &amp; Malhotra, M. S." box="[1085,1387,1765,1792]" pageId="10" pageNumber="499" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra (1980</bibRefCitation>
: fig. 248) and
<bibRefCitation id="EFB84B6EFFC7FF9CFEB2F8824384DD6F" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[260,434,1801,1828]" pageId="10" pageNumber="499" pagination="130 - 142" refId="ref6107" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B95DEA8DFFC7FF9CFE9DF8824353DD6F" box="[299,357,1801,1827]" italics="true" pageId="10" pageNumber="499">et al.</emphasis>
(2016
</bibRefCitation>
: fig. 1G) respectively do not match the original illustrations of
<bibRefCitation id="EFB84B6EFFC7FF9CFB33F881473FDD68" author="Dyal, S." box="[1157,1289,1801,1828]" pageId="10" pageNumber="499" pagination="119 - 252" refId="ref5808" refString="Dyal, S. (1935) Fauna of Lahore. 4. -- Spiders of Lahore. Bulletin of the Department of Zoology of the Panjab University, 1, 119 - 252, pls. 11 - 17." type="journal article" year="1935">Dyal (1935</bibRefCitation>
: fig. 46), but clearly indicate that these authors misidentified specimens that actually belong to
<taxonomicName id="4C294D1CFFC7FF9CFB99F8A646A9DD0B" authorityName="Reimoser" authorityYear="1934" box="[1071,1183,1837,1864]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="499" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B95DEA8DFFC7FF9CFB99F8A646A9DD0B" box="[1071,1183,1837,1864]" italics="true" pageId="10" pageNumber="499">A. lesserti</emphasis>
</taxonomicName>
(compare
<figureCitation id="13122A1AFFC7FF9CFAAEF8A54753DD04" box="[1304,1381,1837,1864]" captionStart="FIGURE 10" captionStartId="12.[151,250,1220,1244]" captionTargetBox="[158,1427,246,1190]" captionTargetId="figure-50@12.[151,1436,237,1197]" captionTargetPageId="12" captionText="FIGURE 10. AB Arctosa lesserti Reimoser, 1934. A female lectotype epigynum, ventral. B female paralectotype epigynum, ventral. C Wadicosa ghatica Kronestedt, 2017, female paralectotype from A. lesserti type series, epigynum, ventral. Scale bars: 0.1 mm. © Christoph Hörweg, NHM, Vienna." figureDoi="http://doi.org/10.5281/zenodo.4447159" httpUri="https://zenodo.org/record/4447159/files/figure.png" pageId="10" pageNumber="499">Fig. 10</figureCitation>
AB to
<bibRefCitation id="EFB84B6EFFC7FF9CFF05F8D943E1DD20" author="Tikader, B. K. &amp; Malhotra, M. S." box="[179,471,1873,1900]" pageId="10" pageNumber="499" pagination="248 - 447" refId="ref6643" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India (Araneae), 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra 1980</bibRefCitation>
: fig. 248;
<bibRefCitation id="EFB84B6EFFC7FF9CFDFEF8DA40DFDD27" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." box="[584,745,1873,1900]" pageId="10" pageNumber="499" pagination="130 - 142" refId="ref6107" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B95DEA8DFFC7FF9CFDD8F8DA4090DD27" box="[622,678,1873,1899]" italics="true" pageId="10" pageNumber="499">et al.</emphasis>
2016
</bibRefCitation>
: fig. 1G).
</paragraph>
</subSubSection>
</treatment>
</document>