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<mods:title id="63E54606D36CDDB14F8112A654DE9923">Notes on species of the genus Chlaenius Bonelli, 1810 (Coleoptera: Carabidae, Chlaeniini) from four agro-climatic zones of Rajasthan, with description of two new species</mods:title>
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<mods:namePart id="00DB9F4FA3AB10C7D1FDC930E76F8811">Chanu, N. Yaiphabi</mods:namePart>
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<paragraph id="26538D42D478FFE0FF46FB055EAC6EE2" blockId="20.[151,1018,1163,1222]" box="[151,1018,1163,1189]" pageId="20" pageNumber="565">
<heading id="7D1B3A2ED478FFE0FF46FB055EAC6EE2" bold="true" box="[151,1018,1163,1189]" fontSize="11" level="1" pageId="20" pageNumber="565" reason="1">
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<taxonomicName id="E1ECF6C1D478FFE0FF46FB055EC36EE2" ID-CoL="69W87" authority="Chanu &amp; Swaminathan" authorityName="Chanu &amp; Swaminathan" box="[151,917,1163,1189]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="565" phylum="Arthropoda" rank="species" species="pseudotristis" status="sp. nov." subGenus="Chlaeniellus">
<emphasis id="14985150D478FFE0FF46FB055C456EE2" bold="true" box="[151,275,1163,1189]" italics="true" pageId="20" pageNumber="565">Chlaenius</emphasis>
(
<emphasis id="14985150D478FFE0FEF5FB055CEB6EE2" bold="true" box="[292,445,1163,1189]" italics="true" pageId="20" pageNumber="565">Chlaeniellus</emphasis>
) 
<emphasis id="14985150D478FFE0FE1FFB055F336EE2" bold="true" box="[462,613,1163,1189]" italics="true" pageId="20" pageNumber="565">pseudotristis</emphasis>
Chanu &amp; Swaminathan
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<taxonomicNameLabel id="0FABEC2BD478FFE0FC4DFB055EAC6EE2" box="[924,1018,1163,1189]" pageId="20" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="BED791C7D478FFE0FF4EFB235DAA6E81" box="[159,252,1197,1222]" captionStart="FIGURES 1 – 14" captionStartId="4.[151,264,1928,1951]" captionTargetBox="[262,1336,242,1856]" captionTargetId="figure@4.[211,1368,193,1881]" captionTargetPageId="4" captionText="FIGURES 1 – 14. Pronotum of Chlaenius species from Rajasthan: 1. C. laevipennis; 2. C. nitidicollis; 3. C. hamifer; 4. C. scapularis; 5. C. malachinus; 6. C. orbicollis; 7. C. posticus; 8. C. distigma; 9. Chlaenius udaipurensis sp. nov.; 10. C. chlorodius; 11. C. pretiosus; 12. C. quadricolor; 13. C. velocipes; 14. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344754/files/figure.png" pageId="20" pageNumber="565">Figs. 14</figureCitation>
, 
<figureCitation id="BED791C7D478FFE0FED9FB205C706E81" box="[264,294,1198,1222]" captionStart="FIGURES 15 – 28" captionStartId="5.[151,264,1941,1964]" captionTargetBox="[252,1360,239,1900]" captionTargetId="figure@5.[207,1379,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 28. Labrum of Chlaenius species from Rajasthan: 15. C. laevipennis; 16. C. nitidicollis; 17. C. hamifer; 18. C. scapularis; 19. C. malachinus; 20. C. orbicollis; 21. C. posticus; 22. C. distigma; 23. Chlaenius udaipurensis sp. nov.; 24. C. chlorodius; 25. C. pretiosus; 26. C. quadricolor; 27. C. velocipes; 28. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344755/files/figure.png" pageId="20" pageNumber="565">28</figureCitation>
, 
<figureCitation id="BED791C7D478FFE0FEE3FB205C076E81" box="[306,337,1198,1222]" captionStart="FIGURES 29 – 42" captionStartId="6.[151,264,1941,1964]" captionTargetBox="[250,1382,237,1884]" captionTargetId="figure@6.[185,1402,193,1921]" captionTargetPageId="6" captionText="FIGURES 29 – 42. Mentum of Chlaenius species from Rajasthan: 29. C. laevipennis; 30. C. nitidicollis; 31. C. hamifer; 32. C. scapularis; 33. C. malachinus; 34. C. orbicollis; 35. C. posticus; 36. C. distigma; 37. Chlaenius udaipurensis sp. nov.; 38. C. chlorodius; 39. C. pretiosus; 40. C. quadricolor; 41. C. velocipes; 42. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344756/files/figure.png" pageId="20" pageNumber="565">42</figureCitation>
, 
<figureCitation id="BED791C7D478FFE0FE8CFB235C2D6E81" box="[349,379,1197,1222]" captionStart="FIGURES 43 – 57" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[233,1286,236,1818]" captionTargetId="figure@7.[181,1405,193,1892]" captionTargetPageId="7" captionText="FIGURES 43 – 57. Maxillary palpi of Chlaenius species from Rajasthan: 43. C. laevipennis; 44. C. nitidicollis; 45. C. hamifer; 46. C. scapularis; 47. C. malachinus; 48. C. orbicollis (♂); 49. C. orbicollis (♀); 50. C. posticus; 51. C. distigma; 52. Chlaenius udaipurensis sp. nov.; 53. C. chlorodius; 54. C. pretiosus; 55. C. quadricolor; 56. C. velocipes; 57. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344757/files/figure.png" pageId="20" pageNumber="565">57</figureCitation>
, 
<figureCitation id="BED791C7D478FFE0FE57FB205CF26E81" box="[390,420,1198,1222]" captionStart="FIGURES 58 – 71" captionStartId="8.[151,264,1941,1964]" captionTargetBox="[324,1264,204,1913]" captionTargetId="figure@8.[258,1329,193,1921]" captionTargetPageId="8" captionText="FIGURES 58 – 71. Labial palpi of Chlaenius species from Rajasthan: 58. C. laevipennis; 59. C. nitidicollis; 60. C. hamifer; 61. C. scapularis; 62. C. malachinus; 63. C. orbicollis; 64. C. posticus; 65. C. distigma; 66. Chlaenius udaipurensis sp. nov.; 67. C. chlorodius; 68. C. pretiosus; 69. C. quadricolor; 70. C. velocipes; 71. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344758/files/figure.png" pageId="20" pageNumber="565">71</figureCitation>
, 
<figureCitation id="BED791C7D478FFE0FE61FB205CA06E81" box="[432,502,1198,1222]" captionStart="FIGURES 72 – 99" captionStartId="9.[151,264,1759,1782]" captionTargetBox="[274,1348,247,1729]" captionTargetId="figure@9.[208,1366,193,1746]" captionTargetPageId="9" captionText="FIGURES 72 – 99. Male genitalia of Chlaenius species from Rajasthan: 72. C. laevipennis (left lateral view); 73. C. laevipennis (dorsal view); 74. C. nitidicollis (left lateral view); 75. C. nitidicollis (dorsal view); 76. C. hamifer (left lateral view); 77. C. hamifer (dorsal view); 78. C. scapularis (left lateral view); 79. C. scapularis (dorsal view); 80. C. malachinus (left lateral view); 81. C. malachinus (dorsal view); 82. C. orbicollis (left lateral view); 83. C. orbicollis (dorsal view); 84. C. posticus (left lateral view); 85. C. posticus (dorsal view); 86. C. distigma (left lateral view); 87. C. distigma (dorsal view); 88. Chlaenius udaipurensis sp. nov. (left lateral view); 89. Chlaenius udaipurensis sp. nov. (dorsal view); 90. C. chlorodius (left lateral view); 91. C. chlorodius (dorsal view); 92. C. pretiosus (left lateral view); 93. C. pretiosus (dorsal view); 94. C. quadricolor (left lateral view); 95. C. quadricolor (dorsal view); 96. C. velocipes (left lateral view); 97. C. velocipes (dorsal view); 98. Chlaenius pseudotristis sp. nov. (left lateral view); 99. Chlaenius pseudotristis sp. nov. (dorsal view)" httpUri="https://zenodo.org/record/344759/files/figure.png" pageId="20" pageNumber="565">98, 99</figureCitation>
&amp; 
<figureCitation id="BED791C7D478FFE0FDCBFB205F126E81" box="[538,580,1198,1222]" captionStart="FIGURES 112 – 113" captionStartId="17.[151,264,1018,1041]" captionTargetBox="[382,1209,227,975]" captionTargetId="figure@17.[321,1266,195,997]" captionTargetPageId="17" captionText="FIGURES 112 – 113. Habitus of Chlaenius taxa from Rajasthan. 112. C. velocipes; 113. Chlaenius pseudotristis sp. nov." httpUri="https://zenodo.org/record/344762/files/figure.png" pageId="20" pageNumber="565">113</figureCitation>
)
</paragraph>
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series.
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: 
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, 
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, RCA 
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Longitude), 
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(white label), 
<taxonomicName id="E1ECF6C1D478FFE0FD1FFA945D996F11" authority="Chanu &amp; Swaminathan 2016" authorityName="Chanu &amp; Swaminathan" authorityYear="2016" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="565" phylum="Arthropoda" rank="species" species="pseudotristis" subGenus="Chlaeniellus">
<emphasis id="14985150D478FFE0FD1FFA9459D26F75" box="[718,1156,1306,1330]" italics="true" pageId="20" pageNumber="565">Chlaenius (Chlaeniellus) pseudotristis</emphasis>
Chanu &amp; Swaminathan 2016
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College of 
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, 
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and 
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, 
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, India.
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<paragraph id="26538D42D478FFE0FF16FA275C656C32" blockId="20.[151,1436,1269,2014]" pageId="20" pageNumber="565">
<emphasis id="14985150D478FFE0FF16FA275C176F85" bold="true" box="[199,321,1449,1474]" pageId="20" pageNumber="565">Diagnosis.</emphasis>
Elytra black with greenish bluish tinge, no fasciae or spots, intervals punctate pubescent; pronotum wider than long, rounded lateral sides, surface confluently coarsely punctate with wrinkles, few setiferous punctures here and there; median tooth of mentum truncate at apex with rounded angles; labrum slightly emarginate at apex; apical segment of maxillary and labial palpi sub-cylindrical and coniform respectively; aedeagus stout, median lobe almost cylindrical, lamella rounded at apex, both sides of median lobe equally tapers towards apex.
</paragraph>
</subSubSection>
<subSubSection id="6EF6DEC9D478FFE1FF16F90F59A86B97" lastPageId="21" lastPageNumber="566" pageId="20" pageNumber="565" type="description">
<paragraph id="26538D42D478FFE0FF16F90F58D26CDE" blockId="20.[151,1436,1269,2014]" box="[199,1412,1665,1690]" pageId="20" pageNumber="565">
<emphasis id="14985150D478FFE0FF16F90F5C0F6CDD" bold="true" box="[199,345,1665,1690]" pageId="20" pageNumber="565">Description.</emphasis>
BL = 
<quantity id="E11420A7D478FFE0FE70F90C5F436CDE" box="[417,533,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="1.302" pageId="20" pageNumber="565" unit="mm" value="13.02">13.02 mm</quantity>
, HW= 
<quantity id="E11420A7D478FFE0FDB7F90F5F9A6CDE" box="[614,716,1665,1690]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="20" pageNumber="565" unit="mm" value="2.6">2.60 mm</quantity>
, PL= 
<quantity id="E11420A7D478FFE0FCDEF90C5E236CDE" box="[783,885,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="2.74" pageId="20" pageNumber="565" unit="mm" value="2.74">2.74 mm</quantity>
, PW= 
<quantity id="E11420A7D478FFE0FC10F90C59716CDE" box="[961,1063,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="565" unit="mm" value="3.8">3.80 mm</quantity>
, EL= 
<quantity id="E11420A7D478FFE0FBBDF90F59846CDE" box="[1132,1234,1665,1690]" metricMagnitude="-3" metricUnit="m" metricValue="7.68" pageId="20" pageNumber="565" unit="mm" value="7.68">7.68 mm</quantity>
, EW= 
<quantity id="E11420A7D478FFE0FAF1F90F58D26CDE" box="[1312,1412,1665,1690]" metricMagnitude="-3" metricUnit="m" metricValue="5.84" pageId="20" pageNumber="565" unit="mm" value="5.84">5.84 mm</quantity>
</paragraph>
<paragraph id="26538D42D478FFE0FF16F92B59186D41" blockId="20.[151,1436,1269,2014]" pageId="20" pageNumber="565">Head and pronotum black with greenish reddish iridescence; elytra black with greenish bluish tinge. Labrum, mandibles, coxae, trochanters and femora dark brown; first 3 antennomeres and tibia pale yellowish brown; remaining antennomeres, maxillary and labial palpi brown. Ventral sternites fuscous.</paragraph>
<paragraph id="26538D42D478FFE0FF16F89F585F6DD1" blockId="20.[151,1436,1269,2014]" pageId="20" pageNumber="565">Head dorsal surface confluently coarsely punctate (small punctures); frontal fovea shallow. Labrum slightly emarginate at apex, apical angles rounded with some setae; clypeus slightly emarginate. Apical segment of maxillary and labial palpi sub-cylindrical and coniform respectively, almost dorsoventrally flattened and slightly tapering towards and truncate at apex. Median tooth of mentum truncate at apex with rounded angles.</paragraph>
<paragraph id="26538D42D478FFE1FF16F82C5CFD6ABF" blockId="20.[151,1436,1269,2014]" lastBlockId="21.[151,1437,151,644]" lastPageId="21" lastPageNumber="566" pageId="20" pageNumber="565">Pronotum quadrate, wider than long, rounded along lateral margins, surface confluently punctate with wrinkles, few punctures with setae here and there, more at basal region; lateral margins bordered, explanate margin rather gently reflexed and increasing in width from apex to base. Basal margin wider than anterior margin; basal angles obtuse but not sharp, apical angles rounded. Median line fine and distinct but do not reach extremities. Basal fovea rounded and deep.</paragraph>
<paragraph id="26538D42D479FFE1FF16FE8A5E936B5A" blockId="21.[151,1437,151,644]" box="[199,965,260,285]" pageId="21" pageNumber="566">Elytra striate punctate; intervals punctate, pubescent and almost flat.</paragraph>
<paragraph id="26538D42D479FFE1FF16FEA65ED16BCF" blockId="21.[151,1437,151,644]" pageId="21" pageNumber="566">Prosternum, proepisternum very finely punctate almost smooth; prosternal process bordered; mesosternum and mesepisternum finely and uniformly punctate; metasternum smooth medially, punctate laterally; metepisternum punctate. Abdominal sternites very finely punctate, almost smooth.</paragraph>
<paragraph id="26538D42D479FFE1FF16FE1A59A86B97" blockId="21.[151,1437,151,644]" pageId="21" pageNumber="566">Aedeagus stout, almost cylindrical; median lobe gradually bent near base; apex of apical lamella rounded; both sides of median lobe equally tapers towards apex on dorsal view; length of apical lamella very short.</paragraph>
</subSubSection>
<subSubSection id="6EF6DEC9D479FFE1FF16FE525837687A" pageId="21" pageNumber="566" type="discussion">
<paragraph id="26538D42D479FFE1FF16FE525837687A" blockId="21.[151,1437,151,644]" pageId="21" pageNumber="566">
<emphasis id="14985150D479FFE1FF16FE525C6D6BB2" bold="true" box="[199,315,476,501]" pageId="21" pageNumber="566">Remarks.</emphasis>
This species appears similar to 
<taxonomicName id="E1ECF6C1D479FFE1FD72FE525E526BB3" box="[675,772,476,500]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="566" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="14985150D479FFE1FD72FE525E526BB3" box="[675,772,476,500]" italics="true" pageId="21" pageNumber="566">C. tristis</emphasis>
</taxonomicName>
; but it differs in having more regular, rounded lateral sides of the pronotum, less pubescence on surface and the presence of wrinkles; the colour of the head, pronotum and elytra is black with a greenish tinge, and antennomeres 1 to 3 are pale yellowish brown, the others are brown.
</paragraph>
</subSubSection>
<subSubSection id="6EF6DEC9D479FFE1FF16FDC95CD368C3" pageId="21" pageNumber="566" type="etymology">
<paragraph id="26538D42D479FFE1FF16FDC95CD368C3" blockId="21.[151,1437,151,644]" pageId="21" pageNumber="566">
<emphasis id="14985150D479FFE1FF16FDC95C186827" bold="true" box="[199,334,583,608]" pageId="21" pageNumber="566">Etymology.</emphasis>
The name of the new species explains the false resemblance with the species 
<taxonomicName id="E1ECF6C1D479FFE1FAFBFDC65C2868C3" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="566" phylum="Arthropoda" rank="species" species="tristis" subGenus="Chlaeniellus">
<emphasis id="14985150D479FFE1FAFBFDC658CA6827" box="[1322,1436,584,608]" italics="true" pageId="21" pageNumber="566">Chlaenius</emphasis>
(
<emphasis id="14985150D479FFE1FF71FDE25C7868C3" box="[160,302,620,644]" italics="true" pageId="21" pageNumber="566">Chlaeniellus</emphasis>
) 
<emphasis id="14985150D479FFE1FEEFFDE35C2868C3" box="[318,382,621,644]" italics="true" pageId="21" pageNumber="566">tristis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>