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<document id="B970C17A89B91A16C4198DDC1F282F3F" ID-DOI="10.1080/00222933.2018.1432777" ID-ISSN="1464-5262" ID-Zenodo-Dep="5178275" ID-ZooBank="33BFE1A2-8C95-4A9D-9E7A-B495CEEC3182" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628622759912" checkinUser="felipe" docAuthor="Song, Sung Joon, Kangtia, Pawana, Khim, Jong Seong &amp; Chullasorn, Supawadee" docDate="2018" docId="03802D3EFFFEFFB8F0BC63F8FF17FAA3" docLanguage="en" docName="JNatHist.52.7-8.377-403.pdf" docOrigin="Journal of Natural History 52 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2018.1432777" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scottolana huysi Song &amp; Kangtia &amp; Khim &amp; Chullasorn 2018, sp. nov." docType="treatment" docUuid="7DD1F9D9-2E91-4EBD-BF86-F00BDE31426C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="399" masterDocId="FFB95546FFEEFFAFF1446632FFF8FF91" masterDocTitle="Two new Asian species of the genus Scottolana Huys, 2009 (Copepoda: Canuelloida: Canuellidae)" masterLastPageNumber="403" masterPageNumber="377" pageNumber="392" updateTime="1699048676939" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3B9AA35772F7581AE61C0100205AADF1">Two new Asian species of the genus Scottolana Huys, 2009 (Copepoda: Canuelloida: Canuellidae)</mods:title>
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<mods:namePart id="CD89CED406E35AC27C80CBF57A383805">Song, Sung Joon</mods:namePart>
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<mods:namePart id="CAE4EEC1B2C6E4FE28B064F7A48961A8">Kangtia, Pawana</mods:namePart>
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<mods:namePart id="AED7DD1C10C2C71A4FE2555B99F0859B">Khim, Jong Seong</mods:namePart>
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<mods:namePart id="B24963C674308D232EEE479E8D028511">Chullasorn, Supawadee</mods:namePart>
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<mods:title id="DBAD7184478450BCDCB64A8209C2A49A">Journal of Natural History</mods:title>
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<mods:date id="56AA115BA9861771FE2281899474DA72">2018</mods:date>
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<mods:number id="BBC5F8045F21DBF0C9934D315C15CA1E">2018-02-14</mods:number>
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<mods:number id="512F28EEBB5A1FAB0B9868C61E614B42">52</mods:number>
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<mods:number id="576DF2D8C1349D023BBA5460BB449BC6">7 - 8</mods:number>
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<treatment id="03802D3EFFFEFFB8F0BC63F8FF17FAA3" ID-DOI="http://doi.org/10.5281/zenodo.5189366" ID-Zenodo-Dep="5189366" LSID="urn:lsid:zoobank.org:act:7DD1F9D9-2E91-4EBD-BF86-F00BDE31426C" httpUri="http://treatment.plazi.org/id/03802D3EFFFEFFB8F0BC63F8FF17FAA3" lastPageId="23" lastPageNumber="399" pageId="16" pageNumber="392">
<subSubSection id="C333CFA3FFFEFFBFF0BC63F8FCD4FA75" box="[504,812,1482,1508]" pageId="16" pageNumber="392" type="nomenclature">
<paragraph id="8B969C28FFFEFFBFF0BC63F8FCD4FA75" blockId="16.[504,812,1482,1542]" box="[504,812,1482,1508]" pageId="16" pageNumber="392">
<heading id="D0DE2B44FFFEFFBFF0BC63F8FCD4FA75" box="[504,812,1482,1508]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="392" reason="5">
<taxonomicName id="4C29E7ABFFFEFFBFF0BC63F8FD3DFA72" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[504,709,1482,1507]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="16" pageNumber="392" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFEFFBFF0BC63F8FD3DFA72" bold="true" box="[504,709,1482,1507]" italics="true" pageId="16" pageNumber="392">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFEFFBFF38B63F9FCD4FA75" box="[719,812,1483,1508]" pageId="16" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C333CFA3FFFEFFBFF37A63DFFD1EF997" box="[574,742,1517,1542]" pageId="16" pageNumber="392" type="description">
<paragraph id="8B969C28FFFEFFBFF37A63DFFD1EF997" blockId="16.[504,812,1482,1542]" box="[574,742,1517,1542]" pageId="16" pageNumber="392">
(
<figureCitation id="131280ADFFFEFFBFF30063DFFD27F997" box="[580,735,1517,1542]" captionStart-0="Figure 9" captionStart-1="Figure 10" captionStart-2="Figure 11" captionStart-3="Figure 12" captionStart-4="Figure 13" captionStart-5="Figure 14" captionStart-6="Figure 15" captionStart-7="Figure 16" captionStart-8="Figure 17" captionStart-9="Figure 18" captionStartId-0="13.[160,225,1478,1501]" captionStartId-1="14.[160,225,1353,1376]" captionStartId-2="15.[160,225,843,866]" captionStartId-3="15.[160,225,1742,1765]" captionStartId-4="16.[160,225,1000,1023]" captionStartId-5="17.[160,225,1055,1078]" captionStartId-6="18.[160,225,1300,1323]" captionStartId-7="19.[160,225,1321,1344]" captionStartId-8="20.[160,225,970,993]" captionStartId-9="21.[160,225,1517,1540]" captionTargetBox-0="[179,1135,408,1447]" captionTargetBox-1="[178,1134,146,1312]" captionTargetBox-2="[180,1138,142,807]" captionTargetBox-3="[179,1137,976,1711]" captionTargetBox-4="[183,1135,143,966]" captionTargetBox-5="[178,1137,147,1021]" captionTargetBox-6="[188,1134,142,1266]" captionTargetBox-7="[178,1135,143,1288]" captionTargetBox-8="[183,1132,142,935]" captionTargetBox-9="[178,1134,142,1486]" captionTargetId-0="figure-30@13.[178,1138,408,1447]" captionTargetId-1="figure-73@14.[178,1138,142,1322]" captionTargetId-2="figure-1@15.[178,1138,142,811]" captionTargetId-3="figure-38@15.[178,1138,975,1711]" captionTargetId-4="figure-194@16.[178,1138,142,969]" captionTargetId-5="figure-145@17.[178,1138,142,1024]" captionTargetId-6="figure-117@18.[178,1138,142,1269]" captionTargetId-7="figure-112@19.[178,1138,142,1290]" captionTargetId-8="figure-261@20.[178,1138,142,938]" captionTargetId-9="figure-54@21.[178,1138,142,1486]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="15" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionTargetPageId-7="19" captionTargetPageId-8="20" captionTargetPageId-9="21" captionText-0="Figure 9. Scottolana huysi sp. nov. Female: (a) habitus, dorsal; (b) habitus, lateral; (c) caudal seta V (scale: 20 μm)." captionText-1="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." captionText-2="Figure 11. Scottolana huysi sp. nov. Female: (a) rostrum, dorsal; (b) antennule; (c) antenna; (d) distal segment of antenna; (e) mandible, ventral; (f) armature of exopodal setae of mandible as shown) (scale: 160 μm)." captionText-3="Figure 12. Scottolana huysi sp. nov. Female: (a) maxillule; (b) maxilla; (c) maxilliped (scale: a, b = 63 μm, c = 42 μm)." captionText-4="Figure 13. Scottolana huysi sp. nov. Female: (a) P1, anterior; (b) P2, anterior (scale: 80 μm)." captionText-5="Figure 14. Scottolana huysi sp. nov. Female: (a) P3, anterior; (b) P4, anterior (scale: 80 μm)." captionText-6="Figure 15. Scottolana huysi sp. nov. Male: (a) habitus, dorsal; (b) habitus, lateral (scale: 200 μm)." captionText-7="Figure 16. Scottolana huysi sp. nov. Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm)." captionText-8="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." captionText-9="Figure 18. Scottolana huysi sp. nov. Male: P4, anterior (scale: 80 μm)." figureDoi-0="http://doi.org/10.5281/zenodo.5178293" figureDoi-1="http://doi.org/10.5281/zenodo.5178295" figureDoi-2="http://doi.org/10.5281/zenodo.5178297" figureDoi-3="http://doi.org/10.5281/zenodo.5178299" figureDoi-4="http://doi.org/10.5281/zenodo.5178301" figureDoi-5="http://doi.org/10.5281/zenodo.5178303" figureDoi-6="http://doi.org/10.5281/zenodo.5178305" figureDoi-7="http://doi.org/10.5281/zenodo.5178307" figureDoi-8="http://doi.org/10.5281/zenodo.5178309" figureDoi-9="http://doi.org/10.5281/zenodo.5178311" httpUri-0="https://zenodo.org/record/5178293/files/figure.png" httpUri-1="https://zenodo.org/record/5178295/files/figure.png" httpUri-2="https://zenodo.org/record/5178297/files/figure.png" httpUri-3="https://zenodo.org/record/5178299/files/figure.png" httpUri-4="https://zenodo.org/record/5178301/files/figure.png" httpUri-5="https://zenodo.org/record/5178303/files/figure.png" httpUri-6="https://zenodo.org/record/5178305/files/figure.png" httpUri-7="https://zenodo.org/record/5178307/files/figure.png" httpUri-8="https://zenodo.org/record/5178309/files/figure.png" httpUri-9="https://zenodo.org/record/5178311/files/figure.png" pageId="16" pageNumber="392">Figures 918</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333CFA3FFFEFFBEF1E46001FE9DFA8B" lastPageId="17" lastPageNumber="393" pageId="16" pageNumber="392" type="materials_examined">
<paragraph id="8B969C28FFFEFFBFF1E46001FECFF9DD" blockId="16.[160,1156,1587,1715]" box="[160,311,1587,1612]" pageId="16" pageNumber="392">
<emphasis id="B95D403AFFFEFFBFF1E46001FECFF9DD" bold="true" box="[160,311,1587,1612]" italics="true" pageId="16" pageNumber="392">Type locality</emphasis>
</paragraph>
<paragraph id="8B969C28FFFEFFBFF1E46064FB7CF9FF" blockId="16.[160,1156,1587,1715]" box="[160,1156,1622,1646]" pageId="16" pageNumber="392">
<materialsCitation id="3B419675FFFEFFBFF1E46064FB78F9FF" box="[160,1152,1622,1646]" country="Thailand" latitude="7.8191667" location="Kata Beach" longLatPrecision="21" longitude="98.297775" municipality="Island" pageId="16" pageNumber="392" specimenCount="1" stateProvince="Phuket">
<location id="8EF6CAF3FFFEFFBFF1E46064FEE5F9FF" LSID="urn:lsid:plazi:treatment:03802D3EFFFEFFB8F0BC63F8FF17FAA3:8EF6CAF3FFFEFFBFF1E46064FEE5F9FF" box="[160,285,1622,1646]" country="Thailand" latitude="7.8191667" longLatPrecision="21" longitude="98.297775" municipality="Island" name="Kata Beach" pageId="16" pageNumber="392" stateProvince="Phuket">Kata Beach</location>
is situated on west coast of
<collectingRegion id="49ED52CAFFFEFFBFF32C6064FD4EF9FF" box="[616,694,1622,1646]" country="Thailand" name="Phuket" pageId="16" pageNumber="392">Phuket</collectingRegion>
<collectingMunicipality id="6BF20652FFFEFFBFF3FB6064FCFCF9FF" box="[703,772,1622,1646]" pageId="16" pageNumber="392">Island</collectingMunicipality>
,
<collectingCountry id="F33EDCB8FFFEFFBFF24B6064FC97F9FF" box="[783,879,1622,1646]" name="Thailand" pageId="16" pageNumber="392">Thailand</collectingCountry>
(
<geoCoordinate id="EE1DFAEFFFFEFFBFF23A6064FC09F9FF" box="[894,1009,1622,1646]" degrees="7" direction="north" minutes="49" orientation="latitude" pageId="16" pageNumber="392" precision="15" seconds="09" value="7.8191667">7°4909″N</geoCoordinate>
,
<geoCoordinate id="EE1DFAEFFFFEFFBFF2B96064FB81F9FF" box="[1021,1145,1622,1646]" degrees="98" direction="east" minutes="17" orientation="longitude" pageId="16" pageNumber="392" precision="15" seconds="52" value="98.297775">98°1752″E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B969C28FFFEFFBFF1E4604BFB9DF922" blockId="16.[160,1156,1587,1715]" pageId="16" pageNumber="392">
The environmental data of the fine sandy beach during low-tide are as follows: dissolved oxygen
<quantity id="4CD131CDFFFEFFBFF1BF60A9FEB2F922" box="[251,330,1691,1715]" metricMagnitude="-6" metricUnit="kg" metricValue="7.4" pageId="16" pageNumber="392" unit="mg" value="7.4">7.4 mg</quantity>
l1, salinity 32 ppt, temperature 32°C, pH 7.64, organic matter 2.30%.
</paragraph>
<caption id="DF56CCA0FFFFFFBEF1E4622DFBCFFBA7" ID-DOI="http://doi.org/10.5281/zenodo.5178303" ID-Zenodo-Dep="5178303" box="[160,1079,1054,1078]" httpUri="https://zenodo.org/record/5178303/files/figure.png" pageId="17" pageNumber="393" startId="17.[160,225,1055,1078]" targetBox="[178,1137,147,1021]" targetPageId="17">
<paragraph id="8B969C28FFFFFFBEF1E4622DFBCFFBA7" blockId="17.[160,1079,1054,1078]" box="[160,1079,1054,1078]" pageId="17" pageNumber="393">
<emphasis id="B95D403AFFFFFFBEF1E4622DFEF1FBA7" bold="true" box="[160,265,1055,1078]" pageId="17" pageNumber="393">Figure 14.</emphasis>
<taxonomicName id="4C29E7ABFFFFFFBEF055622CFE48FBA4" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[273,432,1054,1077]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="17" pageNumber="393" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFFFFBEF055622CFE48FBA4" box="[273,432,1054,1077]" italics="true" pageId="17" pageNumber="393">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFFFFBEF0FD622DFDF0FBA7" box="[441,520,1055,1078]" pageId="17" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
Female: (a) P3, anterior; (b) P4, anterior (scale: 80 μm).
</paragraph>
</caption>
<paragraph id="8B969C28FFFFFFBEF1E46245FE3AFB01" blockId="17.[160,1156,1143,1306]" box="[160,450,1143,1168]" pageId="17" pageNumber="393">
<emphasis id="B95D403AFFFFFFBEF1E46245FE3AFB01" bold="true" box="[160,450,1143,1168]" italics="true" pageId="17" pageNumber="393">Type material examined</emphasis>
</paragraph>
<paragraph id="8B969C28FFFFFFBEF1E462A9FE9DFA8B" blockId="17.[160,1156,1143,1306]" pageId="17" pageNumber="393">
<materialsCitation id="3B419675FFFFFFBEF1E462A9FCE0FB22" box="[160,792,1179,1203]" collectionCode="USNM" pageId="17" pageNumber="393" specimenCode="USNM1136881" specimenCount="1" typeStatus="holotype">
<typeStatus id="5492228AFFFFFFBEF1E462A9FEFFFB22" box="[160,263,1179,1203]" pageId="17" pageNumber="393" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="DB8F3453FFFFFFBEF02F62A9FDEEFB22" box="[363,534,1179,1203]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="393" type="Museum">USNM1136881</specimenCode>
): dissected on 9 slides
</materialsCitation>
.
<materialsCitation id="3B419675FFFFFFBEF26762A9FE72FB44" collectionCode="USNM" pageId="17" pageNumber="393" specimenCode="USNM1136882" specimenCount="1" typeStatus="allotype">
<typeStatus id="5492228AFFFFFFBEF26762A9FC78FB22" box="[803,896,1179,1203]" pageId="17" pageNumber="393" type="allotype">Allotype</typeStatus>
male (
<specimenCode id="DB8F3453FFFFFFBEF28B62A9FB83FB22" box="[975,1147,1179,1203]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="393" type="Museum">USNM1136882</specimenCode>
): dissected on 9 slides
</materialsCitation>
.
<materialsCitation id="3B419675FFFFFFBEF0D0628FFEEAFB69" collectionCode="USNM" pageId="17" pageNumber="393" specimenCode="USNM1136883" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="5492228AFFFFFFBEF0D0628FFDF9FB44" box="[404,513,1213,1237]" pageId="17" pageNumber="393" type="paratype">Paratypes</typeStatus>
(
<specimenCode id="DB8F3453FFFFFFBEF349628FFD41FB44" box="[525,697,1213,1237]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="393" type="Museum">USNM1136883</specimenCode>
):
<specimenCount id="9D2F57A1FFFFFFBEF38D628FFC99FB44" box="[713,865,1213,1237]" pageId="17" pageNumber="393" type="female">three females</specimenCount>
and
<specimenCount id="9D2F57A1FFFFFFBEF2DE628FFBF5FB44" box="[922,1037,1213,1237]" pageId="17" pageNumber="393" type="male">two males</specimenCount>
preserved in alcohol
</materialsCitation>
. All specimens are from the type locality collected by Supawadee Chullasorn on
<date id="FF97BAE8FFFFFFBEF1826330FE99FA8B" box="[198,353,1282,1306]" pageId="17" pageNumber="393" value="1999-04-20">20 April 1999</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C333CFA3FFFFFFBEF1E46375FEFAFA59" pageId="17" pageNumber="393" type="etymology">
<paragraph id="8B969C28FFFFFFBEF1E46375FEE5FAF1" blockId="17.[160,1156,1351,1480]" box="[160,285,1351,1376]" pageId="17" pageNumber="393">
<emphasis id="B95D403AFFFFFFBEF1E46375FEE5FAF1" bold="true" box="[160,285,1351,1376]" italics="true" pageId="17" pageNumber="393">Etymology</emphasis>
</paragraph>
<paragraph id="8B969C28FFFFFFBEF1E46358FEFAFA59" blockId="17.[160,1156,1351,1480]" pageId="17" pageNumber="393">The species was named in honour of Dr Rony Huys, Natural History Museum, London, for his achievements in the study of copepods. The name is a noun in the genitive singular.</paragraph>
</subSubSection>
<subSubSection id="C333CFA3FFFFFFB8F1E463C6FF17FAA3" lastPageId="23" lastPageNumber="399" pageId="17" pageNumber="393" type="description">
<paragraph id="8B969C28FFFFFFBEF1E463C6FD92F99C" blockId="17.[160,1156,1524,1723]" box="[160,618,1524,1549]" pageId="17" pageNumber="393">
<emphasis id="B95D403AFFFFFFBEF1E463C6FD92F99C" bold="true" box="[160,618,1524,1549]" italics="true" pageId="17" pageNumber="393">
Description of adult female (
<typeStatus id="5492228AFFFFFFBEF0BE63C6FD98F99C" box="[506,608,1524,1549]" pageId="17" pageNumber="393" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="8B969C28FFFFFFBEF1E4602AFC5FF92A" blockId="17.[160,1156,1524,1723]" pageId="17" pageNumber="393">
Habitus (
<figureCitation id="131280ADFFFFFFBEF041602AFE9AF9A1" box="[261,354,1560,1584]" captionStart="Figure 9" captionStartId="13.[160,225,1478,1501]" captionTargetBox="[179,1135,408,1447]" captionTargetId="figure-30@13.[178,1138,408,1447]" captionTargetPageId="13" captionText="Figure 9. Scottolana huysi sp. nov. Female: (a) habitus, dorsal; (b) habitus, lateral; (c) caudal seta V (scale: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178293" httpUri="https://zenodo.org/record/5178293/files/figure.png" pageId="17" pageNumber="393">Figure 9</figureCitation>
(a), 9(b)). Total body length
<quantity id="4CD131CDFFFFFFBEF3DE602AFD04F9A1" box="[666,764,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="17" pageNumber="393" unit="mm" value="1.32">1.32 mm</quantity>
measured from anterior margin of rostrum to posterior margin of caudal ramus. Body large, robust, slightly fusiform. Greatest width at posterior border of second pedigerous somite. Body surface ornamented with long sensilla except for anal somite. All somites without distinct posterior hyaline frills, except for anal somite with smooth dorsal hyaline frill.
</paragraph>
<caption id="DF56CCA0FFFCFFBDF1E46326FB92FABA" ID-DOI="http://doi.org/10.5281/zenodo.5178305" ID-Zenodo-Dep="5178305" box="[160,1130,1300,1323]" httpUri="https://zenodo.org/record/5178305/files/figure.png" pageId="18" pageNumber="394" startId="18.[160,225,1300,1323]" targetBox="[188,1134,142,1266]" targetPageId="18">
<paragraph id="8B969C28FFFCFFBDF1E46326FB92FABA" blockId="18.[160,1130,1300,1323]" box="[160,1130,1300,1323]" pageId="18" pageNumber="394">
<emphasis id="B95D403AFFFCFFBDF1E46326FEF1FABA" bold="true" box="[160,265,1300,1323]" pageId="18" pageNumber="394">Figure 15.</emphasis>
<taxonomicName id="4C29E7ABFFFCFFBDF0556326FE48FABA" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[273,432,1300,1323]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="18" pageNumber="394" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFCFFBDF0556326FE48FABA" box="[273,432,1300,1323]" italics="true" pageId="18" pageNumber="394">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFCFFBDF0FD6326FDF0FABA" box="[441,520,1300,1323]" pageId="18" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
Male: (a) habitus, dorsal; (b) habitus, lateral (scale: 200 μm).
</paragraph>
</caption>
<paragraph id="8B969C28FFFCFFBDF184635AFE05F9C1" blockId="18.[160,1156,1384,1720]" pageId="18" pageNumber="394">
Urosome (
<figureCitation id="131280ADFFFCFFBDF072635AFE5AFA11" box="[310,418,1384,1408]" captionStart="Figure 10" captionStartId="14.[160,225,1353,1376]" captionTargetBox="[178,1134,146,1312]" captionTargetId="figure-73@14.[178,1138,142,1322]" captionTargetPageId="14" captionText="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178295" httpUri="https://zenodo.org/record/5178295/files/figure.png" pageId="18" pageNumber="394">Figure 10</figureCitation>
(a), 10(b)) 4-segmented, consisting of P-5 bearing somite, genital double-somite, one free urosomite and anal somite. Genital double-somite slightly wider than long; original segmentation marked by paired dorsolateral and ventrolateral internal cuticular ribs (
<figureCitation id="131280ADFFFCFFBDF02C63E2FE2AFA79" box="[360,466,1488,1512]" captionStart="Figure 10" captionStartId="14.[160,225,1353,1376]" captionTargetBox="[178,1134,146,1312]" captionTargetId="figure-73@14.[178,1138,142,1322]" captionTargetPageId="14" captionText="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178295" httpUri="https://zenodo.org/record/5178295/files/figure.png" pageId="18" pageNumber="394">Figure 10</figureCitation>
(b)). Genital field large, extending to about halfway down the ventral surface of genital double-somite. Copulatory pores paired, posteriorly displaced to inner margin of wing-like processes. Anal operculum (
<figureCitation id="131280ADFFFCFFBDF26C6027FC7CF9BC" box="[808,900,1557,1581]" captionStart="Figure 9" captionStartId="13.[160,225,1478,1501]" captionTargetBox="[179,1135,408,1447]" captionTargetId="figure-30@13.[178,1138,408,1447]" captionTargetPageId="13" captionText="Figure 9. Scottolana huysi sp. nov. Female: (a) habitus, dorsal; (b) habitus, lateral; (c) caudal seta V (scale: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178293" httpUri="https://zenodo.org/record/5178293/files/figure.png" pageId="18" pageNumber="394">Figure 9</figureCitation>
(a)) weakly developed, with transverse row of setules.
</paragraph>
<paragraph id="8B969C28FFFCFFBCF1846069FC22F9E4" blockId="18.[160,1156,1384,1720]" lastBlockId="19.[160,1156,1455,1722]" lastPageId="19" lastPageNumber="395" pageId="18" pageNumber="394">
Caudal rami (
<figureCitation id="131280ADFFFCFFBDF01C6069FE3DF9E2" box="[344,453,1627,1651]" captionStart="Figure 10" captionStartId="14.[160,225,1353,1376]" captionTargetBox="[178,1134,146,1312]" captionTargetId="figure-73@14.[178,1138,142,1322]" captionTargetPageId="14" captionText="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178295" httpUri="https://zenodo.org/record/5178295/files/figure.png" pageId="18" pageNumber="394">Figure 10</figureCitation>
(a), 10(b)) subrectangular, about twice as long as wide; dorsal and ventral surfaces scattered with minute spinules, including characteristic triangular area on ventral surface (
<figureCitation id="131280ADFFFCFFBDF0856092FDC8F929" box="[449,560,1696,1720]" captionStart="Figure 10" captionStartId="14.[160,225,1353,1376]" captionTargetBox="[178,1134,146,1312]" captionTargetId="figure-73@14.[178,1138,142,1322]" captionTargetPageId="14" captionText="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178295" httpUri="https://zenodo.org/record/5178295/files/figure.png" pageId="18" pageNumber="394">Figure 10</figureCitation>
(b)); outer margin and posterioventral surface with dense clusters of strong spinules of different lengths. Each ramus armed with 7 setae: seta I short, spiniform, pinnate, arising ventrally; seta II long, plumose, with inflated base, located halfway inner margin; seta III plumose, displaced dorsally, issuing in the middle of caudal ramus; distal margin of ramus with 3 setae; seta IV and V well developed, strongly spinulose; the latter about 2.5 times longer than the former; seta VI small; seta VII short and smooth, located on dorsal surface between setae II and VI.
</paragraph>
<caption id="DF56CCA0FFFDFFBCF1E4631BFC3FFACF" ID-DOI="http://doi.org/10.5281/zenodo.5178307" ID-Zenodo-Dep="5178307" httpUri="https://zenodo.org/record/5178307/files/figure.png" pageId="19" pageNumber="395" startId="19.[160,225,1321,1344]" targetBox="[178,1135,143,1288]" targetPageId="19">
<paragraph id="8B969C28FFFDFFBCF1E4631BFC3FFACF" blockId="19.[160,1156,1321,1374]" pageId="19" pageNumber="395">
<emphasis id="B95D403AFFFDFFBCF1E4631BFEF0FAD1" bold="true" box="[160,264,1321,1344]" pageId="19" pageNumber="395">Figure 16.</emphasis>
<taxonomicName id="4C29E7ABFFFDFFBCF055631BFE57FAD1" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[273,431,1321,1344]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="19" pageNumber="395" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFDFFBCF055631BFE57FAD1" box="[273,431,1321,1344]" italics="true" pageId="19" pageNumber="395">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFDFFBCF0F3631BFDFEFAD1" box="[439,518,1321,1344]" pageId="19" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm).
</paragraph>
</caption>
<paragraph id="8B969C28FFFDFFBCF184604DFD04F92B" blockId="19.[160,1156,1455,1722]" pageId="19" pageNumber="395">
Rostrum (
<figureCitation id="131280ADFFFDFFBCF069604DFE6CF906" box="[301,404,1663,1687]" captionStart="Figure 11" captionStartId="15.[160,225,843,866]" captionTargetBox="[180,1138,142,807]" captionTargetId="figure-1@15.[178,1138,142,811]" captionTargetPageId="15" captionText="Figure 11. Scottolana huysi sp. nov. Female: (a) rostrum, dorsal; (b) antennule; (c) antenna; (d) distal segment of antenna; (e) mandible, ventral; (f) armature of exopodal setae of mandible as shown) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178297" httpUri="https://zenodo.org/record/5178297/files/figure.png" pageId="19" pageNumber="395">Figure 11</figureCitation>
(a)) very large, bell-shaped, about three-quarters the length of the cephalic shield, with a pair of subapical, tiny sensilla.
</paragraph>
<caption id="DF56CCA0FFFAFFBBF1E465F8FB7CFC70" ID-DOI="http://doi.org/10.5281/zenodo.5178309" ID-Zenodo-Dep="5178309" box="[160,1156,969,993]" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="20" pageNumber="396" startId="20.[160,225,970,993]" targetBox="[183,1132,142,935]" targetPageId="20">
<paragraph id="8B969C28FFFAFFBBF1E465F8FB7CFC70" blockId="20.[160,1156,969,993]" box="[160,1156,969,993]" pageId="20" pageNumber="396">
<emphasis id="B95D403AFFFAFFBBF1E465F8FEFFFC70" bold="true" box="[160,263,970,993]" pageId="20" pageNumber="396">Figure 17.</emphasis>
<taxonomicName id="4C29E7ABFFFAFFBBF05465FBFE5DFC71" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[272,421,969,992]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="20" pageNumber="396" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFAFFBBF05465FBFE5DFC71" box="[272,421,969,992]" italics="true" pageId="20" pageNumber="396">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFAFFBBF0EF65F8FE0CFC70" box="[427,500,970,993]" pageId="20" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm).
</paragraph>
</caption>
<paragraph id="8B969C28FFFAFFBBF1846220FD8CFB46" blockId="20.[160,1156,1042,1724]" pageId="20" pageNumber="396">
Antennule (
<figureCitation id="131280ADFFFAFFBBF0016220FE55FBBB" box="[325,429,1042,1066]" captionStart="Figure 11" captionStartId="15.[160,225,843,866]" captionTargetBox="[180,1138,142,807]" captionTargetId="figure-1@15.[178,1138,142,811]" captionTargetPageId="15" captionText="Figure 11. Scottolana huysi sp. nov. Female: (a) rostrum, dorsal; (b) antennule; (c) antenna; (d) distal segment of antenna; (e) mandible, ventral; (f) armature of exopodal setae of mandible as shown) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178297" httpUri="https://zenodo.org/record/5178297/files/figure.png" pageId="20" pageNumber="396">Figure 11</figureCitation>
(a)) indistinctly 4-segmented. First segment longest and largest, showing 3 incomplete sutures around the posterior margin with 9 spinulose and 4 smooth setae; second segment short, with 1 smooth and 10 spinulose setae, and 2 aesthetascs of unequal length; third segment very short, with 1 spinulose and 2 smooth setae; distal segment with 5 spinulose, 2 plumose, and 6 smooth setae. Armature formula: 1-(13), 2-(11 + 2ae), 3-(3), 4-(13).
</paragraph>
<paragraph id="8B969C28FFFAFFBBF18462D3FEAFFA58" blockId="20.[160,1156,1042,1724]" pageId="20" pageNumber="396">
Antenna (
<figureCitation id="131280ADFFFAFFBBF06A62D3FE6EFB68" box="[302,406,1249,1273]" captionStart="Figure 11" captionStartId="15.[160,225,843,866]" captionTargetBox="[180,1138,142,807]" captionTargetId="figure-1@15.[178,1138,142,811]" captionTargetPageId="15" captionText="Figure 11. Scottolana huysi sp. nov. Female: (a) rostrum, dorsal; (b) antennule; (c) antenna; (d) distal segment of antenna; (e) mandible, ventral; (f) armature of exopodal setae of mandible as shown) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178297" httpUri="https://zenodo.org/record/5178297/files/figure.png" pageId="20" pageNumber="396">Figure 11</figureCitation>
(b)) biramous; comprising a relatively short coxo-basis, with setular patches as shown. Endopod 2-segmented; enp-1 with 1 minute and smooth seta, and 1 long and pinnate seta; enp-2 with incomplete suture showing original segmentation between second and third segments, with 4 lateral pinnate setae, and 7 apical setae, 6 of them pinnate, 1 short and smooth. Exopod 9-segmented; segments 13 and 78 with 1 plumose seta each; segments 46 with 1 pinnate seta each; terminal segment small with 3 pinnate setae.
</paragraph>
<paragraph id="8B969C28FFFAFFBBF18463E6FC79F9E6" blockId="20.[160,1156,1042,1724]" pageId="20" pageNumber="396">
Mandible (
<figureCitation id="131280ADFFFAFFBBF07E63E6FE5CFA7D" box="[314,420,1492,1516]" captionStart="Figure 11" captionStartId="15.[160,225,843,866]" captionTargetBox="[180,1138,142,807]" captionTargetId="figure-1@15.[178,1138,142,811]" captionTargetPageId="15" captionText="Figure 11. Scottolana huysi sp. nov. Female: (a) rostrum, dorsal; (b) antennule; (c) antenna; (d) distal segment of antenna; (e) mandible, ventral; (f) armature of exopodal setae of mandible as shown) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178297" httpUri="https://zenodo.org/record/5178297/files/figure.png" pageId="20" pageNumber="396">Figure 11</figureCitation>
(c)) with well-developed gnathobase bearing 6 strong multicuspidate teeth distally; with additional row of spinules at dorsal margin. Basis with 2 long setae and inner spinular row. Endopod 2-segmented; enp-1 with 1 short and 2 long setae; enp-2 with 2 subapical and 6 apical setae. Exopod 4-segmented; all setae plumose; segments 1 and 4 with 2, segments 23 with 1 seta each.
</paragraph>
<paragraph id="8B969C28FFFAFFBAF18460B0FBC7F92C" blockId="20.[160,1156,1042,1724]" lastBlockId="21.[160,1155,1597,1725]" lastPageId="21" lastPageNumber="397" pageId="20" pageNumber="396">
Maxillule (
<figureCitation id="131280ADFFFAFFBBF07C60B0FE5EF90B" box="[312,422,1666,1690]" captionStart="Figure 12" captionStartId="15.[160,225,1742,1765]" captionTargetBox="[179,1137,976,1711]" captionTargetId="figure-38@15.[178,1138,975,1711]" captionTargetPageId="15" captionText="Figure 12. Scottolana huysi sp. nov. Female: (a) maxillule; (b) maxilla; (c) maxilliped (scale: a, b = 63 μm, c = 42 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178299" httpUri="https://zenodo.org/record/5178299/files/figure.png" pageId="20" pageNumber="396">Figure 12</figureCitation>
(a)). Praecoxa and coxa partly fused. Praecoxal arthrite with 8 spines, 2 pinnate and 2 smooth setae distally; posterior surface with 2 rows of spinules. Coxa with epipodite represented by 2 plumose setae; endite cylindrical, with 3 pinnate and 2 plumose setae. Basis with 6 pinnate setae and posterior row of spinules. Endopod 2-segmented; enp-1 with 5 pinnate setae laterally; enp-2 with 3 outer plumose and 3 inner pinnate setae. Exopod rounded, 1-segmented, with 11 long plumose setae.
</paragraph>
<caption id="DF56CCA0FFFBFFBAF1E463DFFCA1F995" ID-DOI="http://doi.org/10.5281/zenodo.5178311" ID-Zenodo-Dep="5178311" box="[160,857,1516,1540]" httpUri="https://zenodo.org/record/5178311/files/figure.png" pageId="21" pageNumber="397" startId="21.[160,225,1517,1540]" targetBox="[178,1134,142,1486]" targetPageId="21">
<paragraph id="8B969C28FFFBFFBAF1E463DFFCA1F995" blockId="21.[160,857,1516,1540]" box="[160,857,1516,1540]" pageId="21" pageNumber="397">
<emphasis id="B95D403AFFFBFFBAF1E463DFFEF1F995" bold="true" box="[160,265,1517,1540]" pageId="21" pageNumber="397">Figure 18.</emphasis>
<taxonomicName id="4C29E7ABFFFBFFBAF05563DEFE48F992" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[273,432,1516,1539]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="21" pageNumber="397" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFFBFFBAF05563DEFE48F992" box="[273,432,1516,1539]" italics="true" pageId="21" pageNumber="397">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFFBFFBAF0FD63DFFDF0F995" box="[441,520,1517,1540]" pageId="21" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
Male: P4, anterior (scale: 80 μm).
</paragraph>
</caption>
<paragraph id="8B969C28FFF8FFB9F18466BCFDF0FEC5" blockId="22.[160,1156,142,1657]" pageId="22" pageNumber="398">
Maxilla (
<figureCitation id="131280ADFFF8FFB9F06566BCFE68FF37" box="[289,400,142,166]" captionStart="Figure 12" captionStartId="15.[160,225,1742,1765]" captionTargetBox="[179,1137,976,1711]" captionTargetId="figure-38@15.[178,1138,975,1711]" captionTargetPageId="15" captionText="Figure 12. Scottolana huysi sp. nov. Female: (a) maxillule; (b) maxilla; (c) maxilliped (scale: a, b = 63 μm, c = 42 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178299" httpUri="https://zenodo.org/record/5178299/files/figure.png" pageId="22" pageNumber="398">Figure 12</figureCitation>
(b)) comprising praecoxa, coxa, allobasis and 3-segmented endopod. Praecoxa with outer row of spinules; with 2 endites; proximal endite with 1 smooth and 4 pinnate setae; distal endite with 2 pinnate setae. Coxa with 2 cylindrical endites, with 3 pinnate setae each. Allobasis with 3 strong pinnate spines, 2 pinnate and 1 smooth seta. Endopod 3-segmented, all setae smooth; enp-1 and enp-2 with 3 setae each; enp-3 small, with 2 setae.
</paragraph>
<paragraph id="8B969C28FFF8FFB9F184676CFB7CFD90" blockId="22.[160,1156,142,1657]" pageId="22" pageNumber="398">
Maxilliped (
<figureCitation id="131280ADFFF8FFB9F000676CFE57FEE7" box="[324,431,350,374]" captionStart="Figure 12" captionStartId="15.[160,225,1742,1765]" captionTargetBox="[179,1137,976,1711]" captionTargetId="figure-38@15.[178,1138,975,1711]" captionTargetPageId="15" captionText="Figure 12. Scottolana huysi sp. nov. Female: (a) maxillule; (b) maxilla; (c) maxilliped (scale: a, b = 63 μm, c = 42 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178299" httpUri="https://zenodo.org/record/5178299/files/figure.png" pageId="22" pageNumber="398">Figure 12</figureCitation>
(c)) phyllopodial, 2-segmented, comprising undivided protopod and 1-segmented endopod; incomplete suture indicating boundary between syncoxa and basis. Medial margin of protopod with 12 setae/spines (1 praecoxal, 9 coxal, and 2 basal); with rows of spinules as illustrated; outer margin with row of long setules distally. Endopod 1-segmented, with 11 elements (7 inner pinnate, and 4 outer plumose setae).
</paragraph>
<paragraph id="8B969C28FFF8FFB9F184643EFCD4FCEF" blockId="22.[160,1156,142,1657]" pageId="22" pageNumber="398">
P1 (
<figureCitation id="131280ADFFF8FFB9F1A9643EFEA0FDB5" box="[237,344,524,548]" captionStart="Figure 13" captionStartId="16.[160,225,1000,1023]" captionTargetBox="[183,1135,143,966]" captionTargetId="figure-194@16.[178,1138,142,969]" captionTargetPageId="16" captionText="Figure 13. Scottolana huysi sp. nov. Female: (a) P1, anterior; (b) P2, anterior (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178301" httpUri="https://zenodo.org/record/5178301/files/figure.png" pageId="22" pageNumber="398">Figure 13</figureCitation>
(a)) smaller than other swimming legs. Coxa with inner row of setules, and 6 medial spinules; inner coxal seta spinulose and pinnate. Basis with 1 pinnate inner spine, 1 spinulose long outer seta, and row of setules along inner margin. Exopod 3-segmented; exp-1 with 1 bipinnate outer spine, with strong outer spinules and with minute spinules distally; exp-2 with 1 bipinnate outer spine and 1 plumose inner seta; exp-3 outwardly directed, armed with 3 bipinnate outer spines, and 1 long, spinulose spine at outer distal corner, 1 apical and 2 inner plumose setae. Endopod 3-segmented, slightly longer than exopod; enp-1 with 1 plumose inner seta, and long outer setules; enp-2 with 1 plumose inner seta and several inner setules proximally, with outer strong spinules; enp-3 with 2 bipinnate outer spines, 1 spine, 2 apical setae, and 1 inner element, with outer spinules and inner setules as shown.
</paragraph>
<paragraph id="8B969C28FFF8FFB9F18465BBFCA8FB02" blockId="22.[160,1156,142,1657]" pageId="22" pageNumber="398">
P2 (
<figureCitation id="131280ADFFF8FFB9F1A965BBFEA2FC30" box="[237,346,905,929]" captionStart="Figure 13" captionStartId="16.[160,225,1000,1023]" captionTargetBox="[183,1135,143,966]" captionTargetId="figure-194@16.[178,1138,142,969]" captionTargetPageId="16" captionText="Figure 13. Scottolana huysi sp. nov. Female: (a) P1, anterior; (b) P2, anterior (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178301" httpUri="https://zenodo.org/record/5178301/files/figure.png" pageId="22" pageNumber="398">Figure 13</figureCitation>
(b)). Praecoxa without ornamentation. Coxa with rows of spinules and setules as shown; inner coxal seta spinulose. Basis with 1 spinulose outer seta; inner margin with small blunt process. Exopod 3-segmented; exp-1 with bipinnate outer spine, with strong outer spinules; exp-2 with 1 serrate outer spine and 1 plumose inner seta; enp-3 with 4 spines and 3 plumose setae. Endopod 3-segmented; enp-1 smallest, with 1 plumose inner seta, anterior surface produced into long mucroniform process, reaching distal margin of enp-2; enp-2 with 1 plumose inner seta and spinular rows as depicted; enp-3 with 3 spines and 2 plumose setae.
</paragraph>
<paragraph id="8B969C28FFF8FFB9F18462ACFDE4FA17" blockId="22.[160,1156,142,1657]" pageId="22" pageNumber="398">
P3 (
<figureCitation id="131280ADFFF8FFB9F1A962ACFEA0FB27" box="[237,344,1182,1206]" captionStart="Figure 14" captionStartId="17.[160,225,1055,1078]" captionTargetBox="[178,1137,147,1021]" captionTargetId="figure-145@17.[178,1138,142,1024]" captionTargetPageId="17" captionText="Figure 14. Scottolana huysi sp. nov. Female: (a) P3, anterior; (b) P4, anterior (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178303" httpUri="https://zenodo.org/record/5178303/files/figure.png" pageId="22" pageNumber="398">Figure 14</figureCitation>
(a)). Praecoxa without ornamentation. Coxa with strong outer spinules; inner coxal seta spinulose. Basis with 1 spinulose outer seta. Exopod 3-segmented; exp-1 with 1 serrate outer spine and strong spinules as shown; exp-2 with 1 serrate outer spine, 1 spinulose inner seta and strong spinules; exp-3 with 4 spines and 1 plumose setae distally. Endopod 3-segmented, longer than exopod; enp-1 and enp-2 with 1 inner plumose seta and strong spinules as shown; enp-3 with 1 outer medial, 1 outer and 1 inner distal, and 1 inner element.
</paragraph>
<paragraph id="8B969C28FFF8FFB8F18463A3FE1DFE3C" blockId="22.[160,1156,142,1657]" lastBlockId="23.[160,1155,370,498]" lastPageId="23" lastPageNumber="399" pageId="22" pageNumber="398">
P4 (
<figureCitation id="131280ADFFF8FFB9F1AA63A3FEA5FA38" box="[238,349,1425,1449]" captionStart="Figure 14" captionStartId="17.[160,225,1055,1078]" captionTargetBox="[178,1137,147,1021]" captionTargetId="figure-145@17.[178,1138,142,1024]" captionTargetPageId="17" captionText="Figure 14. Scottolana huysi sp. nov. Female: (a) P3, anterior; (b) P4, anterior (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178303" httpUri="https://zenodo.org/record/5178303/files/figure.png" pageId="22" pageNumber="398">Figure 14</figureCitation>
(b)). Praecoxa (not shown) without ornamentation. Coxa with 1 inner spinulose seta and several outer spinules. Basis with 1 outer spinulose seta. Exopod 3- segmented; exp-1 with 1 outer pinnate seta and dense patch of spinules; exp-2 with 1 outer pinnate seta, 1 inner spinulose seta and dense patch of spinules; exp-3 with 1 outer pinnate seta, 2 apical spinulose spines, and 1 inner spinulose seta, with several spinules as shown. Endopod 3-segmented; enp-1 shortest, with 1 very long spinulose inner seta and some spinules distally; enp-2 with dense patch of spinules; enp-3 with 2 spines and 2 spinulose setae. Seta and spine formula of swimming legs of the new species as shown in
<tableCitation id="C6ABA993FFF9FFB8F0CF67A7FE1DFE3C" box="[395,485,405,429]" captionStart="Table 2" captionStartId="23.[160,216,142,165]" captionText="Table 2. Armature formula of swimming legs of Scottolana huysi sp. nov." pageId="23" pageNumber="399">Table 2.</tableCitation>
</paragraph>
<caption id="DF56CCA0FFF9FFB8F1E466BCFC7AFF34" box="[160,898,141,165]" pageId="23" pageNumber="399" startId="23.[160,216,142,165]">
<paragraph id="8B969C28FFF9FFB8F1E466BCFC7AFF34" blockId="23.[160,898,141,165]" box="[160,898,141,165]" pageId="23" pageNumber="399">
<emphasis id="B95D403AFFF9FFB8F1E466BCFF0AFF34" bold="true" box="[160,242,142,165]" pageId="23" pageNumber="399">Table 2.</emphasis>
Armature formula of swimming legs of
<taxonomicName id="4C29E7ABFFF9FFB8F3C866BFFCD3FF35" authority="Song &amp; Kangtia &amp; Khim &amp; Chullasorn, 2018" authorityName="Song &amp; Kangtia &amp; Khim &amp; Chullasorn" authorityYear="2018" box="[652,811,141,164]" class="Hexanauplia" family="Canuellidae" genus="Scottolana" kingdom="Animalia" order="Canuelloida" pageId="23" pageNumber="399" phylum="Arthropoda" rank="species" species="huysi" status="sp. nov.">
<emphasis id="B95D403AFFF9FFB8F3C866BFFCD3FF35" box="[652,811,141,164]" italics="true" pageId="23" pageNumber="399">Scottolana huysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFD41FFF9FFB8F27766BCFC7AFF34" box="[819,898,142,165]" pageId="23" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B969C28FFF9FFB8F31C6680FB8FFEBC" pageId="23" pageNumber="399">
<table id="F9296E88FFF90050F1EF6680FB81FEBC" box="[171,1145,178,301]" colsContinueFrom="13.[171,1145,178,301]" gridcols="3" gridrows="5" pageId="23" pageNumber="399">
<tr id="35199E6AFFF90050F1EF6680FB81FF54" box="[171,1145,178,197]" gridrow="0" pageId="23" pageNumber="399" rowspan-0="1">
<th id="76C8F716FFF90050F3166680FD63FF54" box="[594,667,178,197]" gridcol="1" gridrow="0" pageId="23" pageNumber="399">Exopod</th>
<th id="76C8F716FFF90050F56A6680FB81FF54" box="[1070,1145,178,197]" gridcol="2" gridrow="0" pageId="23" pageNumber="399">Endopod</th>
</tr>
<tr id="35199E6AFFF90050F1EF66E0FB81FF74" box="[171,1145,210,229]" gridrow="1" pageId="23" pageNumber="399">
<th id="76C8F716FFF90050F1EF66E0FF38FF74" box="[171,192,210,229]" gridcol="0" gridrow="1" pageId="23" pageNumber="399">P1</th>
<td id="76C8F716FFF90050F31666E0FD63FF74" box="[594,667,210,229]" gridcol="1" gridrow="1" pageId="23" pageNumber="399">0, 1, 223</td>
<td id="76C8F716FFF90050F56A66E0FB81FF74" box="[1070,1145,210,229]" gridcol="2" gridrow="1" pageId="23" pageNumber="399">1, 1, 123</td>
</tr>
<tr id="35199E6AFFF90050F1EF66D8FB81FF6C" box="[171,1145,234,253]" gridrow="2" pageId="23" pageNumber="399">
<th id="76C8F716FFF90050F1EF66D8FF38FF6C" box="[171,192,234,253]" gridcol="0" gridrow="2" pageId="23" pageNumber="399">P2</th>
<td id="76C8F716FFF90050F31666D8FD63FF6C" box="[594,667,234,253]" gridcol="1" gridrow="2" pageId="23" pageNumber="399">0, 1, 322</td>
<td id="76C8F716FFF90050F56A66D8FB81FF6C" box="[1070,1145,234,253]" gridcol="2" gridrow="2" pageId="23" pageNumber="399">1, 1, 221</td>
</tr>
<tr id="35199E6AFFF90050F1EF6730FB81FE84" box="[171,1145,258,277]" gridrow="3" pageId="23" pageNumber="399">
<th id="76C8F716FFF90050F1EF6730FF38FE84" box="[171,192,258,277]" gridcol="0" gridrow="3" pageId="23" pageNumber="399">P3</th>
<td id="76C8F716FFF90050F3166730FD63FE84" box="[594,667,258,277]" gridcol="1" gridrow="3" pageId="23" pageNumber="399">0, 1, 122</td>
<td id="76C8F716FFF90050F56A6730FB81FE84" box="[1070,1145,258,277]" gridcol="2" gridrow="3" pageId="23" pageNumber="399">1, 1, 121</td>
</tr>
<tr id="35199E6AFFF90050F1EF6728FB81FEBC" box="[171,1145,282,301]" gridrow="4" pageId="23" pageNumber="399">
<th id="76C8F716FFF90050F1EF6728FF38FEBC" box="[171,192,282,301]" gridcol="0" gridrow="4" pageId="23" pageNumber="399">P4</th>
<td id="76C8F716FFF90050F3166728FD63FEBC" box="[594,667,282,301]" gridcol="1" gridrow="4" pageId="23" pageNumber="399">0, 1, 121</td>
<td id="76C8F716FFF90050F56A6728FB81FEBC" box="[1070,1145,282,301]" gridcol="2" gridrow="4" pageId="23" pageNumber="399">1, 0, 121</td>
</tr>
</table>
</paragraph>
<paragraph id="8B969C28FFF9FFB8F1846785FC03FE63" blockId="23.[160,1155,370,498]" pageId="23" pageNumber="399">
P5 (
<figureCitation id="131280ADFFF9FFB8F1AF6785FEAEFE5E" box="[235,342,439,463]" captionStart="Figure 10" captionStartId="14.[160,225,1353,1376]" captionTargetBox="[178,1134,146,1312]" captionTargetId="figure-73@14.[178,1138,142,1322]" captionTargetPageId="14" captionText="Figure 10. Scottolana huysi sp. nov. Female: (a) urosome, dorsal; (b) urosome, ventral; (c) P5, anterior; (d) caudal ramus, ventral; (e) caudal ramus, dorsal (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178295" httpUri="https://zenodo.org/record/5178295/files/figure.png" pageId="23" pageNumber="399">Figure 10</figureCitation>
(b)). Vestigial, incorporated into somite; with 1 innermost smooth and 3 plumose setae; middle one on distal margin longest and with long setules.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F1E4642CFDB3FDA6" blockId="23.[160,1156,542,1330]" box="[160,587,542,567]" pageId="23" pageNumber="399">
<emphasis id="B95D403AFFF9FFB8F1E4642CFDB3FDA6" bold="true" box="[160,587,542,567]" italics="true" pageId="23" pageNumber="399">
Description of adult male (
<typeStatus id="5492228AFFF9FFB8F0A1642CFDBBFDA6" box="[485,579,542,567]" pageId="23" pageNumber="399" type="allotype">allotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="8B969C28FFF9FFB8F1E46470FB91FDEC" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
<typeStatus id="5492228AFFF9FFB8F1E46470FF04FDCB" box="[160,252,578,602]" pageId="23" pageNumber="399" type="allotype">Allotype</typeStatus>
(
<figureCitation id="131280ADFFF9FFB8F04D6470FE89FDCB" box="[265,369,578,602]" captionStart="Figure 15" captionStartId="18.[160,225,1300,1323]" captionTargetBox="[188,1134,142,1266]" captionTargetId="figure-117@18.[178,1138,142,1269]" captionTargetPageId="18" captionText="Figure 15. Scottolana huysi sp. nov. Male: (a) habitus, dorsal; (b) habitus, lateral (scale: 200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178305" httpUri="https://zenodo.org/record/5178305/files/figure.png" pageId="23" pageNumber="399">Figure 15</figureCitation>
(a), 15(b)) slightly smaller and slender than the female. Total body length
<quantity id="4CD131CDFFF9FFB8F1E46457FEFBFDEC" box="[160,259,613,637]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="23" pageNumber="399" unit="mm" value="1.22">1.22 mm</quantity>
. Sexual dimorphism in the antennule, P4, P5, genital somite and caudal ramus.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F18464B5FC8DFCBB" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
Urosome (
<figureCitation id="131280ADFFF9FFB8F07464B5FE6FFD0E" box="[304,407,647,671]" captionStart="Figure 16" captionStartId="19.[160,225,1321,1344]" captionTargetBox="[178,1135,143,1288]" captionTargetId="figure-112@19.[178,1138,142,1290]" captionTargetPageId="19" captionText="Figure 16. Scottolana huysi sp. nov. Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178307" httpUri="https://zenodo.org/record/5178307/files/figure.png" pageId="23" pageNumber="399">Figure 16</figureCitation>
(a), 16(b)) 5-segmented, comprising P5-bearing somite, genital somite 3rd and 4th urosomites, and anal somite. Genital somite (
<figureCitation id="131280ADFFF9FFB8F2646498FC6DFD53" box="[800,917,682,706]" captionStart="Figure 15" captionStartId="18.[160,225,1300,1323]" captionTargetBox="[188,1134,142,1266]" captionTargetId="figure-117@18.[178,1138,142,1269]" captionTargetPageId="18" captionText="Figure 15. Scottolana huysi sp. nov. Male: (a) habitus, dorsal; (b) habitus, lateral (scale: 200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178305" httpUri="https://zenodo.org/record/5178305/files/figure.png" pageId="23" pageNumber="399">Figures 15</figureCitation>
(a), 15(b), 16(a), 16(b)) largest, ornamented with several tiny spinular rows ventrally (
<figureCitation id="131280ADFFF9FFB8F22764FFFC37FD74" box="[867,975,717,741]" captionStart="Figure 16" captionStartId="19.[160,225,1321,1344]" captionTargetBox="[178,1135,143,1288]" captionTargetId="figure-112@19.[178,1138,142,1290]" captionTargetPageId="19" captionText="Figure 16. Scottolana huysi sp. nov. Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178307" httpUri="https://zenodo.org/record/5178307/files/figure.png" pageId="23" pageNumber="399">Figure 16</figureCitation>
(b)), with paired lateral, posteriorly directed spinous extensions covered by tiny spinules and with long setules distally. Third urosomite with a bundle of setules ventrally.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F1846507FF04FCFE" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
Caudal rami (
<figureCitation id="131280ADFFF9FFB8F0126507FE3AFCDC" box="[342,450,821,845]" captionStart="Figure 16" captionStartId="19.[160,225,1321,1344]" captionTargetBox="[178,1135,143,1288]" captionTargetId="figure-112@19.[178,1138,142,1290]" captionTargetPageId="19" captionText="Figure 16. Scottolana huysi sp. nov. Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178307" httpUri="https://zenodo.org/record/5178307/files/figure.png" pageId="23" pageNumber="399">Figure 16</figureCitation>
(a), 16(b)) as in female, except for seta II with normal base, not inflated.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F1846548FD8DFC03" blockId="23.[160,1156,542,1330]" box="[192,629,890,914]" pageId="23" pageNumber="399">
Rostrum (
<figureCitation id="131280ADFFF9FFB8F06A6548FE5FFC03" box="[302,423,890,914]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17a</figureCitation>
) as in the female.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F18465AFFC38FBF3" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
Antennule (
<figureCitation id="131280ADFFF9FFB8F00B65AFFE2AFC24" box="[335,466,925,949]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17b</figureCitation>
) short 5-segmented. First segment (
<figureCitation id="131280ADFFF9FFB8F2D865AFFBE4FC24" box="[924,1052,925,949]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17c</figureCitation>
) with 2 plumose and 1 smooth setae; second segment (
<figureCitation id="131280ADFFF9FFB8F39A658DFCA4FC46" box="[734,860,959,983]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17d</figureCitation>
) with 5 plumose and 11 pinnate setae, and 2 aesthetascs; third segment (
<figureCitation id="131280ADFFF9FFB8F24B65D0FC68FC6B" box="[783,912,994,1018]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17e</figureCitation>
) with 1 smooth, 1 plumose, 1 spinulose and 1 pinnate setae; fourth segment (
<figureCitation id="131280ADFFF9FFB8F2D96237FBE2FB8C" box="[925,1050,1029,1053]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">Figure 17f</figureCitation>
) with 2 plumose and 3 pinnate setae; distal segment (
<figureCitation id="131280ADFFF9FFB8F3A66215FC9CFBAE" box="[738,868,1063,1087]" captionStart="Figure 17" captionStartId="20.[160,225,970,993]" captionTargetBox="[183,1132,142,935]" captionTargetId="figure-261@20.[178,1138,142,938]" captionTargetPageId="20" captionText="Figure 17. Scottolana huysi sp.nov. Male: rostrum + antennule; antennule (disarticulated) (scale: 160 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178309" httpUri="https://zenodo.org/record/5178309/files/figure.png" pageId="23" pageNumber="399">
Figure
<quantity id="4CD131CDFFF9FFB8F27D6215FC9CFBAE" box="[825,868,1063,1087]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="23" pageNumber="399" unit="g" value="17.0">17g</quantity>
</figureCitation>
) with 3 smooth and 2 pinnate setae. Armature formula: 1-(3), 2-(16+2ae), 3-(4), 4-(5), 5-(5).
</paragraph>
<paragraph id="8B969C28FFF9FFB8F184625FFBCEFB14" blockId="23.[160,1156,542,1330]" box="[192,1078,1133,1157]" pageId="23" pageNumber="399">Antenna, mandible, maxillule, maxilla, maxilliped, P1, P2, and P3 as in female.</paragraph>
<paragraph id="8B969C28FFF9FFB8F18462BDFBDAFB7C" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
P4 (
<figureCitation id="131280ADFFF9FFB8F1A862BDFEAEFB36" box="[236,342,1167,1191]" captionStart="Figure 18" captionStartId="21.[160,225,1517,1540]" captionTargetBox="[178,1134,142,1486]" captionTargetId="figure-54@21.[178,1138,142,1486]" captionTargetPageId="21" captionText="Figure 18. Scottolana huysi sp. nov. Male: P4, anterior (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178311" httpUri="https://zenodo.org/record/5178311/files/figure.png" pageId="23" pageNumber="399">Figure 18</figureCitation>
) as in female, except for basis bearing 1 outer smooth seta, and with 4 long setules of unequal length close to joint with endopod, reaching beyond enp-1: from outer 1 pinnate, 1 smooth, 1 longest plumose and 1 short smooth setae.
</paragraph>
<paragraph id="8B969C28FFF9FFB8F18462C5FF17FAA3" blockId="23.[160,1156,542,1330]" pageId="23" pageNumber="399">
P5 (
<figureCitation id="131280ADFFF9FFB8F1A962C5FEA2FA9E" box="[237,346,1271,1295]" captionStart="Figure 16" captionStartId="19.[160,225,1321,1344]" captionTargetBox="[178,1135,143,1288]" captionTargetId="figure-112@19.[178,1138,142,1290]" captionTargetPageId="19" captionText="Figure 16. Scottolana huysi sp. nov. Male: (a) urosome, dorsal; (b) urosome, ventral; (c) caudal seta; (d) P5, ventral; (e) caudal ramus, dorsal; (f) caudal ramus, ventral (scale: 80 μm)." figureDoi="http://doi.org/10.5281/zenodo.5178307" httpUri="https://zenodo.org/record/5178307/files/figure.png" pageId="23" pageNumber="399">Figure 16</figureCitation>
(b)) with armature as in female, but the outermost basal plumose seta longer.
</paragraph>
</subSubSection>
</treatment>
</document>