treatments-xml/data/C2/2C/A7/C22CA730D4030C2A6F43FF7CB2E329DD.xml

<document id="F674F800D89529B15D9F97060510F2E0" ID-DOI="10.5281/zenodo.4525838" ID-ISSN="1638-9387" ID-Zenodo-Dep="4525838" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612879696179" checkinUser="felipe" docAuthor="McLAUGHLIN, Patsy A. &amp; Rahayu, Dwi Listyo" docDate="2008" docId="C22CA730D4030C2A6F43FF7CB2E329DD" docLanguage="en" docName="Zoosystema.30.4.899-916.pdf" docOrigin="Zoosystema 30 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Pteropagurus spina McLaughlin &amp; Rahayu 2006" docType="treatment" docVersion="5" lastPageNumber="909" masterDocId="3E15DF48D4070C206E57FFA4B3302F79" masterDocTitle="Pteropagurus and Catapagurus (Decapoda, Anomura, Paguridae): resource sharing or “ any port in a storm ”?" masterLastPageNumber="916" masterPageNumber="899" pageNumber="903" updateTime="1698869201481" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0652C798EA5B77BEE27CA5A63CD20599">Pteropagurus and Catapagurus (Decapoda, Anomura, Paguridae): resource sharing or “ any port in a storm ”?</mods:title>
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<mods:namePart id="34CBAEB09B9F95F6B5167A4CAC3F85C3">McLAUGHLIN, Patsy A.</mods:namePart>
<mods:affiliation id="4BA2709FC0404BE66F2AC28F099CF5C3">Shannon Point Marine Center, Western Washington University, 1900 Shannon Point Road, Anacortes, WA 98221 - 4042 (USA) hermit @ fidalgo. net</mods:affiliation>
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<mods:namePart id="8817423A441B5A16EFA034048248115A">Rahayu, Dwi Listyo</mods:namePart>
<mods:affiliation id="851EA5323F1C02B1BF1A902253908371">Research Center for Oceanography, Indonesian Institute of Sciences (LIPI), Jl. Pasir Putih 1, Ancol Timur, Jakarta 14430 (Indonesia) dwilistyo @ yahoo. com</mods:affiliation>
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<mods:title id="6F751190F6A96615CF6DFA3B9A2A4F58">Zoosystema</mods:title>
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<mods:date id="733D59F617E752695729CAF5D3BBBA3E">2008</mods:date>
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<taxonomicName id="8D856DA5D4030C246F43FF7CB12D2E6A" ID-CoL="0e31a2af-75da-42ca-898e-05f2ae02d775" authority="McLaughlin &amp; Rahayu, 2006" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="903" phylum="Arthropoda" rank="species" species="spina">
<emphasis id="78F1CA34D4030C246F43FF7CB2D32F8B" bold="true" box="[276,483,216,242]" italics="true" pageId="4" pageNumber="903">Pteropagurus spina</emphasis>
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(
<figureCitation id="D2BE0AA3D4030C246F0BFEB3B2A92E4B" box="[348,409,279,306]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="4" pageNumber="903">Fig. 1</figureCitation>
)
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<treatmentCitation id="CB243037D4030C246E26FEF5B1B22E11" ID-DOI="http://doi.org/10.5281/zenodo.6256853" ID-Zenodo-Dep="6256853" author="MCLAUGHLIN P. A. &amp; RAHAYU D. L." box="[113,642,336,360]" httpUri="http://treatment.plazi.org/id/03EF87F4CC6D954AFEC6FE03FC64A306" page="64" pageId="4" pageNumber="903" year="2006">
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<emphasis id="78F1CA34D4030C246E26FEF5B2152E11" box="[113,293,337,360]" italics="true" pageId="4" pageNumber="903">Pteropagurus spina</emphasis>
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, fig. 5. — 
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<bibRefCitation id="2E146BD7D4030C246E87FECFB2862EFA" author="MCLAUGHLIN P. A." box="[208,438,363,387]" pageId="4" pageNumber="903" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin 2007: 508</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="4A3A1626D4030C246E26FE3FB2822E9E" blockId="4.[112,647,410,487]" pageId="4" pageNumber="903">
<materialsCitation id="FAED1C7BD4030C246E26FE3FB2822E9E" collectingDate="1986-10-09" collectionCode="DW" country="New Caledonia" elevation="285" latitude="-24.746666" location="New Caledonia" longLatPrecision="125" longitude="159.68333" pageId="4" pageNumber="903" specimenCode="DW 274" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="953EA884D4030C246E26FE3FB3EB2EC8" box="[113,219,411,434]" pageId="4" pageNumber="903" type="holotype">HOLOTYPE</typeStatus>
. — 
<emphasis id="78F1CA34D4030C246F52FE3EB2942ECB" bold="true" box="[261,420,410,434]" pageId="4" pageNumber="903">
<collectingCountry id="329256B6D4030C246F52FE3EB2902ECB" box="[261,416,410,434]" name="New Caledonia" pageId="4" pageNumber="903">New Caledonia</collectingCountry>
.
</emphasis>
MUSORSTOM 5, stn 
<specimenCode id="1A23BE5DD4030C246E27FE11B3F62EB4" box="[112,198,437,461]" collectionCode="DW" pageId="4" pageNumber="903">DW 274</specimenCode>
, 
<geoCoordinate id="2FB170E1D4030C246E98FE11B21E2EB4" box="[207,302,437,461]" degrees="24" direction="south" minutes="44.8" orientation="latitude" pageId="4" pageNumber="903" precision="92" value="-24.746666">24°44.8’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4030C246F61FE11B2942EB4" box="[310,420,437,461]" degrees="159" direction="east" minutes="41.0" orientation="longitude" pageId="4" pageNumber="903" precision="92" value="159.68333">159°41.0’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4030C246FFBFE11B2DB2EB4" box="[428,491,437,461]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="903" unit="m" value="285.0">
<elevation id="C1A8F115D4030C246FFBFE11B2DB2EB4" box="[428,491,437,461]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="903" unit="m" value="285.0">285 m</elevation>
</quantity>
, 
<date id="3E3B30E6D4030C246FA4FE11B17F2EB4" box="[499,591,437,461]" pageId="4" pageNumber="903" value="1986-10-09">
<collectingDate id="2E7FC90ED4030C246FA4FE11B17F2EB4" box="[499,591,437,461]" pageId="4" pageNumber="903" value="1986-10-09">9.X.1986</collectingDate>
</date>
, ovig. 
<specimenCount id="5C83DDAFD4030C246E27FE74B3B12E9E" box="[112,129,464,487]" pageId="4" pageNumber="903" type="female">♀</specimenCount>
(
<quantity id="8D7DBBC3D4030C246EC6FE6BB3D02E9E" box="[145,224,463,487]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="903" unit="mm" value="1.2">1.2 mm</quantity>
) (MNHN-Pg 7637).
</materialsCitation>
</paragraph>
<paragraph id="4A3A1626D4030C246E26FE5BB2F82D35" blockId="4.[112,647,511,588]" pageId="4" pageNumber="903">
NEW MATERIAL EXAMINED. — 
<emphasis id="78F1CA34D4030C246FDBFE5BB1162D6E" bold="true" box="[396,550,511,535]" pageId="4" pageNumber="903">
<collectingCountry id="329256B6D4030C246FDBFE5BB1112D6E" box="[396,545,511,535]" name="New Caledonia" pageId="4" pageNumber="903">New Caledonia</collectingCountry>
.
</emphasis>
EBISCO, stn 
<collectionCode id="2C948EE3D4030C246EC6FDBDB38C2D48" box="[145,188,537,561]" pageId="4" pageNumber="903">DW</collectionCode>
2492, 
<geoCoordinate id="2FB170E1D4030C246EABFDBDB26C2D48" box="[252,348,537,561]" degrees="24" direction="south" minutes="44.0" orientation="latitude" pageId="4" pageNumber="903" precision="92" value="-24.733334">24°44.0’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4030C246F32FDBDB2E42D48" box="[357,468,537,561]" degrees="159" direction="east" minutes="41.0" orientation="longitude" pageId="4" pageNumber="903" precision="92" value="159.68333">159°41.0’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4030C246F8BFDBDB12B2D48" box="[476,539,537,561]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="903" unit="m" value="285.0">285 m</quantity>
, 
<date id="3E3B30E6D4030C246C73FDBDB1B02D48" box="[548,640,537,561]" pageId="4" pageNumber="903" value="2005-10-06">6.X.2005</date>
, 
<specimenCount id="5C83DDAFD4030C246E27FD90B3A82D35" box="[112,152,564,588]" pageId="4" pageNumber="903" type="male">1 ♂</specimenCount>
(
<quantity id="8D7DBBC3D4030C246EF1FD90B3C62D35" box="[166,246,564,588]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="903" unit="mm" value="1.3">1.3 mm</quantity>
) (MNHN-Pg 7763).
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4030C246E26FDC0B3872DC9" pageId="4" pageNumber="903" type="distribution">
<paragraph id="4A3A1626D4030C246E26FDC0B3872DC9" blockId="4.[111,647,611,688]" pageId="4" pageNumber="903">
<materialsCitation id="FAED1C7BD4030C246E26FDC0B1B42DEF" country="New Caledonia" location="Chesterfield Plateau" pageId="4" pageNumber="903" specimenCount="1">
DISTRIBUTION. — Presently known only from the 
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locality, 
<location id="4F5A40FDD4030C246EECFDDAB2462DEF" LSID="urn:lsid:plazi:treatment:C22CA730D4030C2A6F43FF7CB2E329DD:4F5A40FDD4030C246EECFDDAB2462DEF" box="[187,374,638,662]" country="New Caledonia" name="Chesterfield Plateau" pageId="4" pageNumber="903">Chesterfield Plateau</location>
, and vicinity, 
<collectingCountry id="329256B6D4030C246FA6FDDAB1B42DEF" box="[497,644,638,662]" name="New Caledonia" pageId="4" pageNumber="903">New Caledonia</collectingCountry>
</materialsCitation>
; 
<specimenCount id="5C83DDAFD4030C246E38FD3CB3872DC9" box="[111,183,664,688]" pageId="4" pageNumber="903" type="generic">
<quantity id="8D7DBBC3D4030C246E38FD3CB3832DC9" box="[111,179,664,688]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="903" unit="m" value="285.0">285 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4030C266E26FD77B1B72CD5" lastPageId="6" lastPageNumber="905" pageId="4" pageNumber="903" type="description">
<paragraph id="4A3A1626D4030C246E26FD77B2552D92" blockId="4.[111,649,723,1700]" box="[113,357,723,749]" pageId="4" pageNumber="903">DESCRIPTION OF MALE</paragraph>
<paragraph id="4A3A1626D4030C246E26FD56B1B92B52" blockId="4.[111,649,723,1700]" pageId="4" pageNumber="903">
Shield (
<figureCitation id="D2BE0AA3D4030C246E95FD56B2212C74" box="[194,273,754,781]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="4" pageNumber="903">Fig. 1A</figureCitation>
) slightly broader than long, weakly calcified; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin roundly truncate. Triangular rostrum produced slightly beyond midlength of ocular acicles; lateral projections each with acute marginal spine. Carapace lateral lobes reaching 0.4 of shield.Posterior carapace with moderately narrow median plate; cardiac sulci reaching to posterior margin. Branchiostegites membranous, unarmed.
</paragraph>
<paragraph id="4A3A1626D4030C246EDFFB8BB3812A71" blockId="4.[111,649,723,1700]" pageId="4" pageNumber="903">Ocular peduncles moderately stout but not noticeably broadened distally, 0.7 length of shield; corneal diameter 0.5 of total peduncular length; ocular acicles narrowly triangular, reaching to or beyond proximal margin of ultimate peduncular segment, each with moderately large submarginal spine.</paragraph>
<paragraph id="4A3A1626D4030C246EDFFAA9B2822A9F" blockId="4.[111,649,723,1700]" pageId="4" pageNumber="903">Antennular peduncles overreaching distal margins of corneas by slightly less than lengths of ultimate peduncular segments; ultimate segment with 1 long, stiff seta on dorsodistal margin; penultimate segment with very few scattered setae; basal segment with acute spine on dorsolateral margin, ventral margin not produced distally.</paragraph>
<paragraph id="4A3A1626D4030C246EDFFA4FB0562EC8" blockId="4.[111,649,723,1700]" lastBlockId="4.[674,1212,216,1700]" pageId="4" pageNumber="903">Antennal peduncles overreaching distal corneal margins by 0.5 lengths of ultimate segments. Fifth and fourth segments with very few scattered setae; third segment with small spine at ventrodistal angle; second segment with dorsolateral distal angle produced, terminating in small spine, dorsomesial angle with prominent spine; first segment with spine at dorsolateral distal margin, ventrolateral margin unarmed. Antennal acicle reaching beyond distal margin of fourth peduncular segment, moderately slender, terminating in simple spine. Antennal flagellum with 1 or 2 very short to moderately long setae every article.</paragraph>
<paragraph id="4A3A1626D4030C246CECFE12B0C22BD0" blockId="4.[674,1212,216,1700]" pageId="4" pageNumber="903">
Chelipeds subequal in length, but right slightly stouter; dactyls and fixed fingers each with small hiatus. Right cheliped (
<figureCitation id="D2BE0AA3D4030C246DF5FE51B0C22D69" box="[930,1010,501,528]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="4" pageNumber="903">Fig. 1B</figureCitation>
) with dactyl equal to length of palm; dorsomesial margin smooth, dorsal surface somewhat convex, unarmed, but with few scattered, short setae mesially and ventrally; cutting edge with 2 low, broad, calcareous teeth, terminating in tiny corneous claw, slightly overlapped by fixed finger. Palm longer than carpus, dorsomesial margin delimited by 3 widely-spaced, acute spines and sparse tufts of setae, dorsal surface weakly convex, unarmed; dorsal surface of fixed finger similarly unarmed, dorsolateral margin smooth and with few scattered short setae; cutting edge with 3 broad, low calcareous teeth, terminating in tiny corneous claw. Carpus approximately equal to length of merus; dorsomesial distal angle with acute spine and 1 additional small spine on dorsomesial margin, dorsal surface with 2 small spines in lateral 0.3, dorsolateral margin not distinctly delimited; remaining surfaces unarmed. Merus with all surfaces unarmed; ventromesial and ventrolateral margins each with small spine at distal angle. Ischium unarmed.
</paragraph>
<paragraph id="4A3A1626D4030C256CECFB0AB24B293C" blockId="4.[674,1212,216,1700]" lastBlockId="5.[111,647,1420,1700]" lastPageId="5" lastPageNumber="904" pageId="4" pageNumber="903">
Left cheliped (
<figureCitation id="D2BE0AA3D4030C246D0DFB0AB09D2BB0" box="[858,941,1198,1225]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="4" pageNumber="903">Fig. 1C</figureCitation>
) approximately equal to length of right, slightly slenderer. Dactyl equal to length of palm; surfaces rounded and unarmed, but with sparsely scattered, moderately long setae ventrally; cutting edge with row of minute calcareous teeth, terminating in small corneous claw. Palm with convex dorsal surface unarmed but with few scattered setae; dorsomesial margin with 3 small spines, dorsolateral margin not delimited; fixed finger rounded and unarmed but with several moderately long setae laterally; cutting edge with row of minute calcareous teeth, terminating in small corneous claw. Carpus equal to length of palm; dorsomesial margin not distinctly delimited but with acute spine at distal angle, without small spine at midlength, dorsal surface with 2 small spines laterad of midline, dorsolateral margin not delimited;remaining surfaces unarmed but with few scattered setae, ventromesial distal angle with small spine. Merus with surfaces unarmed but with few scattered short setae, ventromesial and ventrolateral distal angles each with small spine. Ischium unarmed.
</paragraph>
<caption id="1EFA46AED4020C256E26FB18B0442A46" ID-DOI="http://doi.org/10.5281/zenodo.4525844" ID-Zenodo-Dep="4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="5" pageNumber="904" startId="5.[113,124,1212,1229]" targetBox="[127,1199,215,1171]" targetPageId="5">
<paragraph id="4A3A1626D4020C256E26FB18B0442A46" blockId="5.[113,1211,1212,1343]" pageId="5" pageNumber="904">
FIG. 1. — 
<taxonomicName id="8D856DA5D4020C256E92FB18B1502BB4" authority="McLaughlin &amp; Rahayu, 2006" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[197,608,1212,1229]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="904" phylum="Arthropoda" rank="species" species="spina">
<emphasis id="78F1CA34D4020C256E92FB18B2582BB4" box="[197,360,1212,1229]" italics="true" pageId="5" pageNumber="904">Pteropagurus spina</emphasis>
<bibRefCitation id="2E146BD7D4020C256F39FB18B1502BB4" author="MCLAUGHLIN P. A. &amp; RAHAYU D. L." box="[366,608,1212,1229]" pageId="5" pageNumber="904" pagination="55 - 68" refId="ref10933" refString="MCLAUGHLIN P. A. &amp; RAHAYU D. L. 2006. - A new genus with two new species of hermit crabs (Crustacea, Decapoda, Paguroidea, Paguridae) from an unique habitat. Zootaxa 1116: 55 - 68." type="journal article" year="2006">McLaughlin &amp; Rahayu, 2006</bibRefCitation>
</taxonomicName>
, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): 
<emphasis id="78F1CA34D4020C256E26FB76B34E2B9A" bold="true" box="[113,126,1234,1251]" pageId="5" pageNumber="904">A</emphasis>
, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); 
<emphasis id="78F1CA34D4020C256D1FFB76B0652B9A" bold="true" box="[840,853,1234,1251]" pageId="5" pageNumber="904">B</emphasis>
, carpus and chela of right cheliped, dorsal view (setae omitted); 
<emphasis id="78F1CA34D4020C256F74FB4DB2012B83" bold="true" box="[291,305,1257,1274]" pageId="5" pageNumber="904">C</emphasis>
, carpus and chela of left cheliped, dorsal view (setae omitted); 
<emphasis id="78F1CA34D4020C256D14FB4DB0612B83" bold="true" box="[835,849,1257,1274]" pageId="5" pageNumber="904">D</emphasis>
, right second pereopod, lateral view (setae shown for dactyl only); 
<emphasis id="78F1CA34D4020C256F63FAA4B2702A68" bold="true" box="[308,320,1280,1297]" pageId="5" pageNumber="904">E</emphasis>
, left third pereopod, lateral view (setae shown for dactyl only); 
<emphasis id="78F1CA34D4020C256D19FAA4B0692A68" bold="true" box="[846,857,1280,1297]" pageId="5" pageNumber="904">F</emphasis>
, sternites of second and third pereopods, ventral view (setae omitted); 
<emphasis id="78F1CA34D4020C256F34FAB2B2412A5E" bold="true" box="[355,369,1302,1319]" pageId="5" pageNumber="904">G</emphasis>
, distal three segments of left fourth pereopod, lateral view (setae omitted); 
<emphasis id="78F1CA34D4020C256DBAFAB2B0CB2A5E" bold="true" box="[1005,1019,1302,1319]" pageId="5" pageNumber="904">H</emphasis>
, coxae and sternite of fifth pereopods, ventral view; 
<emphasis id="78F1CA34D4020C256F3FFA89B25E2A47" bold="true" box="[360,366,1325,1342]" pageId="5" pageNumber="904">I</emphasis>
, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm.
</paragraph>
</caption>
<paragraph id="4A3A1626D4020C266EDFF9EEB2DA2EE8" blockId="5.[111,647,1420,1700]" lastBlockId="6.[111,649,216,750]" lastPageId="6" lastPageNumber="905" pageId="5" pageNumber="904">
Second and third pereopods (
<figureCitation id="D2BE0AA3D4020C256FE8F9EEB101291C" box="[447,561,1610,1637]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="5" pageNumber="904">Fig. 1D, E</figureCitation>
) similar in armature but unequal in length, second shorter. Dactyls relatively straight in dorsal and lateral views, slightly shorter to slightly longer than propodi, dorsal surfaces each with row of widely-spaced setae; ventral surfaces each with 4 or 5 corneous spines and few setae. Propodi slightly to considerably longer than carpi; surfaces all unarmed but with sparse setae dorsally and ventrally. Carpi of second pereopods each with dorsodistal spine and 1 additional spine on dorsal surface in proximal 0.5; carpi of third each without minute spinule at dorsodistal margin, other surfaces unarmed but all with sparse setae. Meri unarmed, but each with few widely-spaced setae dorsally. Ischia unarmed. Sternite of third pereopods (
<figureCitation id="D2BE0AA3D4010C266FCCFE93B2D82E2B" box="[411,488,311,338]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="6" pageNumber="905">Fig. 1F</figureCitation>
) not produced posteriorly. Fourth pereopods semichelate; propodal rasp (
<figureCitation id="D2BE0AA3D4010C266EFDFED2B3CD2EE8" box="[170,253,374,401]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="6" pageNumber="905">Fig. 1G</figureCitation>
) with 1 row of scales.
</paragraph>
<paragraph id="4A3A1626D4010C266EDFFE32B2A92D36" blockId="6.[111,649,216,750]" pageId="6" pageNumber="905">
Extremely long sexual tube developed from coxa of left fifth pereopod (
<figureCitation id="D2BE0AA3D4010C266F3DFE12B2F32EA8" box="[362,451,438,465]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="6" pageNumber="905">Fig. 1H</figureCitation>
), directed toward exterior, curved up and over dorsal surface of pleon and ventrally to level of coxa of right fifth pereopod; right coxa with gonopore circumscribed by moderately long, stiff setae.
</paragraph>
<paragraph id="4A3A1626D4010C266EDFFDF0B2912D97" blockId="6.[111,649,216,750]" pageId="6" pageNumber="905">
Uropods symmetrical, each with 2 rows of scales on endopod and exopod. Telson (
<figureCitation id="D2BE0AA3D4010C266FA5FDD0B10F2DF6" box="[498,575,628,655]" captionStart="FIG" captionStartId="5.[113,124,1212,1229]" captionTargetBox="[127,1199,215,1171]" captionTargetId="figure-27@5.[93,1230,162,1188]" captionTargetPageId="5" captionText="FIG. 1. — Pteropagurus spina McLaughlin &amp; Rahayu, 2006, ♂ (1.3 mm), EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7763): A, shield and cephalic appendages, dorsal view (aesthetascs and most setae omitted); B, carpus and chela of right cheliped, dorsal view (setae omitted); C, carpus and chela of left cheliped, dorsal view (setae omitted); D, right second pereopod, lateral view (setae shown for dactyl only); E, left third pereopod, lateral view (setae shown for dactyl only); F, sternites of second and third pereopods, ventral view (setae omitted); G, distal three segments of left fourth pereopod, lateral view (setae omitted); H, coxae and sternite of fifth pereopods, ventral view; I, telson, dorsal view. Scale bar: A-F, H, 0.5 mm; G, I, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525844" httpUri="https://zenodo.org/record/4525844/files/figure.png" pageId="6" pageNumber="905">Fig. 1I</figureCitation>
) with posterior lobes separated by shallow median cleft, oblique terminal margins each with 3 or 4 very small spines, largest at apex.
</paragraph>
<paragraph id="4A3A1626D4010C266E26FCB6B3EF2C55" blockId="6.[113,246,786,845]" box="[113,223,786,812]" pageId="6" pageNumber="905">
<emphasis id="78F1CA34D4010C266E26FCB6B3EF2C55" box="[113,223,786,812]" italics="true" pageId="6" pageNumber="905">Coloration</emphasis>
</paragraph>
<paragraph id="4A3A1626D4010C266E26FC96B3C62C34" blockId="6.[113,246,786,845]" box="[113,246,818,845]" pageId="6" pageNumber="905">Not known.</paragraph>
<paragraph id="4A3A1626D4010C266E26FCD5B3F32CF2" blockId="6.[113,647,881,940]" box="[113,195,881,907]" pageId="6" pageNumber="905">
<emphasis id="78F1CA34D4010C266E26FCD5B3F32CF2" box="[113,195,881,907]" italics="true" pageId="6" pageNumber="905">Habitat</emphasis>
</paragraph>
<paragraph id="4A3A1626D4010C266E26FC35B1B72CD5" blockId="6.[113,647,881,940]" box="[113,647,913,940]" pageId="6" pageNumber="905">
Empty shells of 
<taxonomicName id="8D856DA5D4010C266F40FC35B1B32CD5" authority="(Rang, 1827)" baseAuthorityName="Rang" baseAuthorityYear="1827" box="[279,643,913,940]" class="Gastropoda" family="Cuvierinidae" genus="Cuvierina" kingdom="Animalia" order="Thecosomata" pageId="6" pageNumber="905" phylum="Mollusca" rank="species" species="columnella">
<emphasis id="78F1CA34D4010C266F40FC35B2DE2CD2" box="[279,494,913,939]" italics="true" pageId="6" pageNumber="905">Cuvierina columnella</emphasis>
(Rang, 1827)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4010C266E26FC75B29F2ABF" pageId="6" pageNumber="905" type="discussion">
<paragraph id="4A3A1626D4010C266E26FC75B3E52C90" blockId="6.[111,648,977,1478]" box="[113,213,977,1003]" pageId="6" pageNumber="905">REMARKS</paragraph>
<paragraph id="4A3A1626D4010C266E26FC54B29F2ABF" blockId="6.[111,648,977,1478]" pageId="6" pageNumber="905">
Because of the paucity of material, the mouthparts of 
<taxonomicName id="8D856DA5D4010C266E38FBB4B38C2B50" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[111,188,1040,1065]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="905" phylum="Arthropoda" rank="species" species="spina">
<emphasis id="78F1CA34D4010C266E38FBB4B38C2B50" box="[111,188,1040,1065]" italics="true" pageId="6" pageNumber="905">P. spina</emphasis>
</taxonomicName>
have not been removed for description and illustration, but appear to be in general agreement with those illustrated by 
<bibRefCitation id="2E146BD7D4010C266FD2FBEBB38E2BF3" author="MCLAUGHLIN P. A. &amp; RAHAYU D. L." pageId="6" pageNumber="905" pagination="55 - 68" refId="ref10933" refString="MCLAUGHLIN P. A. &amp; RAHAYU D. L. 2006. - A new genus with two new species of hermit crabs (Crustacea, Decapoda, Paguroidea, Paguridae) from an unique habitat. Zootaxa 1116: 55 - 68." type="journal article" year="2006">McLaughlin &amp; Rahayu (2006)</bibRefCitation>
for 
<taxonomicName id="8D856DA5D4010C266EBAFBCBB2642BF1" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[237,340,1135,1160]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="905" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="78F1CA34D4010C266EBAFBCBB2642BF1" box="[237,340,1135,1160]" italics="true" pageId="6" pageNumber="905">P. inermis</emphasis>
</taxonomicName>
, including the several teeth on the crista dentata of the third maxilliped. The general morphology of the male of 
<taxonomicName id="8D856DA5D4010C266FB9FB0BB10D2BB1" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[494,573,1199,1224]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="905" phylum="Arthropoda" rank="species" species="spina">
<emphasis id="78F1CA34D4010C266FB9FB0BB10D2BB1" box="[494,573,1199,1224]" italics="true" pageId="6" pageNumber="905">P. spina</emphasis>
</taxonomicName>
differs only slightly from the female in proportions and spination of the chelipeds and ambulatory legs. However, the scales of the propodal rasp of the fourth pereopod form only a single row in this male. Having only one specimen of each sex, it is not possible to determine whether the two rows of spines in the female reflect a sexually dimorphic character or simply variation.
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4010C266E29F9AFB39929DA" pageId="6" pageNumber="905" type="reference_group">
<paragraph id="4A3A1626D4010C266E29F9AFB283293C" blockId="6.[126,634,1546,1605]" pageId="6" pageNumber="905">
<taxonomicName id="8D856DA5D4010C266E29F9AFB14A295C" ID-CoL="ed4d9b61-a923-4aa7-bdfa-e4010cf142d8" authority="McLaughlin, 2007" authorityName="McLaughlin" authorityYear="2007" box="[126,634,1546,1573]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="905" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D4010C266E29F9AFB298295C" bold="true" box="[126,424,1546,1573]" italics="true" pageId="6" pageNumber="905">Pteropagurus spinulocarpus</emphasis>
<bibRefCitation id="2E146BD7D4010C266FF8F9AEB14A295C" author="MCLAUGHLIN P. A." box="[431,634,1546,1573]" pageId="6" pageNumber="905" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin, 2007</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D2BE0AA3D4010C266F1BF98EB2A0293C" box="[332,400,1578,1605]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="6" pageNumber="905">Figs 2</figureCitation>
; 
<figureCitation id="D2BE0AA3D4010C266FCBF98EB29A293C" box="[412,426,1578,1605]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="6" pageNumber="905">3</figureCitation>
)
</paragraph>
<paragraph id="4A3A1626D4010C266E26F9D5B39929DA" blockId="6.[113,646,1648,1699]" pageId="6" pageNumber="905">
<treatmentCitationGroup id="6A953108D4010C266E26F9D5B39929DA" pageId="6" pageNumber="905">
<treatmentCitation id="CB243037D4010C266E26F9D5B1B229F1" author="MCLAUGHLIN P. A." box="[113,642,1648,1672]" page="505" pageId="6" pageNumber="905" year="2007">
<taxonomicName id="8D856DA5D4010C266E26F9D5B1B229F1" ID-CoL="ed4d9b61-a923-4aa7-bdfa-e4010cf142d8" authority="McLaughlin, 2007: 505" authorityName="McLaughlin" authorityPageNumber="505" authorityYear="2007" box="[113,642,1648,1672]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="905" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D4010C266E26F9D5B24E29F1" box="[113,382,1648,1672]" italics="true" pageId="6" pageNumber="905">Pteropagurus spinulocarpus</emphasis>
<bibRefCitation id="2E146BD7D4010C266FD0F9D4B1B229F1" author="MCLAUGHLIN P. A." box="[391,642,1648,1672]" pageId="6" pageNumber="905" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin, 2007: 505</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4010C266CF2FF7CB7772D5E" pageId="6" pageNumber="905" type="materials_examined">
<paragraph id="4A3A1626D4010C266CF2FF7CB0F02E46" blockId="6.[675,1209,215,319]" pageId="6" pageNumber="905">
<materialsCitation id="FAED1C7BD4010C266CF2FF7CB0F02E46" collectingDate="1985-08-30" collectionCode="DW" country="New Caledonia" elevation="445" latitude="-22.788334" location="New Caledonia" longLatPrecision="12" longitude="167.23833" pageId="6" pageNumber="905" specimenCode="DW 44" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="953EA884D4010C266CF2FF7CB03E2F97" box="[677,782,216,239]" pageId="6" pageNumber="905" type="holotype">HOLOTYPE</typeStatus>
. — 
<emphasis id="78F1CA34D4010C266D6EFF73B0E92F96" bold="true" box="[825,985,215,239]" pageId="6" pageNumber="905">
<collectingCountry id="329256B6D4010C266D6EFF73B0E52F96" box="[825,981,215,239]" name="New Caledonia" pageId="6" pageNumber="905">New Caledonia</collectingCountry>
.
</emphasis>
BIOCAL, stn 
<specimenCode id="1A23BE5DD4010C266A3EFF73B7862F96" box="[1129,1206,215,239]" collectionCode="DW" pageId="6" pageNumber="905">DW 44</specimenCode>
, 
<geoCoordinate id="2FB170E1D4010C266CF3FF56B0222E73" box="[676,786,242,266]" degrees="22" direction="south" minutes="47.30" orientation="latitude" pageId="6" pageNumber="905" precision="9" value="-22.788334">22°47.30’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266D4DFF56B0A72E73" box="[794,919,242,266]" degrees="167" direction="east" minutes="14.30" orientation="longitude" pageId="6" pageNumber="905" precision="9" value="167.23833">167°14.30’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266DF7FF56B73E2E73" box="[928,1038,242,266]" metricMagnitude="2" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="6" pageNumber="905" unit="m" value="445.0" valueMax="450.0" valueMin="440.0">
<elevation id="C1A8F115D4010C266DF7FF56B73E2E73" box="[928,1038,242,266]" metricMagnitude="2" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="6" pageNumber="905" unit="m" value="445.0" valueMax="450.0" valueMin="440.0">440-450 m</elevation>
</quantity>
, 
<date id="3E3B30E6D4010C266A40FF56B7AC2E73" box="[1047,1180,242,266]" pageId="6" pageNumber="905" value="1985-08-30">
<collectingDate id="2E7FC90ED4010C266A40FF56B7AC2E73" box="[1047,1180,242,266]" pageId="6" pageNumber="905" value="1985-08-30">30.VIII.1985</collectingDate>
</date>
, 
<specimenCount id="5C83DDAFD4010C266AF2FF57B7892E73" box="[1189,1209,243,266]" pageId="6" pageNumber="905" type="male">♂</specimenCount>
(
<quantity id="8D7DBBC3D4010C266CFBFEA8B1CB2E5D" box="[684,763,268,292]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="905" unit="mm" value="1.6">1.6 mm</quantity>
, posterior portion of pleon, uropods and telson missing) (MNHN-Pg 7738).
</materialsCitation>
</paragraph>
<paragraph id="4A3A1626D4010C266CF2FEF3B7772D5E" blockId="6.[674,1211,342,737]" pageId="6" pageNumber="905">
<typeStatus id="953EA884D4010C266CF2FEF3B0202E14" box="[677,784,343,366]" pageId="6" pageNumber="905" type="paratype">PARATYPES</typeStatus>
. — 
<emphasis id="78F1CA34D4010C266D15FEF2B0DC2E17" bold="true" box="[834,1004,342,366]" pageId="6" pageNumber="905">
<collectingCountry id="329256B6D4010C266D15FEF2B0D82E17" box="[834,1000,342,366]" name="New Caledonia" pageId="6" pageNumber="905">New Caledonia</collectingCountry>
.
</emphasis>
MUSORSTOM 4, stn 
<collectionCode id="2C948EE3D4010C266C9AFED5B1C12EF0" box="[717,753,369,393]" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="6" pageNumber="905">CC</collectionCode>
212, 
<geoCoordinate id="2FB170E1D4010C266D66FED5B0942EF0" box="[817,932,369,393]" degrees="22" direction="south" minutes="47.40" orientation="latitude" pageId="6" pageNumber="905" precision="9" value="-22.79">22°47.40’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266DE6FED5B7042EF0" box="[945,1076,369,393]" degrees="167" direction="east" minutes="10.50" orientation="longitude" pageId="6" pageNumber="905" precision="9" value="167.175">167°10.50’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266A16FED5B7862EF0" box="[1089,1206,369,393]" metricMagnitude="2" metricUnit="m" metricValue="3.775" metricValueMax="3.8" metricValueMin="3.75" pageId="6" pageNumber="905" unit="m" value="377.5" valueMax="380.0" valueMin="375.0">375-380 m</quantity>
, 
<date id="3E3B30E6D4010C266CF1FE2FB02A2EDA" box="[678,794,395,419]" pageId="6" pageNumber="905" value="1985-09-28">28.IX.1985</date>
, 1 ovig. 
<specimenCount id="5C83DDAFD4010C266D33FE28B0452EDA" box="[868,885,396,419]" pageId="6" pageNumber="905" type="female">♀</specimenCount>
(
<quantity id="8D7DBBC3D4010C266DD5FE2FB0E12EDA" box="[898,977,395,419]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="905" unit="mm" value="1.5">1.5 mm</quantity>
, right cheliped and right second pereopod missing, left third pereopod regenerating) (MNHN-Pg 7737). — Stn 
<collectionCode id="2C948EE3D4010C266DB4FE64B73F2EA1" box="[995,1039,448,472]" pageId="6" pageNumber="905">DW</collectionCode>
222, 
<geoCoordinate id="2FB170E1D4010C266A10FE64B7852EA1" box="[1095,1205,448,472]" degrees="22" direction="south" minutes="57.60" orientation="latitude" pageId="6" pageNumber="905" precision="9" value="-22.96">22°57.60’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266CF2FE7EB0332E8B" box="[677,771,474,498]" degrees="167" direction="east" minutes="33" orientation="longitude" pageId="6" pageNumber="905" precision="925" value="167.55">167°33’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266D5BFE7EB04C2E8B" box="[780,892,474,498]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="6" pageNumber="905" unit="m" value="425.0" valueMax="440.0" valueMin="410.0">410-440 m</quantity>
, 
<date id="3E3B30E6D4010C266DD1FE7EB0CA2E8B" box="[902,1018,474,498]" pageId="6" pageNumber="905" value="1985-09-30">30.IX.1985</date>
, 
<specimenCount id="5C83DDAFD4010C266A54FE7EB7162E8A" box="[1027,1062,474,499]" pageId="6" pageNumber="905" type="female">1 ♀</specimenCount>
(
<quantity id="8D7DBBC3D4010C266A63FE7EB7B42E8B" box="[1076,1156,474,498]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="905" unit="mm" value="1.5">1.5 mm</quantity>
, with rhizocephalan, missing both chelipeds, left second and third right pereopods) (MNHN-Pg 7701).
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4010C266CF2FD8EB1D72C03" pageId="6" pageNumber="905" type="description">
<paragraph id="4A3A1626D4010C266CF2FD8EB0582D98" blockId="6.[674,1211,342,737]" pageId="6" pageNumber="905">
<materialsCitation id="FAED1C7BD4010C266CF2FD8EB0542D98" collectingDate="1986-10-09" collectingDateMax="1986-10-19" collectingDateMin="1986-10-09" collectionCode="DW, DC" elevation="285" latitude="-19.875" longLatPrecision="12" longitude="158.635" pageId="6" pageNumber="905" specimenCode="DW 274, DC 361" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
MUSORSTOM 5, stn 
<specimenCode id="1A23BE5DD4010C266D20FD8EB0FB2D3B" box="[887,971,554,578]" collectionCode="DW" pageId="6" pageNumber="905">DW 274</specimenCode>
, 
<geoCoordinate id="2FB170E1D4010C266D85FD8EB70A2D3B" box="[978,1082,554,578]" degrees="24" direction="south" minutes="44.84" orientation="latitude" pageId="6" pageNumber="905" precision="9" value="-24.747334">24°44.84’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266A17FD8EB7862D3B" box="[1088,1206,554,578]" degrees="159" direction="east" minutes="41.00" orientation="longitude" pageId="6" pageNumber="905" precision="9" value="159.68333">159°41.00’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266CF5FDE0B1D42D25" box="[674,740,580,604]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="6" pageNumber="905" unit="m" value="285.0">
<elevation id="C1A8F115D4010C266CF5FDE0B1D42D25" box="[674,740,580,604]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="6" pageNumber="905" unit="m" value="285.0">285 m</elevation>
</quantity>
, 
<date id="3E3B30E6D4010C266CB9FDE0B07D2D25" box="[750,845,580,604]" pageId="6" pageNumber="905" value="1986-10-09">
<collectingDate id="2E7FC90ED4010C266CB9FDE0B07D2D25" box="[750,845,580,604]" pageId="6" pageNumber="905" value="1986-10-09">9.X.1986</collectingDate>
</date>
, 
<specimenCount id="5C83DDAFD4010C266D01FDE0B04C2D25" box="[854,892,580,604]" pageId="6" pageNumber="905" type="female">1 ♀</specimenCount>
(
<quantity id="8D7DBBC3D4010C266DDEFDE0B0E92D25" box="[905,985,580,604]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="905" unit="mm" value="1.2">1.2 mm</quantity>
, missing right cheliped and both third pereopods) (MNHN-Pg 7702). — Stn 
<specimenCode id="1A23BE5DD4010C266CF2FDDDB1C72DE8" box="[677,759,633,657]" collectionCode="DC" pageId="6" pageNumber="905">DC 361</specimenCode>
, 
<geoCoordinate id="2FB170E1D4010C266D56FDDDB0402DE8" box="[769,880,633,657]" degrees="19" direction="south" minutes="52.50" orientation="latitude" pageId="6" pageNumber="905" precision="9" value="-19.875">19°52.50’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266D2EFDDDB0C62DE8" box="[889,1014,633,657]" degrees="158" direction="east" minutes="38.10" orientation="longitude" pageId="6" pageNumber="905" precision="9" value="158.635">158°38.10’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266DA8FDDDB7712DE8" box="[1023,1089,633,657]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="905" unit="m" value="400.0">
<elevation id="C1A8F115D4010C266DA8FDDDB7712DE8" box="[1023,1089,633,657]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="905" unit="m" value="400.0">400 m</elevation>
</quantity>
, 
<date id="3E3B30E6D4010C266A1DFDDDB7852DE8" box="[1098,1205,633,657]" pageId="6" pageNumber="905" value="1986-10-19">
<collectingDate id="2E7FC90ED4010C266A1DFDDDB7852DE8" box="[1098,1205,633,657]" pageId="6" pageNumber="905" value="1986-10-19">19.X.1986</collectingDate>
</date>
, 
<specimenCount id="5C83DDAFD4010C266CF2FD30B1FB2DD5" box="[677,715,660,684]" pageId="6" pageNumber="905" type="male">1 ♂</specimenCount>
(
<quantity id="8D7DBBC3D4010C266C8FFD30B0182DD5" box="[728,808,660,684]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="905" unit="mm" value="1.5">1.5 mm</quantity>
, right cheliped regenerating; left cheliped with fixed finger broken off; left third pereopod missing) (MNHN-Pg 7700)
</materialsCitation>
.
</paragraph>
<paragraph id="4A3A1626D4010C266CF2FD5DB1D72C03" blockId="6.[675,1210,760,890]" pageId="6" pageNumber="905">
NEW MATERIAL EXAMINED. — 
<emphasis id="78F1CA34D4010C266DE8FD5CB7692C69" bold="true" box="[959,1113,760,784]" pageId="6" pageNumber="905">
<collectingCountry id="329256B6D4010C266DE8FD5CB7642C69" box="[959,1108,760,784]" name="New Caledonia" pageId="6" pageNumber="905">New Caledonia</collectingCountry>
.
</emphasis>
EBISCO, stn DW 2492, 
<geoCoordinate id="2FB170E1D4010C266D67FCB7B0BF2C52" box="[816,911,787,811]" degrees="24" direction="south" minutes="44.0" orientation="latitude" pageId="6" pageNumber="905" precision="92" value="-24.733334">24°44.0’S</geoCoordinate>
, 
<geoCoordinate id="2FB170E1D4010C266DCFFCB7B7372C52" box="[920,1031,787,811]" degrees="159" direction="east" minutes="41.0" orientation="longitude" pageId="6" pageNumber="905" precision="92" value="159.68333">159°41.0’E</geoCoordinate>
, 
<quantity id="8D7DBBC3D4010C266A58FCB7B77F2C52" box="[1039,1103,787,811]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="6" pageNumber="905" unit="m" value="285.0">285 m</quantity>
, 
<date id="3E3B30E6D4010C266A00FCB7B7832C52" box="[1111,1203,787,811]" pageId="6" pageNumber="905" value="2005-10-06">6.X.2005</date>
, 
<specimenCount id="5C83DDAFD4010C266CF4FC89B1C02C3C" box="[675,752,813,837]" pageId="6" pageNumber="905" type="male">12 ♂♂</specimenCount>
(
<quantity id="8D7DBBC3D4010C266D56FC89B0B12C3C" box="[769,897,813,837]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="6" pageNumber="905" unit="mm" value="1.45" valueMax="1.7" valueMin="1.2">1.2-1.7 mm</quantity>
, 1 with rhizocephalan), 
<specimenCount id="5C83DDAFD4010C266AD7FC89B78A2C3C" box="[1152,1210,813,837]" pageId="6" pageNumber="905" type="female">7 ♀♀</specimenCount>
(
<quantity id="8D7DBBC3D4010C266CFBFCECB0152C19" box="[684,805,840,864]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="6" pageNumber="905" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4-1.7 mm</quantity>
), 15 ovig. 
<specimenCount id="5C83DDAFD4010C266DDEFCEDB09C2C19" box="[905,940,841,864]" pageId="6" pageNumber="905" type="female">♀♀</specimenCount>
(
<quantity id="8D7DBBC3D4010C266DEEFCECB7002C19" box="[953,1072,840,864]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="6" pageNumber="905" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4-1.6 mm</quantity>
) (MNHN-Pg 7764).
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD4010C296CF2FC36B0762B52" lastPageId="9" lastPageNumber="908" pageId="6" pageNumber="905" type="distribution">
<paragraph id="4A3A1626D4010C266CF2FC36B1DC2CA6" blockId="6.[677,1210,914,991]" pageId="6" pageNumber="905">
<materialsCitation id="FAED1C7BD4010C266CF2FC36B7302CBD" country="New Caledonia" location="Isle of Pines" pageId="6" pageNumber="905" specimenCount="1">
DISTRIBUTION. — 
<collectingCountry id="329256B6D4010C266D0FFC36B0DB2CD3" box="[856,1003,914,938]" name="New Caledonia" pageId="6" pageNumber="905">New Caledonia</collectingCountry>
from the Chesterfield Plateau to south of the 
<location id="4F5A40FDD4010C266DDEFC08B7302CBD" LSID="urn:lsid:plazi:treatment:C22CA730D4030C2A6F43FF7CB2E329DD:4F5A40FDD4010C266DDEFC08B7302CBD" box="[905,1024,940,964]" country="New Caledonia" name="Isle of Pines" pageId="6" pageNumber="905">Isle of Pines</location>
</materialsCitation>
; 285-440, possibly 
<specimenCount id="5C83DDAFD4010C266CF2FC63B1DC2CA6" box="[677,748,967,991]" pageId="6" pageNumber="905" type="generic">
<quantity id="8D7DBBC3D4010C266CF2FC63B1D82CA6" box="[677,744,967,991]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="905" unit="m" value="450.0">450 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="4A3A1626D4010C266CF2FBB5B0E92B51" blockId="6.[675,1212,1041,1700]" box="[677,985,1041,1067]" pageId="6" pageNumber="905">SUPPLEMENTAL DESCRIPTION</paragraph>
<paragraph id="4A3A1626D4010C276CF2FB8BB291295C" blockId="6.[675,1212,1041,1700]" lastBlockId="7.[113,649,1515,1700]" lastPageId="7" lastPageNumber="906" pageId="6" pageNumber="905">
Shield (
<figureCitation id="D2BE0AA3D4010C266CA1FB8BB0782B33" box="[758,840,1071,1098]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="6" pageNumber="905">Fig. 2A</figureCitation>
) with rostral lobe slightly to moderately produced, very faintly rounded to broadly and bluntly triangular, unarmed; lateral projections obsolete to roundly triangular, unarmed or with tiny marginal spinule. Ocular peduncles short and stout, appreciably broadened distally; ocular acicles narrowly to ovately triangular, slightly less to slightly more than 0.5 peduncular lengths; each terminating acutely or with prominent terminal spine. Antennular peduncles overreaching distal margins of corneas by 0.2-0.5 lengths of penultimate peduncular segments. Ultimate segment with 1-5 long setae at dorsodistal margin. Antennal peduncles overreaching distal corneal margins by 0.3-0.5 lengths of ultimate segments; second segment with dorsolateral distal angle produced, with acute terminal spine and often 1, occasionally 2, smaller spines on lateral margin subterminally; first segment with small to prominent spine on laterodistal margin; antennal acicle moderately slender, reaching to or beyond distal margin of fourth peduncular segment.
</paragraph>
<caption id="1EFA46AED4000C276E26FAE1B0332AC8" ID-DOI="http://doi.org/10.5281/zenodo.4525846" ID-Zenodo-Dep="4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906" startId="7.[113,124,1349,1366]" targetBox="[126,1187,216,1301]" targetPageId="7">
<paragraph id="4A3A1626D4000C276E26FAE1B0332AC8" blockId="7.[113,1211,1348,1457]" pageId="7" pageNumber="906">
FIG. 2. — 
<taxonomicName id="8D856DA5D4000C276E92FAE0B17E2A2F" authority="McLaughlin, 2007" authorityName="McLaughlin" authorityYear="2007" box="[197,590,1348,1366]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="906" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D4000C276E92FAE0B2802A2C" box="[197,432,1348,1365]" italics="true" pageId="7" pageNumber="906">Pteropagurus spinulocarpus</emphasis>
<bibRefCitation id="2E146BD7D4000C276FE0FAE1B17E2A2F" author="MCLAUGHLIN P. A." box="[439,590,1349,1366]" pageId="7" pageNumber="906" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin, 2007</bibRefCitation>
</taxonomicName>
from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): 
<emphasis id="78F1CA34D4000C276A27FAE0B74D2A2C" bold="true" box="[1136,1149,1348,1365]" pageId="7" pageNumber="906">A</emphasis>
, shield and cephalic appendages, dorsal view (aesthetascs omitted); 
<emphasis id="78F1CA34D4000C276C26FAFFB14E2A15" bold="true" box="[625,638,1371,1388]" pageId="7" pageNumber="906">B</emphasis>
, left mandible, outer view; 
<emphasis id="78F1CA34D4000C276D0BFAFFB05A2A15" bold="true" box="[860,874,1371,1388]" pageId="7" pageNumber="906">C</emphasis>
, left maxillule, outer view; 
<emphasis id="78F1CA34D4000C276A13FAFFB7622A15" bold="true" box="[1092,1106,1371,1388]" pageId="7" pageNumber="906">D</emphasis>
, left maxilla, outer view; 
<emphasis id="78F1CA34D4000C276E87FAD6B3EC2AFA" bold="true" box="[208,220,1394,1411]" pageId="7" pageNumber="906">E</emphasis>
, left first maxilliped, outer view; 
<emphasis id="78F1CA34D4000C276FB4FAD6B2DE2AFA" bold="true" box="[483,494,1394,1411]" pageId="7" pageNumber="906">F</emphasis>
, left second maxilliped,outer view; 
<emphasis id="78F1CA34D4000C276D43FAD6B0122AFA" bold="true" box="[788,802,1394,1411]" pageId="7" pageNumber="906">G</emphasis>
, left third maxilliped, outer view; 
<emphasis id="78F1CA34D4000C276A78FAD6B70D2AFA" bold="true" box="[1071,1085,1394,1411]" pageId="7" pageNumber="906">H</emphasis>
, ischium of left third maxilliped, inner view; 
<emphasis id="78F1CA34D4000C276F0EFA2DB26F2AE3" bold="true" box="[345,351,1417,1434]" pageId="7" pageNumber="906">I</emphasis>
, 
<emphasis id="78F1CA34D4000C276F3EFA2DB2432AE3" bold="true" box="[361,371,1417,1434]" pageId="7" pageNumber="906">J</emphasis>
, sternites of second and third pereopods, ventral view; 
<emphasis id="78F1CA34D4000C276D14FA2DB0602AE3" bold="true" box="[835,848,1417,1434]" pageId="7" pageNumber="906">K</emphasis>
, zoeal telson, dorsal view; 
<emphasis id="78F1CA34D4000C276A66FA2DB70E2AE3" bold="true" box="[1073,1086,1417,1434]" pageId="7" pageNumber="906">A</emphasis>
, 
<emphasis id="78F1CA34D4000C276A1FFA2DB77E2AE3" bold="true" box="[1096,1102,1417,1434]" pageId="7" pageNumber="906">I</emphasis>
, ♂ (1.6 mm); 
<emphasis id="78F1CA34D4000C276E26FA3BB3A42AC9" bold="true" box="[113,148,1439,1457]" pageId="7" pageNumber="906">B -H</emphasis>
, ♂ (1.7 mm); 
<emphasis id="78F1CA34D4000C276F52FA3BB23F2AC9" bold="true" box="[261,271,1439,1456]" pageId="7" pageNumber="906">J</emphasis>
, ♀ (1.7 mm); 
<emphasis id="78F1CA34D4000C276F28FA3BB2BC2AC9" bold="true" box="[383,396,1439,1456]" pageId="7" pageNumber="906">K</emphasis>
, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm.
</paragraph>
</caption>
<paragraph id="4A3A1626D4000C286EDFF98EB2722A9F" blockId="7.[113,649,1515,1700]" lastBlockId="8.[111,647,1451,1700]" lastPageId="8" lastPageNumber="907" pageId="7" pageNumber="906">
Mandible (
<figureCitation id="D2BE0AA3D4000C276EACF98EB279293C" box="[251,329,1578,1605]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2B</figureCitation>
) with 2-segmented palp. Maxillule (
<figureCitation id="D2BE0AA3D4000C276EEBF9EEB225291C" box="[188,277,1610,1637]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2C</figureCitation>
) with external lobe of endopod obsolete, internal lobe with 1 seta; basial endite with very slender tooth-like marginal processes. Maxilla (
<figureCitation id="D2BE0AA3D4000C276D54FA4FB066297F" box="[771,854,1515,1542]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2D</figureCitation>
) with endopod slightly overreaching distal margin of scaphognathite. First maxilliped (
<figureCitation id="D2BE0AA3D4000C276CFDF98EB1C9293C" box="[682,761,1578,1605]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2E</figureCitation>
) with endopod overreaching basial endite; slender exopod with few marginal setae. Second maxilliped (
<figureCitation id="D2BE0AA3D4000C276D71F9CDB04429FD" box="[806,884,1641,1668]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2F</figureCitation>
) with very long exopod. Third maxilliped (
<figureCitation id="D2BE0AA3D4000C276D7CF92DB0B429DD" box="[811,900,1673,1700]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="7" pageNumber="906">Fig. 2G</figureCitation>
) with basis-ischium almost entirely fused; crista dentata (
<figureCitation id="D2BE0AA3D40F0C286FF9FA0FB1342ABF" box="[430,516,1451,1478]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="8" pageNumber="907">Fig. 2H</figureCitation>
) reduced to few very tiny teeth.
</paragraph>
<caption id="1EFA46AED40F0C286E26FB48B0262A16" ID-DOI="http://doi.org/10.5281/zenodo.4525848" ID-Zenodo-Dep="4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="8" pageNumber="907" startId="8.[113,124,1260,1277]" targetBox="[113,1209,217,1217]" targetPageId="8">
<paragraph id="4A3A1626D40F0C286E26FB48B0262A16" blockId="8.[112,1211,1260,1391]" pageId="8" pageNumber="907">
FIG. 3. — 
<taxonomicName id="8D856DA5D40F0C286E96FB48B1762B84" authority="McLaughlin, 2007" authorityName="McLaughlin" authorityYear="2007" box="[193,582,1260,1277]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="907" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40F0C286E96FB48B2992B84" box="[193,425,1260,1277]" italics="true" pageId="8" pageNumber="907">Pteropagurus spinulocarpus</emphasis>
<bibRefCitation id="2E146BD7D40F0C286FF8FB48B1762B84" author="MCLAUGHLIN P. A." box="[431,582,1260,1277]" pageId="8" pageNumber="907" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin, 2007</bibRefCitation>
</taxonomicName>
, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): 
<emphasis id="78F1CA34D40F0C286A79FB48B70B2B84" bold="true" box="[1070,1083,1260,1277]" pageId="8" pageNumber="907">A</emphasis>
, 
<emphasis id="78F1CA34D40F0C286A12FB48B7622B84" bold="true" box="[1093,1106,1260,1277]" pageId="8" pageNumber="907">B</emphasis>
, carpus and chela of right cheliped, dorsal view (setae omitted); 
<emphasis id="78F1CA34D40F0C286C77FAA6B11E2A6A" bold="true" box="[544,558,1282,1299]" pageId="8" pageNumber="907">C</emphasis>
, 
<emphasis id="78F1CA34D40F0C286C6FFAA6B1762A6A" bold="true" box="[568,582,1282,1299]" pageId="8" pageNumber="907">D</emphasis>
, carpus and chela of left chela, dorsal view (setae omitted); 
<emphasis id="78F1CA34D40F0C286A6BFAA6B7782A6A" bold="true" box="[1084,1096,1282,1299]" pageId="8" pageNumber="907">E</emphasis>
, 
<emphasis id="78F1CA34D40F0C286A04FAA6B76E2A6A" bold="true" box="[1107,1118,1282,1299]" pageId="8" pageNumber="907">F</emphasis>
, left chela, mesial view (setae omitted); 
<emphasis id="78F1CA34D40F0C286F0FFABDB2562A53" bold="true" box="[344,358,1305,1322]" pageId="8" pageNumber="907">G</emphasis>
, right second pereopod,lateral view (setae omitted); 
<emphasis id="78F1CA34D40F0C286D40FABDB0152A53" bold="true" box="[791,805,1305,1322]" pageId="8" pageNumber="907">H</emphasis>
, right third pereopod,lateral view (setae omitted); 
<emphasis id="78F1CA34D40F0C286E27FA94B3462A38" bold="true" box="[112,118,1328,1345]" pageId="8" pageNumber="907">I</emphasis>
, dactyl of left third pereopod, mesial view; 
<emphasis id="78F1CA34D40F0C286F89FA94B2D82A38" bold="true" box="[478,488,1328,1345]" pageId="8" pageNumber="907">J</emphasis>
, distal three segments of left fourth pereopod, lateral view (setae omitted); 
<emphasis id="78F1CA34D40F0C286A0DFA94B7572A38" bold="true" box="[1114,1127,1328,1345]" pageId="8" pageNumber="907">K</emphasis>
, 
<emphasis id="78F1CA34D40F0C286A26FA94B74C2A38" bold="true" box="[1137,1148,1328,1345]" pageId="8" pageNumber="907">L</emphasis>
, coxae and sternite of fifth pereopods, ventral view; 
<emphasis id="78F1CA34D40F0C286FBDFAE2B2CB2A2E" bold="true" box="[490,507,1350,1367]" pageId="8" pageNumber="907">M</emphasis>
, 
<emphasis id="78F1CA34D40F0C286C51FAE2B1242A2E" bold="true" box="[518,532,1350,1367]" pageId="8" pageNumber="907">N</emphasis>
, telson, dorsal view; 
<emphasis id="78F1CA34D40F0C286C90FAE2B1E62A2E" bold="true" box="[711,726,1350,1367]" pageId="8" pageNumber="907">O</emphasis>
, cluster of stalked protozoans from shell interior; 
<emphasis id="78F1CA34D40F0C286A20FAE2B7B42A2E" bold="true" box="[1143,1156,1350,1367]" pageId="8" pageNumber="907">A</emphasis>
, 
<emphasis id="78F1CA34D40F0C286AD8FAE2B7AD2A2E" bold="true" box="[1167,1181,1350,1367]" pageId="8" pageNumber="907">C</emphasis>
, 
<emphasis id="78F1CA34D40F0C286AFFFAE2B7842A2E" bold="true" box="[1192,1204,1350,1367]" pageId="8" pageNumber="907">E</emphasis>
, 
<emphasis id="78F1CA34D40F0C286E26FAF9B3A42A17" bold="true" box="[113,148,1373,1391]" pageId="8" pageNumber="907">G -K</emphasis>
, 
<emphasis id="78F1CA34D40F0C286EC8FAF9B3802A17" bold="true" box="[159,176,1373,1390]" pageId="8" pageNumber="907">M</emphasis>
, ♂ (1.6 mm); 
<emphasis id="78F1CA34D40F0C286F76FAF9B21E2A17" bold="true" box="[289,302,1373,1390]" pageId="8" pageNumber="907">B</emphasis>
, 
<emphasis id="78F1CA34D40F0C286F6FFAF9B2762A17" bold="true" box="[312,326,1373,1390]" pageId="8" pageNumber="907">D</emphasis>
, 
<emphasis id="78F1CA34D40F0C286F06FAF9B26C2A17" bold="true" box="[337,348,1373,1390]" pageId="8" pageNumber="907">F</emphasis>
, 
<emphasis id="78F1CA34D40F0C286F31FAF9B2412A17" bold="true" box="[358,369,1373,1390]" pageId="8" pageNumber="907">L</emphasis>
, 
<emphasis id="78F1CA34D40F0C286F2BFAF9B2BA2A17" bold="true" box="[380,394,1373,1390]" pageId="8" pageNumber="907">N</emphasis>
, ♀ (1.7 mm); 
<emphasis id="78F1CA34D40F0C286FAEFAF9B1382A17" bold="true" box="[505,520,1373,1390]" pageId="8" pageNumber="907">O</emphasis>
, from shell. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="4A3A1626D40F0C296EDFFA4FB2CC2E0B" blockId="8.[111,647,1451,1700]" lastBlockId="9.[111,649,216,1700]" lastPageId="9" lastPageNumber="908" pageId="8" pageNumber="907">
Right cheliped (
<figureCitation id="D2BE0AA3D40F0C286F66FA4FB2AF297F" box="[305,415,1515,1542]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="8" pageNumber="907">Fig. 3A, B</figureCitation>
) slightly longer to appreciably shorter than left, but considerably stouter; dactyl and fixed finger with or without slight hiatus. Dactyl 0.6-0.7 length of palm; dorsomesial margin rounded, all surfaces unarmed, but with few scattered setae, particularly ventrally; cutting edge with 2 or 3 moderate to large calcareous teeth. Palm slightly shorter to slightly longer than carpus; dorsomesial and dorsolateral margins rounded, unarmed, convex dorsal and ventral surfaces also unarmed. Carpus slightly longer than merus; dorsomesial and dorsolateral margins each with row of 4-9 small spines, number increasing with increased animal size, other surfaces unarmed, but distomesial margin often with 2 or 3 smaller spines. Merus glabrous or with row of widely-spaced, individual bristles on dorsal surface; other surfaces unarmed but with few scattered setae; ventromesial and ventrolateral distal angles each usually with small spine.
</paragraph>
<paragraph id="4A3A1626D40E0C296EDFFED2B2462C74" blockId="9.[111,649,216,1700]" pageId="9" pageNumber="908">
Left cheliped (
<figureCitation id="D2BE0AA3D40E0C296F49FED2B2682EE8" box="[286,344,374,401]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3</figureCitation>
C-F) long, slender, considerably longer than right with tips prominently curved in larger male specimens; dactyl and fixed finger upwardly flexed (
<figureCitation id="D2BE0AA3D40E0C296F79FE71B24F2E89" box="[302,383,469,496]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3F</figureCitation>
) and with slight hiatus; dactyl shorter to approximately 1.2 length of palm; surfaces all unarmed but with sparse scattering of short setae particularly ventrally. Palm long, slen- der, surfaces unarmed but with few scattered setae. Carpus approximately equal to length of merus; dorsomesial and dorsolateral margins each with row of small spines increasing in number with increasing body size, other surfaces unarmed but with few scattered setae.
</paragraph>
<paragraph id="4A3A1626D40E0C296EDFFCB6B2682A1E" blockId="9.[111,649,216,1700]" pageId="9" pageNumber="908">
Second (
<figureCitation id="D2BE0AA3D40E0C296EB7FCB6B2002C54" box="[224,304,786,813]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig.3G</figureCitation>
) and third (
<figureCitation id="D2BE0AA3D40E0C296FFEFCB6B13F2C54" box="[425,527,786,813]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3H, I</figureCitation>
) pereopods similar in armature, but dissimilar in length and proportions, third appreciably longer and stouter. Dactyls slightly shorter to approximately equal to lengths of propodi; terminating in small corneous claws; mesial faces each with row of widely-spaced small bristles or corneous spinules adjacent to ventral margin (
<figureCitation id="D2BE0AA3D40E0C296F70FC54B2442B72" box="[295,372,1008,1035]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3I</figureCitation>
). Propodi approximately twice length of carpi, unarmed or with very low protuberances and sparse row of setae on each dorsal surface. Carpi each with row of small spines on dorsal surface. Meri of second pereopods each usually with dorsodistal spine, occasionally unarmed; third each with 1 dorsodistal and 1-3 subdistal spinulose protuberances or small spines and 1 or 2 setae, ventrolateral distal angles each with tiny spinule or small spine. Fourth pereopods (
<figureCitation id="D2BE0AA3D40E0C296C60FAA9B1B02A51" box="[567,640,1293,1320]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3J</figureCitation>
) semichelate; propodal rasp with 1 row of 3-5 spiniform corneous scales.
</paragraph>
<paragraph id="4A3A1626D40E0C296EDFFAC8B2E1295C" blockId="9.[111,649,216,1700]" pageId="9" pageNumber="908">
Sternites of second and third pereopods (
<figureCitation id="D2BE0AA3D40E0C296C72FAC8B14F2AFE" box="[549,639,1388,1415]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="9" pageNumber="908">Fig. 2I, J</figureCitation>
) broad; third broadly subrectangular anteriorly and drawn out posteriorly into broad, terminally rounded plate in both sexes. Sternite of fifth pereopods (
<figureCitation id="D2BE0AA3D40E0C296E95FA4FB204297F" box="[194,308,1515,1542]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3K, L</figureCitation>
) with simple or faintly bilobed median projection in both sexes.
</paragraph>
<paragraph id="4A3A1626D40E0C296EDFF98EB1B829DD" blockId="9.[111,649,216,1700]" pageId="9" pageNumber="908">
Male with coxae of fifth pereopods approximately equal, right with long, stout sexual tube developed as posterior coxal extension (
<figureCitation id="D2BE0AA3D40E0C296C7DF9CDB1B029FD" box="[554,640,1641,1668]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3K</figureCitation>
) directed posteriorly and exteriorly and curving over
</paragraph>
<paragraph id="4A3A1626D40E0C296CF4FF7CB0972E2B" blockId="9.[675,1212,216,496]" pageId="9" pageNumber="908">
anterior portion of dorsal pleon; coxa of left with gonopore and sometimes papilla. Coxae of fifth pereopods (
<figureCitation id="D2BE0AA3D40E0C296D70FEB3B04B2E4B" box="[807,891,279,306]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3L</figureCitation>
) in females each with dense setae on ventral surface.
</paragraph>
<paragraph id="4A3A1626D40E0C296CECFEF3B0242E89" blockId="9.[675,1212,216,496]" pageId="9" pageNumber="908">
Telson (
<figureCitation id="D2BE0AA3D40E0C296D5AFEF3B0B32E0B" box="[781,899,343,370]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3M, N</figureCitation>
) with V- or U-shaped median cleft, or only slight median concavity; terminal margins oblique, each with 2-4 tiny blunt or acute spines; lateral margins each with 1 or 2 moderately long setae.
</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FDB1B0232D56" blockId="9.[677,794,533,591]" box="[677,787,533,559]" pageId="9" pageNumber="908">
<emphasis id="78F1CA34D40E0C296CF2FDB1B0232D56" box="[677,787,533,559]" italics="true" pageId="9" pageNumber="908">Coloration</emphasis>
</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FD90B02A2D36" blockId="9.[677,794,533,591]" box="[677,794,564,591]" pageId="9" pageNumber="908">Unknown.</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FDD0B1C72DF7" blockId="9.[677,1210,628,686]" box="[677,759,628,654]" pageId="9" pageNumber="908">
<emphasis id="78F1CA34D40E0C296CF2FDD0B1C72DF7" box="[677,759,628,654]" italics="true" pageId="9" pageNumber="908">Habitat</emphasis>
</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FD37B78A2DD7" blockId="9.[677,1210,628,686]" box="[677,1210,659,686]" pageId="9" pageNumber="908">
Empty shells of 
<taxonomicName id="8D856DA5D40E0C296D1DFD30B7862DD7" authority="(Rang, 1827)" baseAuthorityName="Rang" baseAuthorityYear="1827" box="[842,1206,659,686]" class="Gastropoda" family="Cuvierinidae" genus="Cuvierina" kingdom="Animalia" order="Thecosomata" pageId="9" pageNumber="908" phylum="Mollusca" rank="species" species="columnella">
<emphasis id="78F1CA34D40E0C296D1DFD30B7122DD4" box="[842,1058,659,686]" italics="true" pageId="9" pageNumber="908">Cuvierina columnella</emphasis>
(Rang, 1827)
</taxonomicName>
.
</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FD77B05F2D94" blockId="9.[676,1212,723,1067]" box="[677,879,723,749]" pageId="9" pageNumber="908">
<emphasis id="78F1CA34D40E0C296CF2FD77B05F2D94" box="[677,879,723,749]" italics="true" pageId="9" pageNumber="908">Larval development</emphasis>
</paragraph>
<paragraph id="4A3A1626D40E0C296CF2FD56B0762B52" blockId="9.[676,1212,723,1067]" pageId="9" pageNumber="908">
Only limited information available from prematurely hatched zoeae. Carapace with no rostral spine apparent; posterolateral angles rounded. Appendages typical of first stage zoeae, but none with setae extruded. Pleon with 5 somites and telson; posterior telsonal margin (
<figureCitation id="D2BE0AA3D40E0C296A75FC35B7492CD5" box="[1058,1145,913,940]" captionStart="FIG" captionStartId="7.[113,124,1349,1366]" captionTargetBox="[126,1187,216,1301]" captionTargetId="figure-42@7.[93,1230,195,1324]" captionTargetPageId="7" captionText="FIG. 2. — Pteropagurus spinulocarpus McLaughlin, 2007 from EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, shield and cephalic appendages, dorsal view (aesthetascs omitted);B, left mandible, outer view; C, left maxillule, outer view; D, left maxilla, outer view; E, left first maxilliped, outer view;F, left second maxilliped,outer view; G, left third maxilliped, outer view;H, ischium of left third maxilliped, inner view; I, J, sternites of second and third pereopods, ventral view; K, zoeal telson, dorsal view; A, I, ♂ (1.6 mm); B-H, ♂ (1.7 mm); J, ♀ (1.7 mm); K, zoea. Scale bars: A-J, 0.5 mm; K, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4525846" httpUri="https://zenodo.org/record/4525846/files/figure.png" pageId="9" pageNumber="908">Fig. 2K</figureCitation>
) with small U-shaped median cleft and 7+7 processes: outermost – small spine, second – anomuran hair, third through seventh – plumodenticulate setae, fourth longest.
</paragraph>
</subSubSection>
<subSubSection id="029F45ADD40E0C2A6CF2FBEBB2E329DD" lastPageId="10" lastPageNumber="909" pageId="9" pageNumber="908" type="discussion">
<paragraph id="4A3A1626D40E0C296CF2FBEBB0392B1E" blockId="9.[675,1211,1103,1700]" box="[677,777,1103,1129]" pageId="9" pageNumber="908">REMARKS</paragraph>
<paragraph id="4A3A1626D40E0C2A6CF2FBCBB2B52D36" blockId="9.[675,1211,1103,1700]" lastBlockId="10.[111,648,216,1700]" lastPageId="10" lastPageNumber="909" pageId="9" pageNumber="908">
The mouthparts of 
<taxonomicName id="8D856DA5D40E0C296D2EFBCBB72F2BF1" authorityName="McLaughlin" authorityYear="2007" box="[889,1055,1134,1160]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40E0C296D2EFBCBB72F2BF1" box="[889,1055,1134,1160]" italics="true" pageId="9" pageNumber="908">P. spinulocarpus</emphasis>
</taxonomicName>
are in general agreement with those described for 
<taxonomicName id="8D856DA5D40E0C296A1CFB2BB7892BD1" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[1099,1209,1167,1192]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="78F1CA34D40E0C296A1CFB2BB7892BD1" box="[1099,1209,1167,1192]" italics="true" pageId="9" pageNumber="908">P. inermis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="2E146BD7D40E0C296C9FFB0AB72C2BB0" author="MCLAUGHLIN P. A. &amp; RAHAYU D. L." box="[712,1052,1198,1225]" pageId="9" pageNumber="908" pagination="55 - 68" refId="ref10933" refString="MCLAUGHLIN P. A. &amp; RAHAYU D. L. 2006. - A new genus with two new species of hermit crabs (Crustacea, Decapoda, Paguroidea, Paguridae) from an unique habitat. Zootaxa 1116: 55 - 68." type="journal article" year="2006">McLaughlin &amp; Rahayu (2006)</bibRefCitation>
as representative of the genus. However, the crista dentata of 
<taxonomicName id="8D856DA5D40E0C296CF1FB4AB0682A7E" authorityName="McLaughlin" authorityYear="2007" box="[678,856,1261,1287]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40E0C296CF1FB4AB0682A7E" box="[678,856,1261,1287]" italics="true" pageId="9" pageNumber="908">P. spinulocarpus</emphasis>
</taxonomicName>
consists of considerably fewer teeth than were illustrated but not specified for 
<taxonomicName id="8D856DA5D40E0C296CF2FA89B03C2A3F" authorityName="McLaughlin &amp; Rahayu" authorityYear="2006" box="[677,780,1325,1350]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="78F1CA34D40E0C296CF2FA89B03C2A3F" box="[677,780,1325,1350]" italics="true" pageId="9" pageNumber="908">P. inermis</emphasis>
</taxonomicName>
. Although the right chelipeds of males of 
<taxonomicName id="8D856DA5D40E0C296C97FAE9B0512A1F" authorityName="McLaughlin" authorityYear="2007" box="[704,865,1356,1382]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40E0C296C97FAE9B0512A1F" box="[704,865,1356,1382]" italics="true" pageId="9" pageNumber="908">P. spinulocarpus</emphasis>
</taxonomicName>
are generally a little stouter than females, both are very similar morphologically and do not change with growth. 
<bibRefCitation id="2E146BD7D40E0C296D82FA28B7882ADE" author="MCLAUGHLIN P. A." box="[981,1208,1420,1447]" pageId="9" pageNumber="908" pagination="503 - 513" refId="ref10884" refString="MCLAUGHLIN P. A. 2007. - A new species of Pteropagurus McLaughlin &amp; Rahayu, 2006 and a new genus and species of scaphopod dweller (Crustacea, Decapoda, Anomura, Paguroidea, Paguridae). Zoosystema 29 (3): 503 - 513." type="journal article" year="2007">McLaughlin’s (2007)</bibRefCitation>
description of the left chela being ventrally curved in both sexes is misleading. In lateral or mesial view, the dactyl and fixed finger are flexed upward in relation to the ventral margin of the palm (
<figureCitation id="D2BE0AA3D40E0C296A31F9AEB783295C" box="[1126,1203,1546,1573]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="9" pageNumber="908">Fig. 3F</figureCitation>
) with the ventral margins of the former slightly curved. This flexion readily distinguishes the left cheliped of 
<taxonomicName id="8D856DA5D40E0C296D7BF9CEB0E829FA" authorityName="McLaughlin" authorityYear="2007" box="[812,984,1641,1667]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="908" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40E0C296D7BF9CEB0E829FA" box="[812,984,1641,1667]" italics="true" pageId="9" pageNumber="908">P. spinulocarpus</emphasis>
</taxonomicName>
from the otherwise structurally similar and sexually dimorphic left cheliped of 
<taxonomicName id="8D856DA5D40D0C2A6EA0FF7CB2A02F8B" authorityName="de Saint Laurent &amp; McLaughlin" authorityYear="2000" box="[247,400,216,243]" class="Malacostraca" family="Paguridae" genus="Catapagurus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinicarpus">
<emphasis id="78F1CA34D40D0C2A6EA0FF7CB2392F8B" box="[247,265,216,242]" italics="true" pageId="10" pageNumber="909">C</emphasis>
. 
<emphasis id="78F1CA34D40D0C2A6F4DFF7DB2A02F8B" box="[282,400,217,242]" italics="true" pageId="10" pageNumber="909">spinicarpus</emphasis>
</taxonomicName>
. Nonetheless, the left chelipeds of 
<taxonomicName id="8D856DA5D40D0C2A6EA8FF5CB2982E68" authorityName="McLaughlin" authorityYear="2007" box="[255,424,247,273]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40D0C2A6EA8FF5CB2982E68" box="[255,424,247,273]" italics="true" pageId="10" pageNumber="909">P. spinulocarpus</emphasis>
</taxonomicName>
exhibit comparable sexual dimorphism to that seen in 
<taxonomicName id="8D856DA5D40D0C2A6FBCFEBCB14F2E48" authorityName="de Saint Laurent &amp; McLaughlin" authorityYear="2000" box="[491,639,280,306]" class="Malacostraca" family="Paguridae" genus="Catapagurus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinicarpus">
<emphasis id="78F1CA34D40D0C2A6FBCFEBCB14F2E48" box="[491,639,280,306]" italics="true" pageId="10" pageNumber="909">C. spinicarpus</emphasis>
</taxonomicName>
, such as an increase in length with increased animal size and the prominent ventral curvature of the distal portions of the dactyl and fixed finger, which accompanies the increase in overall length in males (
<figureCitation id="D2BE0AA3D40D0C2A6E80FE12B2792EA8" box="[215,329,438,465]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="10" pageNumber="909">Fig. 3C, E</figureCitation>
), but not in females (
<figureCitation id="D2BE0AA3D40D0C2A6C7CFE12B3B42E89" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="10" pageNumber="909">Fig. 3D, F</figureCitation>
). Sexual dimorphism is exhibited by females of 
<taxonomicName id="8D856DA5D40D0C2A6E25FE52B2212D76" authorityName="McLaughlin" authorityYear="2007" box="[114,273,501,527]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40D0C2A6E25FE52B2212D76" box="[114,273,501,527]" italics="true" pageId="10" pageNumber="909">P. spinulocarpus</emphasis>
</taxonomicName>
in the presence of moderately dense setation on the ventral surfaces of the coxae of the fifth pereopods (
<figureCitation id="D2BE0AA3D40D0C2A6F72FD90B2482D36" box="[293,376,564,591]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="10" pageNumber="909">Fig. 3L</figureCitation>
).
</paragraph>
<paragraph id="4A3A1626D40D0C2A6EDFFDF0B1052C54" blockId="10.[111,648,216,1700]" pageId="10" pageNumber="909">
The increases in the number of spines on the dorsal surfaces of the carpi of the chelipeds and ambulatory legs occur with increased body size in both sexes and appear to be entirely functions of growth. The variations seen in the breadth, median cleft, or armature of the telson (
<figureCitation id="D2BE0AA3D40D0C2A6FEEFD56B1012C74" box="[441,561,754,781]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="10" pageNumber="909">Fig. 3M, N</figureCitation>
) do not appear to be dimorphic or growth related.
</paragraph>
<paragraph id="4A3A1626D40D0C2A6EDFFC96B2E329DD" blockId="10.[111,648,216,1700]" pageId="10" pageNumber="909">
<bibRefCitation id="2E146BD7D40D0C2A6EDFFC96B2EA2C34" author="MCLAUGHLIN P. A. &amp; RAHAYU D. L." box="[136,474,818,845]" pageId="10" pageNumber="909" pagination="55 - 68" refId="ref10933" refString="MCLAUGHLIN P. A. &amp; RAHAYU D. L. 2006. - A new genus with two new species of hermit crabs (Crustacea, Decapoda, Paguroidea, Paguridae) from an unique habitat. Zootaxa 1116: 55 - 68." type="journal article" year="2006">McLaughlin &amp; Rahayu (2006)</bibRefCitation>
speculated that the longer third pereopods of 
<taxonomicName id="8D856DA5D40D0C2A6FE7FCF6B1072C12" authorityName=", McLaughlin &amp; Rahayu" authorityYear="2006" box="[432,567,850,875]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="genus">
<emphasis id="78F1CA34D40D0C2A6FE7FCF6B1072C12" box="[432,567,850,875]" italics="true" pageId="10" pageNumber="909">Pteropagurus</emphasis>
</taxonomicName>
species enabled the animals to elevate the shells above the substrate when walking, as none of the four shells examined in their original sample had any marks on the exterior surfaces. The numerous carcinoecia of 
<taxonomicName id="8D856DA5D40D0C2A6EC7FC55B20D2B73" authorityName="McLaughlin" authorityYear="2007" box="[144,317,1008,1035]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40D0C2A6EC7FC55B3902B73" box="[144,160,1009,1034]" italics="true" pageId="10" pageNumber="909">P</emphasis>
. 
<emphasis id="78F1CA34D40D0C2A6EE7FC54B20D2B73" box="[176,317,1008,1034]" italics="true" pageId="10" pageNumber="909">spinulocarpus</emphasis>
</taxonomicName>
in the recent sample support their supposition. The external shell surfaces are clear and smooth, showing no signs of scuffing from being dragged across the substrate. The apertures are usually smooth and unbroken. In the preserved state, the appendages of the hermits often were at least partially extended; however, the entire animals and their appendages could be retracted completely into the carcinoecia. On the internal surfaces of several of the pteropod shells occupied by 
<taxonomicName id="8D856DA5D40D0C2A6EA8FA89B2952A3E" authorityName="McLaughlin" authorityYear="2007" box="[255,421,1325,1351]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40D0C2A6EA8FA89B2952A3E" box="[255,421,1325,1351]" italics="true" pageId="10" pageNumber="909">P. spinulocarpus</emphasis>
</taxonomicName>
were what appeared to be clusters of a stalked protozoan (
<figureCitation id="D2BE0AA3D40D0C2A6C73FAE8B14E2A1E" box="[548,638,1356,1383]" captionStart="FIG" captionStartId="8.[113,124,1260,1277]" captionTargetBox="[113,1209,217,1217]" captionTargetId="figure-38@8.[85,1222,188,1241]" captionTargetPageId="8" captionText="FIG. 3. — Pteropagurus spinulocarpus McLaughlin, 2007, EBISCO, stn DW 2492, New Caledonia (MNHN-Pg 7764): A, B, carpus and chela of right cheliped, dorsal view (setae omitted); C, D, carpus and chela of left chela, dorsal view (setae omitted); E, F, left chela, mesial view (setae omitted);G, right second pereopod,lateral view (setae omitted);H, right third pereopod,lateral view (setae omitted); I, dactyl of left third pereopod, mesial view; J, distal three segments of left fourth pereopod, lateral view (setae omitted); K, L, coxae and sternite of fifth pereopods, ventral view; M, N, telson, dorsal view; O, cluster of stalked protozoans from shell interior; A, C, E, G-K, M, ♂ (1.6 mm); B, D, F, L, N, ♀ (1.7 mm); O, from shell. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4525848" httpUri="https://zenodo.org/record/4525848/files/figure.png" pageId="10" pageNumber="909">Fig. 3O</figureCitation>
) probably of the phylum Ciliophora. Although 
<bibRefCitation id="2E146BD7D40D0C2A6E23FA28B2D12ADE" author="WILLIAMS J. D. &amp; MCDERMOTT J. J." box="[116,481,1420,1447]" pageId="10" pageNumber="909" pagination="11 - 128" refId="ref11252" refString="WILLIAMS J. D. &amp; MCDERMOTT J. J. 2004. - Hermit crab biocoenoses: a worldwide review of the diversity and natural history of hermit crab associates. Journal of Experimental Marine Biology and Ecology 305: 11 - 128." type="journal article" year="2004">Williams &amp; McDermott (2004)</bibRefCitation>
indicated that protozoan attachments included shell surfaces and other symbionts as well as the exoskeletons of the crabs, Fernandez-Leborans &amp; Tato-Porto (2000a, b) and Fernandez-Leborans (2001) reported such attachments only on the gills, integuments, and setae of several hermit crabs. Clusters similar to those on the shell walls occasionally also were observed on the bodies of 
<taxonomicName id="8D856DA5D40D0C2A6F7FF92EB2FC29DA" authorityName="McLaughlin" authorityYear="2007" box="[296,460,1673,1699]" class="Malacostraca" family="Paguridae" genus="Pteropagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="909" phylum="Arthropoda" rank="species" species="spinulocarpus">
<emphasis id="78F1CA34D40D0C2A6F7FF92EB2FC29DA" box="[296,460,1673,1699]" italics="true" pageId="10" pageNumber="909">P. spinulocarpus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>