448 lines
55 KiB
XML
448 lines
55 KiB
XML
<document id="BAFE8D28928AA9C698DBF6CB99798A37" ID-CLB-Dataset="3953" ID-DOI="10.5281/zenodo.1196003" ID-GBIF-Dataset="673e368f-06c4-465a-b40d-68dbedd81138" ID-ISSN="0035-418" ID-Zenodo-Dep="1196003" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634783046189" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N." docDate="2018" docId="BC2887A2FFA199215DA0F0454EC1F975" docLanguage="en" docName="1_Anisyutkin_1.pdf" docOrigin="Revue suisse de Zoologie 125 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Morphnodes Hebard 1930" docType="treatment" docVersion="6" lastPageNumber="9" masterDocId="4011FFDAFFA599295E10F6034F12FF98" masterDocTitle="Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d'histoire naturelle de Genève. Part 3" masterLastPageNumber="16" masterPageNumber="1" pageNumber="5" updateTime="1699460453851" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="633EFECE78FFE92FE4F539B08BB71D62">
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<mods:title id="8DC868BB3A30F8596468CDC6D9BD00CA">Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d'histoire naturelle de Genève. Part 3</mods:title>
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<mods:namePart id="2C9EA199BD1CA48FD06C3E323E889FAE">Anisyutkin, Leonid N.</mods:namePart>
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<mods:affiliation id="6E06032A9E4BF8915901D7635CFA0070">Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia. E-mail: leonid. dictyoptera @ gmail. com, Leonid. Anisyutkin @ zin. ru</mods:affiliation>
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<mods:typeOfResource id="67C2FFA8CA55B57EBFBBAB4BA25AF8F6">text</mods:typeOfResource>
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<mods:title id="4CF4D93FB2F234A426F0C6754F0589A4">Revue suisse de Zoologie</mods:title>
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<mods:part id="5B58AEDE005AB07192B3FEE2E5AF89E0">
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<mods:date id="C0F6EA89C2290B5A9FA0D77478DED3AA">2018</mods:date>
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<mods:number id="3C8FCD6EC2A4E5626D0F9CE9F81E4D9D">2018-03-31</mods:number>
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<mods:number id="733B0BC6E1ED1A5F7FCB8CB932A287AC">125</mods:number>
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<mods:identifier id="9B1FA74B9923571FD28FAF0C13AFD746" type="DOI">10.5281/zenodo.1196003</mods:identifier>
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<treatment id="BC2887A2FFA199215DA0F0454EC1F975" ID-DOI="http://doi.org/10.5281/zenodo.5589900" ID-GBIF-Taxon="189149526" ID-Zenodo-Dep="5589900" LSID="urn:lsid:plazi:treatment:BC2887A2FFA199215DA0F0454EC1F975" httpUri="http://treatment.plazi.org/id/BC2887A2FFA199215DA0F0454EC1F975" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
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<subSubSection id="7C9B653FFFA1992D5DA0F0454A39F9C5" box="[944,1323,1605,1629]" pageId="4" pageNumber="5" type="nomenclature">
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||
<paragraph id="343E36B4FFA1992D5DA0F0454A39F9C5" blockId="4.[944,1323,1605,1629]" box="[944,1323,1605,1629]" pageId="4" pageNumber="5">
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<heading id="6F7681D8FFA1992D5DA0F0454A39F9C5" bold="true" box="[944,1323,1605,1629]" centered="true" level="2" pageId="4" pageNumber="5" reason="2">
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||
<emphasis id="06F5EAA6FFA1992D5DA0F0454A39F9C5" bold="true" box="[944,1323,1605,1629]" pageId="4" pageNumber="5">
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Genus
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<taxonomicName id="F3814D37FFA1992D5DEFF0464A39F9C5" ID-CoL="8N7CJ" authority="Hebard, 1930" authorityName="Hebard" authorityYear="1930" box="[1023,1323,1605,1629]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5DEFF0464B98F9C5" bold="true" box="[1023,1162,1605,1629]" italics="true" pageId="4" pageNumber="5">Morphnodes</emphasis>
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<bibRefCitation id="50104B45FFA1992D5A81F0454A39F9C5" author="Hebard M." box="[1169,1323,1606,1629]" pageId="4" pageNumber="5" pagination="1 - 109" refId="ref10294" refString="Hebard M. 1930. Studies in Malayan Blattidae (Orthoptera). Proceedings of the Academy of Natural Sciences of Philadelphia (1929) 81: 1 - 109." type="journal article" year="1930">Hebard, 1930</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7C9B653FFFA1992D5D2FF0764B0EF934" pageId="4" pageNumber="5" type="type_taxon">
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<paragraph id="343E36B4FFA1992D5D2FF0764B0EF934" blockId="4.[831,1436,1653,1708]" pageId="4" pageNumber="5">
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<emphasis id="06F5EAA6FFA1992D5D2FF0764CF6F914" bold="true" box="[831,996,1653,1676]" pageId="4" pageNumber="5">
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<typeStatus id="EB3A8816FFA1992D5D2FF0764C6AF914" box="[831,888,1653,1676]" pageId="4" pageNumber="5">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="F3814D37FFA1992D5DE7F0764A78F914" ID-CoL="6FT6H" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" box="[1015,1386,1653,1676]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vasta">
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<emphasis id="06F5EAA6FFA1992D5DE7F0764BAAF914" box="[1015,1208,1653,1676]" italics="true" pageId="4" pageNumber="5">Epilampra vasta</emphasis>
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<bibRefCitation id="50104B45FFA1992D5ADBF0764A78F914" author="Walker F." box="[1227,1386,1653,1676]" pageId="4" pageNumber="5" refId="ref10895" refString="Walker F. 1868. Catalogue of the specimens of Blattariae in the collection of the British Museum. British Museum, London, 239 pp." type="book" year="1868">Walker, 1868</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="7C9B653FFFA199215D2FF0C74EC1F975" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="discussion">
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<paragraph id="343E36B4FFA1992C5D2FF0C74D69F863" blockId="4.[831,1436,1732,2077]" lastBlockId="5.[151,756,1887,2077]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<emphasis id="06F5EAA6FFA1992D5D2FF0C74CA0F943" bold="true" box="[831,946,1732,1755]" pageId="4" pageNumber="5">Remarks:</emphasis>
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The genus
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<taxonomicName id="F3814D37FFA1992D5A48F0C74BF3F943" authorityName="Hebard" authorityYear="1930" box="[1112,1249,1732,1755]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5A48F0C74BF3F943" box="[1112,1249,1732,1755]" italics="true" pageId="4" pageNumber="5">Morphnodes</emphasis>
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</taxonomicName>
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was described as closely related to
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<taxonomicName id="F3814D37FFA1992D5A22F0E74B84F963" authorityName="Shelford" authorityYear="1910" box="[1074,1174,1764,1787]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5A22F0E74B84F963" box="[1074,1174,1764,1787]" italics="true" pageId="4" pageNumber="5">Morphna</emphasis>
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</taxonomicName>
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, but differs from it as follows: “Pronotal disk with a weak to moderately developed impression latero-cephalad. Tegmina showing distinct distal truncation. Hind metatarsus biseriately spined ventrad.” (
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<bibRefCitation id="50104B45FFA1992D5A6FF1664A55F8E4" author="Hebard M." box="[1151,1351,1893,1916]" pageId="4" pageNumber="5" pagination="1 - 109" refId="ref10294" refString="Hebard M. 1930. Studies in Malayan Blattidae (Orthoptera). Proceedings of the Academy of Natural Sciences of Philadelphia (1929) 81: 1 - 109." type="journal article" year="1930">Hebard, 1930: 93</bibRefCitation>
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). Later,
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<emphasis id="06F5EAA6FFA1992D5D2FF1864CA8F804" box="[831,954,1925,1948]" italics="true" pageId="4" pageNumber="5">Morpnodes</emphasis>
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was synonymized with
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<taxonomicName id="F3814D37FFA1992D5AF2F1864CD6F824" authority="Kirby, 1903" authorityName="Kirby" authorityYear="1903" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5AF2F1864A8EF804" box="[1250,1436,1925,1948]" italics="true" pageId="4" pageNumber="5">Pseudophoraspis</emphasis>
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<bibRefCitation id="50104B45FFA1992D5D2FF1A64CD6F824" author="Kirby W. F." box="[831,964,1957,1980]" pageId="4" pageNumber="5" pagination="273 - 280" refId="ref10325" refString="Kirby W. F. 1903. Notes on Blattidae & c., with description of new genera and species in the collection of the British Museum, South Kensington. No 2. Annals and Magazine of Natural History (Series 7) 12: 273 - 280." type="journal article" year="1903">Kirby, 1903</bibRefCitation>
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</taxonomicName>
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by
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<bibRefCitation id="50104B45FFA1992D5DE5F1A64B87F824" author="Princis K." box="[1013,1173,1957,1980]" pageId="4" pageNumber="5" pagination="59 - 75" refId="ref10461" refString="Princis K. 1958. Revision der Walkerschen und Kirbyschen Blattarientypen im British Museum of Natural History, London. II. Opuscula entomologica 23: 59 - 75." type="journal article" year="1958">Princis (1958)</bibRefCitation>
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. It was argued that the
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<typeStatus id="EB3A8816FFA1992D5D2FF1C64C7FF844" box="[831,877,1989,2012]" pageId="4" pageNumber="5">type</typeStatus>
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species of
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<taxonomicName id="F3814D37FFA1992D5DE4F1C64B6EF844" authorityName="Hebard" authorityYear="1930" box="[1012,1148,1989,2012]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5DE4F1C64B6EF844" box="[1012,1148,1989,2012]" italics="true" pageId="4" pageNumber="5">Morphnodes</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="F3814D37FFA1992D5A9EF1C64BF8F844" authorityName="Walker" authorityYear="1868" box="[1166,1258,1989,2012]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vasta">
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<emphasis id="06F5EAA6FFA1992D5A9EF1C64B8CF844" box="[1166,1182,1989,2012]" italics="true" pageId="4" pageNumber="5">E</emphasis>
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.
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<emphasis id="06F5EAA6FFA1992D5AA1F1C64BF8F844" box="[1201,1258,1989,2012]" italics="true" pageId="4" pageNumber="5">vasta</emphasis>
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</taxonomicName>
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, belongs to the genus
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<taxonomicName id="F3814D37FFA1992D5D97F1E64B2DF864" authorityName="Kirby" authorityYear="1903" box="[903,1087,2021,2044]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA1992D5D97F1E64B2DF864" box="[903,1087,2021,2044]" italics="true" pageId="4" pageNumber="5">Pseudophoraspis</emphasis>
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</taxonomicName>
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as stated by
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<bibRefCitation id="50104B45FFA1992D5AC1F1E64A92F864" author="Shelford R." box="[1233,1408,2021,2044]" pageId="4" pageNumber="5" pagination="1 - 21" refId="ref10799" refString="Shelford R. 1910. Orthoptera. Fam. Blattidae. Subfam. Epilamprinae. Genera Insectorum 101: 1 - 21." type="journal article" year="1910">Shelford (1910)</bibRefCitation>
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. It must be noted that
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<bibRefCitation id="50104B45FFA1992D5A06FE054BFCF785" author="Shelford R." box="[1046,1262,2054,2077]" pageId="4" pageNumber="5" pagination="1 - 21" refId="ref10799" refString="Shelford R. 1910. Orthoptera. Fam. Blattidae. Subfam. Epilamprinae. Genera Insectorum 101: 1 - 21." type="journal article" year="1910">Shelford (1910: 12)</bibRefCitation>
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did not discuss the taxonomical position of
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<taxonomicName id="F3814D37FFA0992C5FCDF15C4D28F8EE" authorityName="Walker" authorityYear="1868" box="[477,570,1887,1910]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vasta">
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<emphasis id="06F5EAA6FFA0992C5FCDF15C4EFFF8EE" box="[477,493,1887,1910]" italics="true" pageId="5" pageNumber="6">E</emphasis>
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.
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<emphasis id="06F5EAA6FFA0992C5C11F15C4D28F8EE" box="[513,570,1887,1910]" italics="true" pageId="5" pageNumber="6">vasta</emphasis>
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</taxonomicName>
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, but only listed this species under the genus
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<taxonomicName id="F3814D37FFA0992C5FC7F1834D9DF80F" authorityName="Kirby" authorityYear="1903" box="[471,655,1920,1943]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA0992C5FC7F1834D9DF80F" box="[471,655,1920,1943]" italics="true" pageId="5" pageNumber="6">Pseudophoraspis</emphasis>
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</taxonomicName>
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. In 1967 Princis listed
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<taxonomicName id="F3814D37FFA0992C5F3BF1A14E91F821" authorityName="Walker" authorityYear="1868" box="[299,387,1954,1977]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vasta">
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<emphasis id="06F5EAA6FFA0992C5F3BF1A14E29F821" box="[299,315,1954,1977]" italics="true" pageId="5" pageNumber="6">E</emphasis>
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.
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<emphasis id="06F5EAA6FFA0992C5F5BF1A14E91F821" box="[331,387,1954,1977]" italics="true" pageId="5" pageNumber="6">vasta</emphasis>
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</taxonomicName>
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in the genus
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<taxonomicName id="F3814D37FFA0992C5C07F1A14FDEF842" authority="Kirby, 1903" authorityName="Kirby" authorityYear="1903" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA0992C5C07F1A14DBAF821" box="[535,680,1954,1977]" italics="true" pageId="5" pageNumber="6">Rhabdoblatta</emphasis>
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<bibRefCitation id="50104B45FFA0992C5CA1F1A14FDEF842" author="Kirby W. F." pageId="5" pageNumber="6" pagination="273 - 280" refId="ref10325" refString="Kirby W. F. 1903. Notes on Blattidae & c., with description of new genera and species in the collection of the British Museum, South Kensington. No 2. Annals and Magazine of Natural History (Series 7) 12: 273 - 280." type="journal article" year="1903">Kirby, 1903</bibRefCitation>
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</taxonomicName>
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and placed
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<taxonomicName id="F3814D37FFA0992C5F47F1C04ECCF842" authorityName="Hebard" authorityYear="1930" box="[343,478,1987,2010]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="06F5EAA6FFA0992C5F47F1C04ECCF842" box="[343,478,1987,2010]" italics="true" pageId="5" pageNumber="6">Morphnodes</emphasis>
|
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</taxonomicName>
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in the synonymy of that genus without giving reasons (
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<bibRefCitation id="50104B45FFA0992C5FCDF1E74D7DF863" author="Princis K." box="[477,623,2020,2043]" pageId="5" pageNumber="6" pagination="615 - 710" refId="ref10491" refString="Princis K. 1967. Blattariae: Subordo Epilamproidea. Fam.: Nyctiboridae, Epilampridae. In: Beier M. (ed.). Orthopterorum Catalogus 11: 615 - 710." type="journal article" year="1967">Princis, 1967</bibRefCitation>
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).
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</paragraph>
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<caption id="60FE663CFFA0992C5E83F08C4C82F88F" ID-DOI="http://doi.org/10.5281/zenodo.5589898" ID-Zenodo-Dep="5589898" httpUri="https://zenodo.org/record/5589898/files/figure.png" pageId="5" pageNumber="6" startId="5.[147,188,1679,1700]" targetBox="[161,1418,260,1635]" targetPageId="5">
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||
<paragraph id="343E36B4FFA0992C5E83F08C4C82F88F" blockId="5.[147,1437,1679,1815]" pageId="5" pageNumber="6">
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Figs 21-24.
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<taxonomicName id="F3814D37FFA0992C5F1AF08C4DEEF93C" authority="(Brunner von Wattenwyl, 1865)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1865" box="[266,764,1679,1700]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculata">
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<emphasis id="06F5EAA6FFA0992C5F1AF08C4ED1F93C" box="[266,451,1679,1700]" italics="true" pageId="5" pageNumber="6">Morphna maculata</emphasis>
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(
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<bibRefCitation id="50104B45FFA0992C5FDFF08C4DE7F93C" author="Brunner von Wattenwyl C." box="[463,757,1679,1700]" pageId="5" pageNumber="6" refId="ref10170" refString="Brunner von Wattenwyl C. 1865. Nouveau systeme des Blattaires. Carl Ueberreuter, Wien, 426 pp." type="book" year="1865">Brunner von Wattenwyl, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female specimen No. 141012/04. (21) Abdominal apex, ventral view, genital plate removed. (22) Ovipositor and adjacent structures, ventral view. (23) Basal part of ovipositor, dorsal view. (24) Vestibular sclerite. Dotted areas show membranous parts, except for valves of ovipositor. Abbreviations:
|
||
<emphasis id="06F5EAA6FFA0992C5B1CF0CA4A28F946" box="[1292,1338,1737,1758]" italics="true" pageId="5" pageNumber="6">bd.s.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5B57F0CA4A7EF946" box="[1351,1388,1737,1758]" italics="true" pageId="5" pageNumber="6">bsv.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5B68F0CA4A85F946" box="[1400,1431,1737,1758]" italics="true" pageId="5" pageNumber="6">gg.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5F1AF0E64E23F962" box="[266,305,1765,1786]" italics="true" pageId="5" pageNumber="6">par.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5F2CF0E64E47F962" box="[316,341,1765,1786]" italics="true" pageId="5" pageNumber="6">pl.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5F71F0E64EB2F962" box="[353,416,1765,1786]" italics="true" pageId="5" pageNumber="6">teVIII.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5FBBF0E64EC8F962" box="[427,474,1765,1786]" italics="true" pageId="5" pageNumber="6">teIX.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5FF5F0E64D11F962" box="[485,515,1765,1786]" italics="true" pageId="5" pageNumber="6">v.I.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5C1EF0E64D26F962" box="[526,564,1765,1786]" italics="true" pageId="5" pageNumber="6">v.II.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5C50F0E64D7CF962" box="[576,622,1765,1786]" italics="true" pageId="5" pageNumber="6">v.III.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5C69F0E64D86F962" box="[633,660,1765,1786]" italics="true" pageId="5" pageNumber="6">vs.</emphasis>
|
||
– see paragraph “abbreviation used in figures”;
|
||
<emphasis id="06F5EAA6FFA0992C5A74F0E64B69F962" box="[1124,1147,1765,1786]" italics="true" pageId="5" pageNumber="6">IX</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA0992C5A97F0E64B84F962" box="[1159,1174,1765,1786]" italics="true" pageId="5" pageNumber="6">X</emphasis>
|
||
- abdominal tergites IX-X; for details see text. Scale bars 1 mm: a = 21, b = 22, c = 23, d = 24.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="343E36B4FFA099215E87FE054D87FC63" blockId="5.[151,756,1887,2077]" lastBlockId="8.[151,757,258,1661]" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
|
||
<taxonomicName id="F3814D37FFA0992C5E87FE054D37F785" authority="(Shelford, 1906)" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[151,549,2054,2077]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFA0992C5E87FE054E7DF785" box="[151,367,2054,2077]" italics="true" pageId="5" pageNumber="6">Morphnodes goliath</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFA0992C5F6BFE054D0FF785" author="Shelford R." box="[379,541,2054,2077]" pageId="5" pageNumber="6" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, redescribed below, is similar to
|
||
<taxonomicName id="F3814D37FFA0992C5DD2F15C4B9DF8EE" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1865" box="[962,1167,1887,1910]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="06F5EAA6FFA0992C5DD2F15C4B9DF8EE" box="[962,1167,1887,1910]" italics="true" pageId="5" pageNumber="6">Morphna maculata</emphasis>
|
||
</taxonomicName>
|
||
, but differs from this and other representatives of the genus
|
||
<taxonomicName id="F3814D37FFA0992C5A8CF1834BEDF80F" authorityName="Shelford" authorityYear="1910" box="[1180,1279,1920,1943]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFA0992C5A8CF1834BEDF80F" box="[1180,1279,1920,1943]" italics="true" pageId="5" pageNumber="6">Morphna</emphasis>
|
||
</taxonomicName>
|
||
in the structure of the hind tarsi – a character mentioned by Hebard in the original description of the genus
|
||
<taxonomicName id="F3814D37FFA0992C5B1FF1C04A84F842" authorityName="Hebard" authorityYear="1930" box="[1295,1430,1987,2010]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFA0992C5B1FF1C04A84F842" box="[1295,1430,1987,2010]" italics="true" pageId="5" pageNumber="6">Morphnodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F3814D37FFA0992C5D2FF1E74B32F863" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[831,1056,2020,2043]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFA0992C5D2FF1E74B32F863" box="[831,1056,2020,2043]" italics="true" pageId="5" pageNumber="6">Morphnodes goliath</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="F3814D37FFA0992C5AD3F1E74B21F785" authority="(Walker, 1868)" baseAuthorityName="Walker" baseAuthorityYear="1868" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="praecipua">
|
||
<emphasis id="06F5EAA6FFA0992C5AD3F1E74C68F785" italics="true" pageId="5" pageNumber="6">Rhabdoblatta praecipua</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFA0992C5D80FE054B39F785" author="Walker F." box="[912,1067,2054,2077]" pageId="5" pageNumber="6" refId="ref10895" refString="Walker F. 1868. Catalogue of the specimens of Blattariae in the collection of the British Museum. British Museum, London, 239 pp." type="book" year="1868">Walker, 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="EB3A8816FFA0992C5A64FE054BB0F785" box="[1140,1186,2054,2077]" pageId="5" pageNumber="6">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="F3814D37FFAD99215E87F7014E3AFE81" authorityName="Kirby" authorityYear="1903" box="[151,296,258,281]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215E87F7014E3AFE81" box="[151,296,258,281]" italics="true" pageId="8" pageNumber="9">Rhabdoblatta</emphasis>
|
||
</taxonomicName>
|
||
(see redescription in
|
||
<bibRefCitation id="50104B45FFAD99215C35F7014DFAFE81" author="Anisyutkin L. N." box="[549,744,258,281]" pageId="8" pageNumber="9" pagination="301 - 332" refId="ref9866" refString="Anisyutkin L. N. 2014. On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa. Zootaxa 3847 (3): 301 - 332." type="journal article" year="2014">Anisyutkin, 2014</bibRefCitation>
|
||
), and other species of this genus (
|
||
<bibRefCitation id="50104B45FFAD99215FFDF7214DB9FEA1" author="Anisyutkin L. N." box="[493,683,290,313]" pageId="8" pageNumber="9" pagination="190 - 208" refId="ref9788" refString="Anisyutkin L. N. 2000. New Cockroach species of the genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Southeast Asia: I. Entomological Review 80 (2): 190 - 208." type="journal article" year="2000">Anisyutkin, 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="50104B45FFAD99215CA6F7214DFFFEA1" author="Anisyutkin L. N." box="[694,749,290,313]" pageId="8" pageNumber="9" pagination="540 - 556" refId="ref9825" refString="Anisyutkin L. N. 2003. New and little known cockroaches of the genus Rhabdoblatta Kirby (Dictyoptera, Blaberidae) from Vietnam and southern China: II. Entomological Review 83 (5): 540 - 556." type="journal article" year="2003">2003</bibRefCitation>
|
||
) in the presence of a well developed “dorsal outgrowth” of the apical part of sclerite L2D (
|
||
<figureCitation id="ACBA2A31FFAD99215C31F7614D89FEE1" box="[545,667,354,377]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 56-61</figureCitation>
|
||
) and in the structure of fore and mid tarsi: presence of large euplantulae and absence of any spines (
|
||
<figureCitation id="ACBA2A31FFAD99215C79F7A14DFAFE21" box="[617,744,418,441]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 46-47</figureCitation>
|
||
). In all representatives of
|
||
<taxonomicName id="F3814D37FFAD99215FA9F7C04D58FE42" authorityName="Kirby" authorityYear="1903" box="[441,586,451,474]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215FA9F7C04D58FE42" box="[441,586,451,474]" italics="true" pageId="8" pageNumber="9">Rhabdoblatta</emphasis>
|
||
</taxonomicName>
|
||
known to the author all metatarsal euplantulae are small and apical.
|
||
<taxonomicName id="F3814D37FFAD99215E87F4004E66FD82" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[151,372,515,538]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFAD99215E87F4004E66FD82" box="[151,372,515,538]" italics="true" pageId="8" pageNumber="9">Morphnodes goliath</emphasis>
|
||
</taxonomicName>
|
||
is similar to representatives of the genus
|
||
<taxonomicName id="F3814D37FFAD99215EF4F4204E8EFDA2" authorityName="Kirby" authorityYear="1903" box="[228,412,547,570]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215EF4F4204E8EFDA2" box="[228,412,547,570]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis</emphasis>
|
||
</taxonomicName>
|
||
in the presence of a “dorsal outgrowth” on the apical part of sclerite L2D, but differs from species of
|
||
<taxonomicName id="F3814D37FFAD99215F44F4604D1EFDE2" authorityName="Kirby" authorityYear="1903" box="[340,524,611,634]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215F44F4604D1EFDE2" box="[340,524,611,634]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the “dorsal outgrowth”. In
|
||
<taxonomicName id="F3814D37FFAD99215F9CF4804EE9FD02" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[396,507,643,666]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFAD99215F9CF4804EB0FD02" box="[396,418,643,666]" italics="true" pageId="8" pageNumber="9">M</emphasis>
|
||
.
|
||
<emphasis id="06F5EAA6FFAD99215FBFF4804EE9FD02" box="[431,507,643,666]" italics="true" pageId="8" pageNumber="9">goliath</emphasis>
|
||
</taxonomicName>
|
||
the “dorsal outgrowth” is short and directed upward (
|
||
<figureCitation id="ACBA2A31FFAD99215FDBF4A04D52FD22" box="[459,576,675,698]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 56-61</figureCitation>
|
||
,
|
||
<emphasis id="06F5EAA6FFAD99215C59F4A04D63FD22" box="[585,625,675,698]" italics="true" pageId="8" pageNumber="9">d.o.</emphasis>
|
||
), in contrast to the long and cranial-directed “dorsal outgrowth” of
|
||
<taxonomicName id="F3814D37FFAD99215E87F4E04EB5FD62" box="[151,423,739,762]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="06F5EAA6FFAD99215E87F4E04E5DFD62" box="[151,335,739,762]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="50104B45FFAD99215FF7F4E04DB5FD62" author="Anisyutkin L. N." box="[487,679,739,762]" pageId="8" pageNumber="9" pagination="434 - 454" refId="ref9755" refString="Anisyutkin L. N. 1999. Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula. Entomological Review 79 (4): 434 - 454." type="journal article" year="1999">Anisyutkin, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="50104B45FFAD99215CA6F4E04E33FC83" author="Wang Z. & Wu K. & Che Y." pageId="8" pageNumber="9" pagination="1 - 14" refId="ref10922" refString="Wang Z., Wu K., Che Y. 2013. New record of the cockroach genus Pseudophoraspis (Blaberidae, Epilamprinae) from China with descriptions of three new species. ZooKeys 273: 1 - 14." type="journal article" year="2013">
|
||
Wang
|
||
<emphasis id="06F5EAA6FFAD99215E87F5074FC1FC83" box="[151,211,772,795]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
). The head structure is also different: in species of
|
||
<taxonomicName id="F3814D37FFAD99215F1FF5274ED5FCA3" authorityName="Kirby" authorityYear="1903" box="[271,455,804,827]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215F1FF5274ED5FCA3" box="[271,455,804,827]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis</emphasis>
|
||
</taxonomicName>
|
||
the facial part of the head has a distinct longitudinal impression (see
|
||
<bibRefCitation id="50104B45FFAD99215C67F5474FDDFCE3" author="Anisyutkin L. N." pageId="8" pageNumber="9" pagination="434 - 454" refId="ref9755" refString="Anisyutkin L. N. 1999. Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula. Entomological Review 79 (4): 434 - 454." type="journal article" year="1999">Anisyutkin, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="50104B45FFAD99215EF0F5674EA6FCE3" author="Wang Z. & Wu K. & Che Y." box="[224,436,868,891]" pageId="8" pageNumber="9" pagination="1 - 14" refId="ref10922" refString="Wang Z., Wu K., Che Y. 2013. New record of the cockroach genus Pseudophoraspis (Blaberidae, Epilamprinae) from China with descriptions of three new species. ZooKeys 273: 1 - 14." type="journal article" year="2013">
|
||
Wang
|
||
<emphasis id="06F5EAA6FFAD99215F3BF5674E75FCE3" box="[299,359,868,891]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
). The apical part of sclerite L2D in
|
||
<taxonomicName id="F3814D37FFAD99215EEBF5874DE6FC03" authority="Shelford, 1910" authorityName="Shelford" authorityYear="1910" box="[251,756,900,923]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fruhstorferi">
|
||
<emphasis id="06F5EAA6FFAD99215EEBF5874D2FFC03" box="[251,573,900,923]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis fruhstorferi</emphasis>
|
||
<bibRefCitation id="50104B45FFAD99215C5DF5874DE6FC03" author="Shelford R." box="[589,756,900,923]" pageId="8" pageNumber="9" pagination="1 - 21" refId="ref10799" refString="Shelford R. 1910. Orthoptera. Fam. Blattidae. Subfam. Epilamprinae. Genera Insectorum 101: 1 - 21." type="journal article" year="1910">Shelford, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3814D37FFAD99215ED8F5A74D28FC23" authority="Anisyutkin, 1999" authorityName="Anisyutkin" authorityYear="1999" box="[200,570,932,955]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tramlapensis">
|
||
<emphasis id="06F5EAA6FFAD99215ED8F5A74FCAFC23" box="[200,216,932,955]" italics="true" pageId="8" pageNumber="9">P</emphasis>
|
||
.
|
||
<emphasis id="06F5EAA6FFAD99215EFAF5A74E66FC23" box="[234,372,932,955]" italics="true" pageId="8" pageNumber="9">tramlapensis</emphasis>
|
||
<bibRefCitation id="50104B45FFAD99215F6DF5A74D28FC23" author="Anisyutkin L. N." box="[381,570,932,955]" pageId="8" pageNumber="9" pagination="434 - 454" refId="ref9755" refString="Anisyutkin L. N. 1999. Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula. Entomological Review 79 (4): 434 - 454." type="journal article" year="1999">Anisyutkin, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
have no “dorsal outgrowth” (
|
||
<bibRefCitation id="50104B45FFAD99215F32F5C74EF1FC43" author="Anisyutkin L. N." box="[290,483,964,987]" pageId="8" pageNumber="9" pagination="434 - 454" refId="ref9755" refString="Anisyutkin L. N. 1999. Cockroaches of the subfamily Epilamprinae (Dictyoptera, Blaberidae) from the Indochina Peninsula. Entomological Review 79 (4): 434 - 454." type="journal article" year="1999">Anisyutkin, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="50104B45FFAD99215FE1F5C74DAEFC43" author="Wang Z. & Wu K. & Che Y." box="[497,700,964,987]" pageId="8" pageNumber="9" pagination="1 - 14" refId="ref10922" refString="Wang Z., Wu K., Che Y. 2013. New record of the cockroach genus Pseudophoraspis (Blaberidae, Epilamprinae) from China with descriptions of three new species. ZooKeys 273: 1 - 14." type="journal article" year="2013">
|
||
Wang
|
||
<emphasis id="06F5EAA6FFAD99215C29F5C74D60FC43" box="[569,626,964,987]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
), but these species probably belong to another genus.
|
||
</paragraph>
|
||
<caption id="60FE663CFFA3992F5E87F1234EC9F784" ID-DOI="http://doi.org/10.5281/zenodo.5589904" ID-Zenodo-Dep="5589904" httpUri="https://zenodo.org/record/5589904/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,192,1824,1845]" targetBox="[162,1432,258,1769]" targetPageId="6">
|
||
<paragraph id="343E36B4FFA3992F5E87F1234EC9F784" blockId="6.[151,1436,1824,2076]" pageId="6" pageNumber="7">
|
||
Figs 25-43.
|
||
<taxonomicName id="F3814D37FFA3992F5F1EF1234DE8F8AD" authority="(Brunner von Wattenwyl, 1865)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1865" box="[270,762,1824,1845]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="06F5EAA6FFA3992F5F1EF1234ED7F8AD" box="[270,453,1824,1845]" italics="true" pageId="6" pageNumber="7">Morphna maculata</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFA3992F5FC1F1234DE1F8AD" author="Brunner von Wattenwyl C." box="[465,755,1824,1845]" pageId="6" pageNumber="7" refId="ref10170" refString="Brunner von Wattenwyl C. 1865. Nouveau systeme des Blattaires. Carl Ueberreuter, Wien, 426 pp." type="book" year="1865">Brunner von Wattenwyl, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, males (25-26, 33, 37-43: specimen in BMNH, 27-32, 34-36: specimen in ZIN, No. 141012/03). (25) Facial part of head. (26) Pronotum, dorsal view. (27) Abdominal apex, dorsal view. (28) Abdominal apex, ventral view (only right sides of VII and VIII tergites are shown). (29) Paraprocts, posterior view. (30) Hypandrium, ventral view. (31) Right phallomere, ventral view. (32) Same, dorsal view. (33) Sclerite L2D, dorsal view, bristles not shown. (34, 37) Posterior part of sclerite L2D, lateral view. (35, 38) Same, dorsal view. (36, 39) The same, contralateral view. (40) Sclerite L3. (41-43) Apex of sclerite L3. Dotted areas show membranous parts. Abbreviations:
|
||
<emphasis id="06F5EAA6FFA3992F5F1EF1CE4E4BF87A" box="[270,345,1997,2018]" italics="true" pageId="6" pageNumber="7">c.p.R1T</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5F75F1CE4E9BF87A" box="[357,393,1997,2018]" italics="true" pageId="6" pageNumber="7">d.o.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5F84F1CE4EA9F87A" box="[404,443,1997,2018]" italics="true" pageId="6" pageNumber="7">par.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5FD5F1CE4ECDF87A" box="[453,479,1997,2018]" italics="true" pageId="6" pageNumber="7">R2</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5FFBF1CE4D14F87A" box="[491,518,1997,2018]" italics="true" pageId="6" pageNumber="7">R3</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5C02F1CE4D3FF87A" box="[530,557,1997,2018]" italics="true" pageId="6" pageNumber="7">R4</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5C28F1CE4D41F87A" box="[568,595,1997,2018]" italics="true" pageId="6" pageNumber="7">R5</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5C4FF1CE4D69F87A" box="[607,635,1997,2018]" italics="true" pageId="6" pageNumber="7">s.t.</emphasis>
|
||
– see paragraph “abbreviation used in figures”;
|
||
<emphasis id="06F5EAA6FFA3992F5A58F1CE4B75F87A" box="[1096,1127,1997,2018]" italics="true" pageId="6" pageNumber="7">VII</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5A62F1CE4B8BF87A" box="[1138,1177,1997,2018]" italics="true" pageId="6" pageNumber="7">VIII</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5AB4F1CE4BA9F87A" box="[1188,1211,1997,2018]" italics="true" pageId="6" pageNumber="7">IX</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA3992F5AD7F1CE4BC4F87A" box="[1223,1238,1997,2018]" italics="true" pageId="6" pageNumber="7">X</emphasis>
|
||
- abdominal tergites VII-X; for details see text. Scale bars 1 mm: a = 25, b = 26, c = 27, d = 28-29, e = 30, f = 31-32, g = 33, h = 34-36, i = 37- 39, j = 40, k = 41-43.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="60FE663CFFA2992E5EB3F1254A89F79D" ID-DOI="http://doi.org/10.5281/zenodo.5589908" ID-Zenodo-Dep="5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="7" pageNumber="8" startId="7.[163,204,1830,1851]" targetBox="[165,1423,257,1781]" targetPageId="7">
|
||
<paragraph id="343E36B4FFA2992E5EB3F1254A89F79D" blockId="7.[163,1435,1830,2053]" pageId="7" pageNumber="8">
|
||
Figs 44-65.
|
||
<taxonomicName id="F3814D37FFA2992E5F0AF1254D95F8A3" authority="(Shelford, 1906)" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[282,647,1830,1851]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFA2992E5F0AF1254ECCF8A3" box="[282,478,1830,1851]" italics="true" pageId="7" pageNumber="8">Morphnodes goliath</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFA2992E5FFCF1254D92F8A3" author="Shelford R." box="[492,640,1830,1851]" pageId="7" pageNumber="8" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts. Abbreviations:
|
||
<emphasis id="06F5EAA6FFA2992E5FF6F1D04D1AF870" box="[486,520,2003,2024]" italics="true" pageId="7" pageNumber="8">a.s.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5C03F1D04D2CF870" box="[531,574,2003,2024]" italics="true" pageId="7" pageNumber="8">b.L3</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5C59F1D04D86F870" box="[585,660,2003,2024]" italics="true" pageId="7" pageNumber="8">c.p.R1T</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5C8FF1D04DD6F870" box="[671,708,2003,2024]" italics="true" pageId="7" pageNumber="8">d.o.</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5CDFF1D04DFBF870" box="[719,745,2003,2024]" italics="true" pageId="7" pageNumber="8">R2</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5CE5F1D04C1DF870" box="[757,783,2003,2024]" italics="true" pageId="7" pageNumber="8">R3</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5D0BF1D04C24F870" box="[795,822,2003,2024]" italics="true" pageId="7" pageNumber="8">R4</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5D51F1D04C4EF870" box="[833,860,2003,2024]" italics="true" pageId="7" pageNumber="8">R5</emphasis>
|
||
,
|
||
<emphasis id="06F5EAA6FFA2992E5D77F1D04C91F870" box="[871,899,2003,2024]" italics="true" pageId="7" pageNumber="8">s.t.</emphasis>
|
||
see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="343E36B4FFAD99215E87F2074D68FBE4" blockId="8.[151,757,258,1661]" pageId="8" pageNumber="9">
|
||
Taking into account the aforementioned, I prefer to restore the genus
|
||
<taxonomicName id="F3814D37FFAD99215F5FF2274EC4FBA3" authorityName="Hebard" authorityYear="1930" box="[335,470,1060,1083]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215F5FF2274EC4FBA3" box="[335,470,1060,1083]" italics="true" pageId="8" pageNumber="9">Morphnodes</emphasis>
|
||
</taxonomicName>
|
||
from synonymy. To clarify the status of the genus
|
||
<taxonomicName id="F3814D37FFAD99215F8FF2474D34FBC3" authorityName="Hebard" authorityYear="1930" box="[415,550,1092,1115]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215F8FF2474D34FBC3" box="[415,550,1092,1115]" italics="true" pageId="8" pageNumber="9">Morphnodes</emphasis>
|
||
</taxonomicName>
|
||
, it is necessary to redescribe the
|
||
<typeStatus id="EB3A8816FFAD99215F23F2664E73FBE4" box="[307,353,1125,1148]" pageId="8" pageNumber="9">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="F3814D37FFAD99215FD2F2664D60FBE4" authorityName="Walker" authorityYear="1868" box="[450,626,1125,1148]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vasta">
|
||
<emphasis id="06F5EAA6FFAD99215FD2F2664D60FBE4" box="[450,626,1125,1148]" italics="true" pageId="8" pageNumber="9">Epilampra vasta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="343E36B4FFAD99215E87F2864DBEFAE4" blockId="8.[151,757,258,1661]" pageId="8" pageNumber="9">
|
||
The genera
|
||
<taxonomicName id="F3814D37FFAD99215F32F2864E97FB04" authorityName="Shelford" authorityYear="1910" box="[290,389,1157,1180]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215F32F2864E97FB04" box="[290,389,1157,1180]" italics="true" pageId="8" pageNumber="9">Morphna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3814D37FFAD99215FD6F2864D5FFB04" authorityName="Hebard" authorityYear="1930" box="[454,589,1157,1180]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215FD6F2864D5FFB04" box="[454,589,1157,1180]" italics="true" pageId="8" pageNumber="9">Morphnodes</emphasis>
|
||
</taxonomicName>
|
||
are similar in appearance and in the structure of the male genitalia. They are probably sister-groups with the following synapomorphies: (1) wide costal field of tegmina; (2) fore and mid tarsi with large euplantulae; (3) fore and mid tarsi with reduced spines; (4) “dorsal outgrowth” of apical part of sclerite L2D large and directed upward (large and directed cranially in
|
||
<taxonomicName id="F3814D37FFAD99215FF5F3664D8FFAE4" authorityName="Kirby" authorityYear="1903" box="[485,669,1381,1404]" class="Insecta" family="Blaberidae" genus="Pseudophoraspis" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06F5EAA6FFAD99215FF5F3664D8FFAE4" box="[485,669,1381,1404]" italics="true" pageId="8" pageNumber="9">Pseudophoraspis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="343E36B4FFAD99215E87F3864DC1FA25" blockId="8.[151,757,258,1661]" pageId="8" pageNumber="9">
|
||
A species with a similar venation of the tegmina was described from the Paleocene (
|
||
<bibRefCitation id="50104B45FFAD99215FF0F3A54DDAFA25" author="Vrsansky P. & Vidlicka L. & Barna P. & Bugdaeva E. & Markevich V." box="[480,712,1446,1469]" pageId="8" pageNumber="9" pagination="117 - 126" refId="ref10846" refString="Vrsansky P., Vidlicka L., Barna P., Bugdaeva E., Markevich V. 2013. Paleocene origin of the cockroach families Blaberidae and Corydiidae: Evidence from Amur River region of Russia. Zootaxa 3635 (2): 117 - 126." type="journal article" year="2013">
|
||
Vršanský
|
||
<emphasis id="06F5EAA6FFAD99215C5BF3A54D93FA25" box="[587,641,1446,1469]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="343E36B4FFAD99215E87F3C54FE9F9E5" blockId="8.[151,757,258,1661]" pageId="8" pageNumber="9">
|
||
In the catalogues of
|
||
<bibRefCitation id="50104B45FFAD99215F60F3C54D1AFA45" author="Princis K." box="[368,520,1478,1501]" pageId="8" pageNumber="9" pagination="615 - 710" refId="ref10491" refString="Princis K. 1967. Blattariae: Subordo Epilamproidea. Fam.: Nyctiboridae, Epilampridae. In: Beier M. (ed.). Orthopterorum Catalogus 11: 615 - 710." type="journal article" year="1967">Princis (1967)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="50104B45FFAD99215C2CF3C54DE6FA45" author="Beccaloni G. W." box="[572,756,1478,1501]" pageId="8" pageNumber="9" refId="ref10070" refString="Beccaloni G. W. 2014. Cockroach Species File Online. Version 5.0 / 5.0. World Wide Web electronic publication. Available at http: // Cockroach. SpeciesFile. org (accessed 22 March 2017)." type="url" year="2014">Beccaloni (2014)</bibRefCitation>
|
||
the date of publication of Hebard’s paper is incorrectly given. Volume 81 of the Proceedings of the Academy of Natural Sciences of Philadelphia for the year 1929 was in fact published in 1930, as stated on the first page of the issue.
|
||
</paragraph>
|
||
<paragraph id="343E36B4FFAD99215E87F0964EC1F975" blockId="8.[151,756,1685,1773]" pageId="8" pageNumber="9">
|
||
<emphasis id="06F5EAA6FFAD99215E87F0964E4FF934" bold="true" box="[151,349,1685,1708]" pageId="8" pageNumber="9">Included species:</emphasis>
|
||
<taxonomicName id="F3814D37FFAD99215F76F0964DD8F934" authority="Walker, 1868" authorityName="Walker" authorityYear="1868" box="[358,714,1685,1708]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vasta">
|
||
<emphasis id="06F5EAA6FFAD99215F76F0964D3CF934" box="[358,558,1685,1708]" italics="true" pageId="8" pageNumber="9">Morphnodes vasta</emphasis>
|
||
<bibRefCitation id="50104B45FFAD99215C27F0964DD8F934" author="Walker F." box="[567,714,1685,1708]" pageId="8" pageNumber="9" refId="ref10895" refString="Walker F. 1868. Catalogue of the specimens of Blattariae in the collection of the British Museum. British Museum, London, 239 pp." type="book" year="1868">Walker, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3814D37FFAD99215CC7F0964EBAF954" authority="(Stal, 1877)" baseAuthorityName="Stal" baseAuthorityYear="1877" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="imperatoria">
|
||
<emphasis id="06F5EAA6FFAD99215CC7F0964DFFF934" box="[727,749,1685,1708]" italics="true" pageId="8" pageNumber="9">M</emphasis>
|
||
.
|
||
<emphasis id="06F5EAA6FFAD99215E87F0B64E05F954" box="[151,279,1717,1740]" italics="true" pageId="8" pageNumber="9">imperatoria</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFAD99215F3BF0B64EB2F954" author="Stal C." box="[299,416,1717,1740]" pageId="8" pageNumber="9" pagination="33 - 58" refId="ref10822" refString="Stal C. 1877. Orthoptera nova ex insulis Philippinis descripsit. Ofversigt ap Kongl. Vetenskaps-Akademiens Forhandlingar 10: 33 - 58." type="journal article" year="1877">Stål, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3814D37FFAD99215FA8F0B64DE2F954" authority="(Shelford, 1906)" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[440,752,1717,1740]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="goliath">
|
||
<emphasis id="06F5EAA6FFAD99215FA8F0B64EDCF954" box="[440,462,1717,1740]" italics="true" pageId="8" pageNumber="9">M</emphasis>
|
||
.
|
||
<emphasis id="06F5EAA6FFAD99215FF0F0B64D3EF954" box="[480,556,1717,1740]" italics="true" pageId="8" pageNumber="9">goliath</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFAD99215C2FF0B64DFAF954" author="Shelford R." box="[575,744,1717,1740]" pageId="8" pageNumber="9" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3814D37FFAD99215E87F0D54EDDF975" authority="(Shelford, 1906)" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[151,463,1750,1773]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="miranda">
|
||
<emphasis id="06F5EAA6FFAD99215E87F0D54FBFF975" box="[151,173,1750,1773]" italics="true" pageId="8" pageNumber="9">M</emphasis>
|
||
.
|
||
<emphasis id="06F5EAA6FFAD99215EABF0D54E07F975" box="[187,277,1750,1773]" italics="true" pageId="8" pageNumber="9">miranda</emphasis>
|
||
(
|
||
<bibRefCitation id="50104B45FFAD99215F33F0D54EDAF975" author="Shelford R." box="[291,456,1750,1773]" pageId="8" pageNumber="9" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |