131 lines
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131 lines
16 KiB
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<document id="1CAAC0EE667C9FD42DD4648CB2A0577D" ID-CLB-Dataset="33557" ID-DOI="http://dx.doi.org/10.3897/dez.62.5093" ID-GBIF-Dataset="1d374c05-3d43-4b84-8695-23a5978148ef" ID-Pensoft-Pub="1860-1324-1-101" ID-ZooBank="BF4238DAC6A24AF0AB80697A1FFF3374" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1860-1324-1-101" ModsDocOrigin="Deutsche Entomologische Zeitschrift 62 (1)" ModsDocTitle="Contributions to the taxonomy of the ant genus Echinopla Smith, 1857 (Hymenoptera, Formicidae)" checkinTime="1433500152900" checkinUser="pensoft" docAuthor="Zettel, Herbert & Laciny, Alice" docDate="2015" docId="B970FC52A6D3BAB80EDC232D8B36F193" docLanguage="en" docName="DeutEntomolZeit 62(1): 101-121" docOrigin="Deutsche Entomologische Zeitschrift 62 (1)" docSource="http://dx.doi.org/10.3897/dez.62.5093" docTitle="Echinopla wardi Zettel & Laciny, 2015, sp. n." docType="treatment" docUuid="044DDB4D-67C1-4A69-8FF8-677588D569F9" docUuidSource="ZooBank" docVersion="7" lastPageNumber="104" masterDocId="FC1B5475D767FFE2FFD41E00FF8CFFD8" masterDocTitle="Contributions to the taxonomy of the ant genus Echinopla Smith, 1857 (Hymenoptera, Formicidae)" masterLastPageNumber="121" masterPageNumber="101" pageNumber="102" updateTime="1701381044664" updateUser="ExternalLinkService">
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<mods:mods id="B9CD6FB2E952C07125D2D2C8C42EE8AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EA733B6F56B29444BC79290F8E52DD5F">
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<mods:title id="A6A5C6917BD52D7DE207CAE0CF1895D9">Contributions to the taxonomy of the ant genus Echinopla Smith, 1857 (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="4D91E332B1C6AD340D5791AE28977258">Zettel, Herbert</mods:namePart>
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<mods:namePart id="D60779039E391289E99A6444A836070A">Laciny, Alice</mods:namePart>
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<mods:typeOfResource id="51C07D66042B63A5998CF6BED65B025A">text</mods:typeOfResource>
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<mods:title id="624A76158A259EC2B16555A66FDE336D">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="316C020EA0FD3F421015424BB5349B82">
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<mods:date id="6AE5C7DDEAA529D5FD892A1B51C22981">2015</mods:date>
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<mods:detail id="B28719447EFADA9F3842C3E2B145DCD3" type="volume">
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<mods:number id="02AD727D85D800B4279D89B72FA604FF">62</mods:number>
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<mods:number id="143BDBB974FBE4B6453C7BB0805E1514">1</mods:number>
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<mods:start id="F707F9DEB4B5E2603E0A4D652BB54475">101</mods:start>
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<mods:url id="14C00D201F56D9444FDD305AA93B1727">http://dx.doi.org/10.3897/dez.62.5093</mods:url>
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<mods:classification id="912061F5A1F64B886CE98BD3DB2427F4">journal article</mods:classification>
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<mods:identifier id="A2C27A20318BFDA0062CEDF4622FC190" type="DOI">http://dx.doi.org/10.3897/dez.62.5093</mods:identifier>
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<mods:identifier id="A5E9E43D5456EB5C4A24F1EFAAB30415" type="Pensoft-Pub">1860-1324-1-101</mods:identifier>
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<treatment id="B970FC52A6D3BAB80EDC232D8B36F193" ID-GBIF-Taxon="127900107" LSID="urn:lsid:zoobank.org:act:044DDB4D-67C1-4A69-8FF8-677588D569F9" httpUri="http://treatment.plazi.org/id/B970FC52A6D3BAB80EDC232D8B36F193" lastPageId="3" lastPageNumber="104" pageId="1" pageNumber="102">
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<subSubSection id="E422C484B16F75BA48A6AD77C6A9F944" pageId="1" pageNumber="102" type="multiple">
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<paragraph id="3DA4ADA2BCC8D167CED24BDF4C590BA3" pageId="1" pageNumber="102">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
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</subSubSection>
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<subSubSection id="A84F0E7AB5954241E4BA7E510EF8241C" pageId="1" pageNumber="102" type="nomenclature">
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<paragraph id="8F96CF7D05255B0201450572ED05DB40" pageId="1" pageNumber="102">
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<taxonomicName id="CB477BC36E6BAE19647FF7F3DAB5F839" LSID="http://zoobank.org/044DDB4D-67C1-4A69-8FF8-677588D569F9" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla wardi" order="Hymenoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="wardi">Echinopla wardi</taxonomicName>
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<taxonomicNameLabel id="F8C01E2859FB32F20E03E66DEEA0A3E9" pageId="1" pageNumber="102">sp. n.</taxonomicNameLabel>
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Figs 5-8
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</paragraph>
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</subSubSection>
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<subSubSection id="8107A73DC839E657249C92E287D52386" pageId="1" pageNumber="102" type="type material">
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<paragraph id="254A1E05028C4513A935E1B759FC9F35" pageId="1" pageNumber="102">Type material.</paragraph>
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<paragraph id="B0EF8D99B18500F7C8190581E2B8525E" pageId="1" pageNumber="102">
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Holotype (worker, NHMW) from Malaysia, Johor, 13 km NE Kota Tinggi,
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<geoCoordinate id="91E7F839C20139EA0384A042BBE93D81" direction="north" orientation="latitude" precision="925" value="1.8166666">1°49'N</geoCoordinate>
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;
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<geoCoordinate id="AAD8D0518EEC8A5228A87F5F024EFA91" direction="east" orientation="longitude" precision="925" value="103.833336">103°50'E</geoCoordinate>
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, 50 m a.s.l., 22.XI.1988, on low vegetation in second-growth rain forest, leg. Philip S. Ward (PSW09591-12; ANTWEB CASENT 0280337). Paratype (worker, ZCW) from West Malaysia, Pahang, 70 km SW Kuala Rompin, Endau Rompin National Park, Gunung Beremban, 600 m a.s.l., 13.IV.-3.V.2010, leg. Petr
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<normalizedToken id="296708731AB0CE48636203515DC7A8A7" originalValue="Čechovský">Cechovsky</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA9899731820D27258C65BE476792362" pageId="1" pageNumber="102" type="diagnosis">
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<paragraph id="539CC99CB369994064F2749D3FFC9FB1" pageId="1" pageNumber="102">Diagnosis</paragraph>
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<paragraph id="06503E713A2A6A81BC9002D939EC5ABA" pageId="1" pageNumber="102">(worker). Predominantly black, small species, TL = 3.8-4.3. Surface polyporous, dorsal margins of mesosoma with sharp tubercles. Head wider than long (CI = 110). Mesosoma roughly 1.25 times as long as pronotal width (MI 122-126), with sharp and deep incision in front of propodeum. Pronotal width equal to head width, if eyes excluded. Propodeum shorter than promesonotum. Petiole dentate, with three sharp teeth laterally below lateral spine. Gaster tergite 1 about one fourth longer than wide, anteriorly with relatively large pores and shiny interspaces; disk with dense subcumbent pilosity. Setae on dorsal surface, legs and scape relatively short. Setae on tibia reaching or surpassing base of distally following setae.</paragraph>
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</subSubSection>
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<subSubSection id="5138F6FF1010D03FFCDF8105F5B64087" lastPageId="3" lastPageNumber="104" pageId="1" pageNumber="102" type="description">
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<paragraph id="B73B1F31D0C9322C1AB9F10DBED52C6A" pageId="1" pageNumber="102">Description.</paragraph>
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<paragraph id="6AAB98300D09C0CE546F7197214A949F" pageId="1" pageNumber="102">Measurements of holotype worker: TL 3.8; HW1 1.11; HW2 1.02; HL 1.01; EL 0.23; SL 0.96; SW 0.12; HaL 0.16; PML 0.72; PMW 1.02; PpL 0.57; PpW 0.91; PH 0.45; PL 0.40; PW 0.91; GL 1.27; GW 1.11. Indices: CI 110; SI 80; MI 126.</paragraph>
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<paragraph id="F52B9BB79DAAB6D586EED8B234A19E9C" pageId="2" pageNumber="103">
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<pageBreakToken id="D95F1F5C39DEE5AF634198F7F890311F" pageId="2" pageNumber="103" start="start">Measurements</pageBreakToken>
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of paratype worker: TL 4.3; HW1 1.17; HW2 1.11; HL 1.07; EL 0.23; SL 0.96; SW 0.12; HaL 0.14; PML 0.76; PMW 1.11; PpL 0.59; PpW 1.00; PH 0.51; PL 0.38; PW 1.03; GL 1.37; GW 1.24. Indices: CI 110; SI 81; MI 122.
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</paragraph>
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<paragraph id="4B7ADCD03D76C243416476265023D8E5" pageId="2" pageNumber="103">Structures: Head wider than long, subtrapezoidal, with slightly convex sides; dorsally and laterally polyporous, with closely set pores, posterolaterally with a few minute tubercles, matt; ventral surface smooth and shiny. Compound eye relatively small, moderately protruding, positioned slightly behind mid-length of head. Frons with prominent median carina; frontal lobes chiefly horizontally oriented, largely covering antennal fossae in dorsal aspect, maximum distance of margins slightly behind mid-length, as great as half of HW2. Clypeus with distinct median carina, anterior margin almost straight. Mandibles striate, masticatory margin with five teeth. Antennal scape moderately long, weakly s-curved, steadily widened from base to apex; antennomeres 8-10 approximately as wide as long.</paragraph>
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<paragraph id="5D95240ECC3866EE8D68CE2C68839520" pageId="2" pageNumber="103">Mesosoma stout, length ca. 1.25 times pronotum width; propodeum shorter than promesonotum. Surface polyporous, with closely set pores, dorsal margins with sharp tubercles. Pronotum with distinct lateral angles, as wide as head excluding eyes. Promesonotal suture weak, indicated by a convex row of deeper pits. Mesometanotal suture sharp and deep, medially narrow, laterally widened. In dorsal aspect mesosoma with waist-like incision in front of propodeum. Legs moderately long; femora not much widened.</paragraph>
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<paragraph id="3CE5DC81BF4F3FAA0BF07BA4984BE096" pageId="2" pageNumber="103">Petiole wide and stout, subtriangular in lateral view, strongly transverse in dorsal aspect; surface structure as on mesosoma; sharp dorsal crest bearing four sharp teeth medially and one pair of small denticles laterally; lateral tooth strongly developed; below lateral tooth with one long and two short, sharp teeth. Gaster tergite 1 clearly longer than wide, completely covering the following tergites; hind margin moderately convex, finely serrate; surface polyporous, interspaces smooth and shiny; distances of pores mostly smaller than their diameters.</paragraph>
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<paragraph id="5CBD5A1F09D0556B730296CC04788C7B" pageId="2" pageNumber="103">Pilosity: Entire trunk dorsally and laterally with dense, appressed white pilosity and moderately long white standing setae; only the standing setae on anterior part of gaster tergite 1 distinctly longer. White standing setae on scape about as long as the setae on head. Legs with fine appressed pilosity. Femora with some long standing setae on flexor side. Tibiae and first tarsomeres of middle and hind leg with rows of oblique white setae; their length as long as their distance.</paragraph>
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<paragraph id="AC67EB7181496DB5142499E997E241FF" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
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Colour: Trunk black, without metallic shimmer, appearing grey by whitish pilosity. Antenna with black scape; funiculus basally black, becoming gradually paler
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<pageBreakToken id="34D1459C3771736094D8AE251A59BF8D" pageId="3" pageNumber="104" start="start">towards</pageBreakToken>
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testaceous apex. Mandible basally black, distally brown; other mouthparts yellowish. Legs chiefly blackish, but coxae of middle and hind legs brown, all trochanters dark testaceous, and tarsi distally reddish testaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="BB0F3350D1E2DD93F26525DD744426BF" pageId="3" pageNumber="104" type="comparative notes">
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<paragraph id="44AE2BB48CED09F24DF569676BE2F007" pageId="3" pageNumber="104">Comparative notes.</paragraph>
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<paragraph id="9ADFB643D67FBEFF3E09DB436917590B" pageId="3" pageNumber="104">
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<taxonomicName id="2970C09EAE1B4BB4A23F78CE0CD562B5" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla wardi" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="wardi">Echinopla wardi</taxonomicName>
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sp. n. is similar to
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<taxonomicName id="8EC617BF0DB6915F01140E97238C80C4" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla australis" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="australis">Echinopla australis</taxonomicName>
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Forel, 1901 from Australia, but this species has yellow legs and funiculus, relatively long pilosity, and pronotal width distinctly smaller than head width. Images of
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<taxonomicName id="721814340AD33CCE8CB1DC6BA71C4015" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla australis" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="australis">Echinopla australis</taxonomicName>
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are presented by
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<bibRefCitation id="746E7663638C4503A5D137CBA0387677" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antweb.org" year="2015">Antweb (2015)</bibRefCitation>
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. According to the original description (
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<bibRefCitation id="46668AE1D76F854F7DCB0E4BB5A2BDE3" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" pagination="46 - 55" title="Voyage de M. Chaper a Borneo. Catalogue des fourmis et description des especes nouvelles." volume="5" year="1892">
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<normalizedToken id="21227701327A482E700CA9082CA0AB16" originalValue="André">Andre</normalizedToken>
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1892
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</bibRefCitation>
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),
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<taxonomicName id="FD29A4930FE1A18B40D7276D3EB41BB0" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla rugosa" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="rugosa">Echinopla rugosa</taxonomicName>
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<normalizedToken id="E918597591B34BE0E8B59C517E3098ED" originalValue="André">Andre</normalizedToken>
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, 1892 from Borneo has similar structures of mesosoma and petiole, but differs by uniformly black colour with bronze shimmer and slightly smaller size. Unfortunately the holotype depository of this enigmatic species is unknown and no further specimen has become known. See also notes for
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<taxonomicName id="537232669960A9715B3AE25E839338F9" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla madli" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="madli">Echinopla madli</taxonomicName>
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sp. n. and
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<taxonomicName id="B5CC7BFCDBB0FA7180F5B5EDAC36BE22" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla brevisetosa" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="brevisetosa">Echinopla brevisetosa</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph id="A9F4498288A15E443DC3B95BC4CA79EE" pageId="3" pageNumber="104">
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Figures 5-8 are taken from the paratype whereas the holotype is illustrated by
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<bibRefCitation id="89DCA0638318997DCC61DF25D80C907A" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antweb.org" year="2015">Antweb (2015)</bibRefCitation>
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with specimen number CASENT 0280337.
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</paragraph>
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<caption id="5A29FB8A83391DFC5368C9CE85EA7FD8" pageId="3" pageNumber="104">
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<paragraph id="1E3D05CA2400088CCF23B91BDA8CB0DC" pageId="3" pageNumber="104">
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Figures 5-8.
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<taxonomicName id="0840ADAB0CDBF573220CB7ABC5EE5016" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla wardi" order="Hymenoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="wardi">Echinopla wardi</taxonomicName>
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sp. n., paratype: (5) Head, full face view. (6) Habitus, lateral view. (7) Habitus, dorsal view. (8) Labels.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="9C1B992C49DCC4B8D16E4AF304AF9213" pageId="3" pageNumber="104" type="etymology">
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<paragraph id="D525DC96734FCB19A7E24F1F683F1CB0" pageId="3" pageNumber="104">Etymology.</paragraph>
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<paragraph id="DE7344D33ADC33EEC390B02220C0F468" pageId="3" pageNumber="104">Dedicated to Phil S. Ward (University of California) who collected the holotype and kindly donated it to NHMW.</paragraph>
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</subSubSection>
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</treatment>
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</document> |