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<mods:title id="DAF73AFFCF278EBF43D7513C034C19EC">Revision of Indian wolf spiders: I. Genus Arctosa C. L. Koch, 1847 (Araneae Lycosidae, Tricassinae)</mods:title>
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<mods:namePart id="CB2C798E880D5A50897214FF1069F202">Caleb, John T. D.</mods:namePart>
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<mods:affiliation id="B299742AF28AA86051EFBAF314FEA038">Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India. caleb 87 woodgate @ gmail. com; https: // orcid. org / 0000 - 0002 - 9471 - 9467</mods:affiliation>
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<mods:namePart id="80ED86A35DCFE51D87E544F3DA2AD3F5">Sebastian, Pothalil A.</mods:namePart>
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<mods:affiliation id="C7182AECFBDC7BA670B9830EFAD03361">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India. &amp; drpothalil @ rediffmail. com; https: // orcid. org / 0000 - 0002 - 4936 - 4310</mods:affiliation>
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<paragraph id="8B96369FFFC0FF9BFF21FE94467BDB7B" blockId="13.[151,1101,284,347]" box="[151,1101,284,311]" pageId="13" pageNumber="502">
<heading id="D0DE81F3FFC0FF9BFF21FE94467BDB7B" bold="true" box="[151,1101,284,311]" fontSize="11" level="1" pageId="13" pageNumber="502" reason="1">
<emphasis id="B95DEA8DFFC0FF9BFF21FE94467BDB7B" bold="true" box="[151,1101,284,311]" pageId="13" pageNumber="502">
<taxonomicName id="4C294D1CFFC0FF9BFF21FE9441F6DB7A" ID-CoL="98Z92" authority="(Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012)" authorityName="Sankaran &amp; Caleb &amp; Sebastian" authorityYear="2021" baseAuthorityName="Dhali, Roy, Sen, Saha &amp; Raychaudhuri" baseAuthorityYear="2012" box="[151,960,284,311]" class="Arachnida" family="Lycosidae" genus="Ovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="species" species="quinquedens" status="comb. nov.">
<emphasis id="B95DEA8DFFC0FF9BFF21FE944345DB7B" bold="true" box="[151,371,284,311]" italics="true" pageId="13" pageNumber="502">Ovia quinquedens</emphasis>
(
<bibRefCitation id="EFB84B6EFFC0FF9BFE34FE944181DB7A" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[386,951,284,311]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012</bibRefCitation>
)
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<taxonomicNameLabel id="A26E57F6FFC0FF9BFC71FE95467BDB7B" box="[967,1101,285,311]" pageId="13" pageNumber="502" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C3336514FFC0FF9BFF21FEC842D0DB17" box="[151,230,320,347]" pageId="13" pageNumber="502" type="description">
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<figureCitation id="13122A1AFFC0FF9BFF21FEC842D0DB17" box="[151,230,320,347]" captionStart="FIGURE 11" captionStartId="13.[151,250,1979,2003]" captionTargetBox="[159,1426,1082,1950]" captionTargetId="figure-317@13.[156,1432,1074,1954]" captionTargetPageId="13" captionText="FIGURE 11. Ovia quinquedens (Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012) comb. nov. A female holotype epigynum, ventral. B same, ventral. Figures not to scale and 11A used with the permission of Dr Dhruba Chandra Dhali." figureDoi="http://doi.org/10.5281/zenodo.4447161" httpUri="https://zenodo.org/record/4447161/files/figure.png" pageId="13" pageNumber="502">Fig. 11</figureCitation>
</paragraph>
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<treatmentCitation id="0A88108EFFC0FF9BFF21FE084053DBD4" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[151,613,384,408]" page="1200" pageId="13" pageNumber="502" year="2012">
<taxonomicName id="4C294D1CFFC0FF9BFF21FE084053DBD4" ID-CoL="GBHT" authority="Dhali et al., 2012: 1200" authorityName="Dhali et al., 2012:" authorityPageNumber="1200" authorityYear="1200" box="[151,613,384,408]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B95DEA8DFFC0FF9BFF21FE08435DDBD4" box="[151,363,384,408]" italics="true" pageId="13" pageNumber="502">Arctosa quinquedens</emphasis>
<bibRefCitation id="EFB84B6EFFC0FF9BFEC4FE084053DBD4" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[370,613,384,408]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFE06FE0843D2DBD4" box="[432,484,384,408]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
, 2012: 1200
</bibRefCitation>
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, figs 16, 18.
<treatmentCitation id="0A88108EFFC0FF9BFD42FE0841FEDBD4" author="Dhali, D. C. &amp; Saha, S. &amp; Raychaudhuri, D." box="[756,968,384,408]" page="78" pageId="13" pageNumber="502" year="2017">
<bibRefCitation id="EFB84B6EFFC0FF9BFD42FE0841FEDBD4" author="Dhali, D. C. &amp; Saha, S. &amp; Raychaudhuri, D." box="[756,968,384,408]" pageId="13" pageNumber="502" pagination="1 - 242" refId="ref5648" refString="Dhali, D. C., Saha, S. &amp; Raychaudhuri, D. (2017) Litter and ground dwelling spiders (Araneae: Arachnida) of reserve forests of Dooars, West Bengal. World Scientific News, 63, 1 - 242." type="journal article" year="2017">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFC85FE084150DBD4" box="[819,870,384,408]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
2017: 78
</bibRefCitation>
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, plate XXIII, figs 394398.
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</paragraph>
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<subSubSection id="C3336514FFC0FF9BFF21FE494351D86E" pageId="13" pageNumber="502" type="materials_examined">
<paragraph id="8B96369FFFC0FF9BFF21FE494351D86E" blockId="13.[151,1437,448,1051]" pageId="13" pageNumber="502">
<emphasis id="B95DEA8DFFC0FF9BFF21FE49437CDB96" bold="true" box="[151,330,448,475]" pageId="13" pageNumber="502">Type material.</emphasis>
<materialsCitation id="3B413CC2FFC0FF9BFEE0FE484355D86E" ID-GBIF-Occurrence="3023039312" collectingDate="2009-04-09" collectorName="D. C. Dhali" country="India" county="Female" elevation="81" latitude="26.615997" location="Buxa Tiger Reserve" longLatPrecision="1" longitude="89.532394" municipality="Jalpaiguri" pageId="13" pageNumber="502" specimenCount="1" stateProvince="West Bengal" typeStatus="holotype">
<collectingCounty id="62F74E13FFC0FF9BFEE0FE48439EDB96" box="[342,424,448,474]" pageId="13" pageNumber="502">Female</collectingCounty>
<typeStatus id="5492883DFFC0FF9BFE03FE484021DB96" box="[437,535,448,474]" pageId="13" pageNumber="502" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F33E760FFFC0FF9BFDD0FE49408CDB97" box="[614,698,449,475]" name="India" pageId="13" pageNumber="502">INDIA</collectingCountry>
:
<collectingRegion id="49EDF87DFFC0FF9BFD7FFE494157DB96" box="[713,865,448,475]" country="India" name="West Bengal" pageId="13" pageNumber="502">West Bengal</collectingRegion>
:
<collectingMunicipality id="6BF2ACE5FFC0FF9BFCC6FE4841D5DB96" box="[880,995,448,474]" pageId="13" pageNumber="502">Jalpaiguri</collectingMunicipality>
:
<location id="8EF66044FFC0FF9BFC45FE4946D6DB97" LSID="urn:lsid:plazi:treatment:03808789FFC0FF9BFF21FE944145DE56:8EF66044FFC0FF9BFC45FE4946D6DB97" box="[1011,1248,448,475]" country="India" county="Female" latitude="26.615997" longLatPrecision="1" longitude="89.532394" municipality="Jalpaiguri" name="Buxa Tiger Reserve" pageId="13" pageNumber="502" stateProvince="West Bengal">Buxa Tiger Reserve</location>
:
<location id="8EF66044FFC0FF9BFB46FE4847ABDB96" LSID="urn:lsid:plazi:treatment:03808789FFC0FF9BFF21FE944145DE56:8EF66044FFC0FF9BFB46FE4847ABDB96" box="[1264,1437,448,474]" country="India" county="Female" latitude="26.615997" longLatPrecision="1" longitude="89.532394" municipality="Jalpaiguri" name="Rajabhatkhawa" pageId="13" pageNumber="502" stateProvince="West Bengal">Rajabhatkhawa</location>
(
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,
<geoCoordinate id="EE1D5058FFC0FF9BFED6FE6C4026DBB3" box="[352,528,484,511]" degrees="89" direction="east" minutes="31" orientation="longitude" pageId="13" pageNumber="502" precision="1" seconds="56.62" value="89.532394">89°3156.62E</geoCoordinate>
),
<quantity id="4CD19B7AFFC0FF9BFD90FE6D4056DBB3" box="[550,608,485,511]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="502" unit="m" value="81.0">
<elevation id="0004D1ACFFC0FF9BFD90FE6D4056DBB3" box="[550,608,485,511]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="502" unit="m" value="81.0">81 m</elevation>
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alt.;
<collectorName id="26DC5349FFC0FF9BFD2AFE6D4121DBB2" box="[668,791,484,511]" pageId="13" pageNumber="502">D.C. Dhali</collectorName>
leg.;
<date id="FF97105FFFC0FF9BFCEFFE6D41DADBB3" box="[857,1004,484,511]" pageId="13" pageNumber="502" value="2009-04-09">
<collectingDate id="EFD3E9B7FFC0FF9BFCEFFE6D41DADBB3" box="[857,1004,484,511]" pageId="13" pageNumber="502" value="2009-04-09">9 April 2009</collectingDate>
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; repository EZC (0025-12), examined from photographs
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.
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<subSubSection id="C3336514FFC0FF9BFF71FDA44625D8C3" pageId="13" pageNumber="502" type="diagnosis">
<paragraph id="8B96369FFFC0FF9BFF71FDA44625D8C3" blockId="13.[151,1437,448,1051]" pageId="13" pageNumber="502">
<emphasis id="B95DEA8DFFC0FF9BFF71FDA44374D80A" bold="true" box="[199,322,556,582]" pageId="13" pageNumber="502">Diagnosis.</emphasis>
Females of
<taxonomicName id="4C294D1CFFC0FF9BFE7EFDA54041D80A" authorityName="Dhali et al., 2012:" authorityYear="1200" box="[456,631,556,583]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="B95DEA8DFFC0FF9BFE7EFDA54041D80A" box="[456,631,556,583]" italics="true" pageId="13" pageNumber="502">O. quinquedens</emphasis>
</taxonomicName>
closely resemble the females of
<taxonomicName id="4C294D1CFFC0FF9BFC69FDA5476ED80B" authority="(Yu &amp; Song, 1988)" baseAuthorityName="Yu &amp; Song" baseAuthorityYear="1988" box="[991,1368,557,583]" class="Arachnida" family="Lycosidae" genus="Ovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="species" species="procurva">
<emphasis id="B95DEA8DFFC0FF9BFC69FDA546B7D80A" box="[991,1153,557,583]" italics="true" pageId="13" pageNumber="502">Ovia procurva</emphasis>
(Yu &amp; Song, 1988)
</taxonomicName>
in the general appearance of the epigynum, but can be easily distinguished from the latter by large epigynal atria, which are small in
<taxonomicName id="4C294D1CFFC0FF9BFE97FDFD439FD8C2" baseAuthorityName="Yu &amp; Song" baseAuthorityYear="1988" box="[289,425,629,655]" class="Arachnida" family="Lycosidae" genus="Ovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="species" species="procurva">
<emphasis id="B95DEA8DFFC0FF9BFE97FDFD439FD8C2" box="[289,425,629,655]" italics="true" pageId="13" pageNumber="502">O. procurva</emphasis>
</taxonomicName>
(compare
<figureCitation id="13122A1AFFC0FF9BFD96FDFC405AD8C3" box="[544,620,628,655]" captionStart="FIGURE 11" captionStartId="13.[151,250,1979,2003]" captionTargetBox="[159,1426,1082,1950]" captionTargetId="figure-317@13.[156,1432,1074,1954]" captionTargetPageId="13" captionText="FIGURE 11. Ovia quinquedens (Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012) comb. nov. A female holotype epigynum, ventral. B same, ventral. Figures not to scale and 11A used with the permission of Dr Dhruba Chandra Dhali." figureDoi="http://doi.org/10.5281/zenodo.4447161" httpUri="https://zenodo.org/record/4447161/files/figure.png" pageId="13" pageNumber="502">Fig. 11</figureCitation>
AB to
<bibRefCitation id="EFB84B6EFFC0FF9BFD71FDFC4187D8C3" author="Sankaran, P. M. &amp; Malamel, J. J. &amp; Sebastian, P. A." box="[711,945,628,655]" pageId="13" pageNumber="502" pagination="366 - 376" refId="ref6363" refString="Sankaran, P. M., Malamel, J. J. &amp; Sebastian, P. A. (2017) On the new monotypic wolf spider genus Ovia gen. nov. (Araneae: Lycosidae, Lycosinae). Zootaxa, 4221 (3), 366 - 376. https: // doi. org / 10.11646 / zootaxa. 4221.3.5" type="journal article" year="2017">
Sankaran
<emphasis id="B95DEA8DFFC0FF9BFC81FDFD4159D8C2" box="[823,879,628,654]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
2017
</bibRefCitation>
: fig. 5J).
</paragraph>
</subSubSection>
<subSubSection id="C3336514FFC0FF9BFF71FD104145DE56" pageId="13" pageNumber="502" type="description">
<paragraph id="8B96369FFFC0FF9BFF71FD1040DFD8FF" blockId="13.[151,1437,448,1051]" box="[199,745,664,691]" pageId="13" pageNumber="502">
<emphasis id="B95DEA8DFFC0FF9BFF71FD10436FD8FE" bold="true" box="[199,345,664,690]" pageId="13" pageNumber="502">Description.</emphasis>
For female, see
<bibRefCitation id="EFB84B6EFFC0FF9BFDA4FD1040D3D8FF" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[530,741,664,691]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFDEEFD1140A6D8FE" box="[600,656,664,690]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
(2012)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B96369FFFC0FF9BFF71FD3443B3D89A" blockId="13.[151,1437,448,1051]" box="[199,389,700,727]" pageId="13" pageNumber="502">
<emphasis id="B95DEA8DFFC0FF9BFF71FD344337D89A" box="[199,257,700,726]" italics="true" pageId="13" pageNumber="502">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B96369FFFC0FF9BFF71FD684145DE56" blockId="13.[151,1437,448,1051]" pageId="13" pageNumber="502">
<emphasis id="B95DEA8DFFC0FF9BFF71FD68402DD8B6" bold="true" box="[199,539,736,763]" pageId="13" pageNumber="502">Justification of the transfer.</emphasis>
<bibRefCitation id="EFB84B6EFFC0FF9BFD93FD6840CBD8B7" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[549,765,736,763]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFDDBFD69409ED8B6" box="[621,680,736,762]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
(2012)
</bibRefCitation>
described this species based on a single female specimen collected in
<collectingRegion id="49EDF87DFFC0FF9BFE96FC8D4384D952" box="[288,434,772,799]" country="India" name="West Bengal" pageId="13" pageNumber="502">West Bengal</collectingRegion>
. The original genitalia illustrations (
<bibRefCitation id="EFB84B6EFFC0FF9BFCFFFC8C463AD953" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[841,1036,772,799]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFC27FC8D41FFD952" box="[913,969,772,798]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
2012
</bibRefCitation>
: figs 56) of this species are highly schematic and do not facilitate identification. The paired anterior hoods of the epigynum (
<figureCitation id="13122A1AFFC0FF9BFBC9FCA046D4D90F" box="[1151,1250,808,835]" captionStart="FIGURE 11" captionStartId="13.[151,250,1979,2003]" captionTargetBox="[159,1426,1082,1950]" captionTargetId="figure-317@13.[156,1432,1074,1954]" captionTargetPageId="13" captionText="FIGURE 11. Ovia quinquedens (Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012) comb. nov. A female holotype epigynum, ventral. B same, ventral. Figures not to scale and 11A used with the permission of Dr Dhruba Chandra Dhali." figureDoi="http://doi.org/10.5281/zenodo.4447161" httpUri="https://zenodo.org/record/4447161/files/figure.png" pageId="13" pageNumber="502">Fig. 11B</figureCitation>
) clearly indicate that this species is misplaced in
<taxonomicName id="4C294D1CFFC0FF9BFDBFFCC54069D92B" authorityName="C. L. Koch" authorityYear="1847" box="[521,607,845,871]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8DFFC0FF9BFDBFFCC54069D92B" box="[521,607,845,871]" italics="true" pageId="13" pageNumber="502">Arctosa</emphasis>
</taxonomicName>
, as species of this genus lack anterior epigynal hoods and have well-developed epigynal median septum (
<bibRefCitation id="EFB84B6EFFC0FF9BFDA8FCF8410CD9C7" author="Dondale, C. D. &amp; Redner, J. H." box="[542,826,880,907]" pageId="13" pageNumber="502" pagination="1 - 30" refId="ref5757" refString="Dondale, C. D. &amp; Redner, J. H. (1983) Revision of the wolf spiders of the genus Arctosa C. L. Koch in North and Central America (Araneae: Lycosidae). Journal of Arachnology, 11, 1 - 30." type="journal article" year="1983">Dondale &amp; Redner 1983</bibRefCitation>
;
<bibRefCitation id="EFB84B6EFFC0FF9BFCF1FCF941CAD9C7" author="Yoo, J. S. &amp; Framenau, V. W. &amp; Kim, J. P." box="[839,1020,880,907]" pageId="13" pageNumber="502" pagination="171 - 180" refId="ref6958" refString="Yoo, J. S., Framenau, V. W. &amp; Kim, J. P. (2007) Arctosa stigmosa and A. subamylacea are two different species (Araneae, Lycosidae). Journal of Arachnology, 35, 171 - 180. https: // doi. org / 10.1636 / H 06 - 24.1" type="journal article" year="2007">
Yoo
<emphasis id="B95DEA8DFFC0FF9BFCCBFCF94181D9C6" box="[893,951,880,906]" italics="true" pageId="13" pageNumber="502">et al.</emphasis>
2007
</bibRefCitation>
). Instead, the general female habitus and colour pattern, the female epigynum with paired wide, longitudinal atria separated by a narrow median septum and the paired conspicuous anterior epigynal hoods (
<bibRefCitation id="EFB84B6EFFC0FF9BFD5DFC30419BD99F" author="Dhali, D. C. &amp; Roy, T. K. &amp; Sen, S. &amp; Saha, S. &amp; Raychaudhuri, D." box="[747,941,952,979]" pageId="13" pageNumber="502" pagination="1199 - 1213" refId="ref5696" refString="Dhali, D. C., Roy, T. K., Sen, S., Saha, S. &amp; Raychaudhuri, D. (2012) Wolf spiders (Araneae: Lycosidae) of the reserve forests of Dooars, West Bengal, India. Munis Entomology and Zoology, 7, 1199 - 1213." type="journal article" year="2012">
Dhali
<emphasis id="B95DEA8DFFC0FF9BFC87FC314154D99E" box="[817,866,952,978]" italics="true" pageId="13" pageNumber="502">et al</emphasis>
. 2012
</bibRefCitation>
: fig. 18;
<figureCitation id="13122A1AFFC0FF9BFBA7FC30466BD99F" box="[1041,1117,952,979]" captionStart="FIGURE 11" captionStartId="13.[151,250,1979,2003]" captionTargetBox="[159,1426,1082,1950]" captionTargetId="figure-317@13.[156,1432,1074,1954]" captionTargetPageId="13" captionText="FIGURE 11. Ovia quinquedens (Dhali, Roy, Sen, Saha &amp; Raychaudhuri, 2012) comb. nov. A female holotype epigynum, ventral. B same, ventral. Figures not to scale and 11A used with the permission of Dr Dhruba Chandra Dhali." figureDoi="http://doi.org/10.5281/zenodo.4447161" httpUri="https://zenodo.org/record/4447161/files/figure.png" pageId="13" pageNumber="502">Fig. 11</figureCitation>
A-B) indicate that it may be a member of
<taxonomicName id="4C294D1CFFC0FF9BFE98FC554117D9BB" authority="Sankaran, Malamel &amp; Sebastian, 2017" authorityName="Sankaran, Malamel &amp; Sebastian" authorityYear="2017" box="[302,801,988,1015]" class="Arachnida" family="Lycosidae" genus="Ovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8DFFC0FF9BFE98FC554352D9BB" box="[302,356,989,1015]" italics="true" pageId="13" pageNumber="502">Ovia</emphasis>
<bibRefCitation id="EFB84B6EFFC0FF9BFEDAFC544117D9BB" author="Sankaran, P. M. &amp; Malamel, J. J. &amp; Sebastian, P. A." box="[364,801,988,1015]" pageId="13" pageNumber="502" pagination="366 - 376" refId="ref6363" refString="Sankaran, P. M., Malamel, J. J. &amp; Sebastian, P. A. (2017) On the new monotypic wolf spider genus Ovia gen. nov. (Araneae: Lycosidae, Lycosinae). Zootaxa, 4221 (3), 366 - 376. https: // doi. org / 10.11646 / zootaxa. 4221.3.5" type="journal article" year="2017">Sankaran, Malamel &amp; Sebastian, 2017</bibRefCitation>
</taxonomicName>
. However, it will be confirmed only after examining the vulva of this species, which could not be examined for this study.
</paragraph>
</subSubSection>
</treatment>
</document>