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<document id="D6E8C2AC2FE6029A7CE7685AFF7AA220" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE50B63EDD9E45DFFA470609" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius" docType="key" docVersion="12" lastPageNumber="115" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="114" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="26D3ADD55ACB70013348FF50343A22E8">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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Key to males of 
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<emphasis id="B91CEAE5AE50B63DDC5645C0FE2E00D9" bold="true" box="[351,501,655,681]" italics="true" pageId="13" pageNumber="114">Parachorius</emphasis>
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<taxonomicName id="4C684D74AE50B63DDD9E4599FEF8009F" box="[151,291,726,751]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE50B63DDD9E4599FEF8009F" box="[151,291,726,751]" italics="true" pageId="13" pageNumber="114">Parachorius</emphasis>
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species are reliably identified only using males. A separate key to the identifiable females is provided in the next section. The species are illustrated in 
<figureCitation id="13532A72AE50B63DDFB045B5FCF40163" box="[697,815,762,787]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-10="FIGURE 16" captionStart-11="FIGURE 17" captionStart-12="FIGURE 18" captionStart-13="FIGURE 19" captionStart-14="FIGURE 20" captionStart-15="FIGURE 21" captionStart-16="FIGURE 22" captionStart-17="FIGURE 23" captionStart-18="FIGURE 24" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStart-5="FIGURE 11" captionStart-6="FIGURE 12" captionStart-7="FIGURE 13" captionStart-8="FIGURE 14" captionStart-9="FIGURE 15" captionStartId-0="16.[151,250,1359,1381]" captionStartId-1="17.[151,250,1493,1515]" captionStartId-10="30.[151,250,803,825]" captionStartId-11="31.[151,250,1575,1597]" captionStartId-12="33.[151,250,1485,1507]" captionStartId-13="35.[151,250,1502,1524]" captionStartId-14="36.[151,250,1708,1730]" captionStartId-15="37.[151,250,1909,1931]" captionStartId-16="39.[151,250,1152,1174]" captionStartId-17="40.[151,250,1887,1909]" captionStartId-18="41.[151,250,1951,1973]" captionStartId-2="19.[151,250,1935,1957]" captionStartId-3="21.[151,250,1325,1347]" captionStartId-4="22.[151,250,1898,1920]" captionStartId-5="23.[151,250,1984,2006]" captionStartId-6="25.[151,250,1061,1083]" captionStartId-7="26.[151,250,1259,1281]" captionStartId-8="27.[151,250,1980,2002]" captionStartId-9="29.[151,250,1247,1269]" captionTargetBox-0="[151,1436,193,1325]" captionTargetBox-1="[153,1407,193,1475]" captionTargetBox-10="[346,1241,307,778]" captionTargetBox-11="[151,1416,301,1538]" captionTargetBox-12="[151,1431,628,1457]" captionTargetBox-13="[151,1436,473,1484]" captionTargetBox-14="[151,1420,733,1677]" captionTargetBox-15="[151,1436,973,1878]" captionTargetBox-16="[151,1430,195,1127]" captionTargetBox-17="[151,1423,840,1815]" captionTargetBox-18="[168,1425,1036,1927]" captionTargetBox-2="[151,1419,233,1853]" captionTargetBox-3="[151,1436,339,1299]" captionTargetBox-4="[151,1428,799,1834]" captionTargetBox-5="[151,1410,826,1961]" captionTargetBox-6="[151,1426,303,1030]" captionTargetBox-7="[151,1416,193,1211]" captionTargetBox-8="[164,1415,1417,1938]" captionTargetBox-9="[151,1436,267,1223]" captionTargetId-0="figure@16.[151,1436,193,1338]" captionTargetId-1="figure@17.[151,1436,193,1479]" captionTargetId-10="figure@30.[340,1247,303,781]" captionTargetId-11="figure@31.[151,1436,301,1553]" captionTargetId-12="figure@33.[151,1436,628,1464]" captionTargetId-13="figure@35.[151,1436,473,1491]" captionTargetId-14="figure@36.[151,1436,733,1687]" captionTargetId-15="figure@37.[151,1436,973,1888]" captionTargetId-16="figure@39.[151,1436,195,1131]" captionTargetId-17="figure@40.[151,1436,840,1830]" captionTargetId-18="figure@41.[168,1432,1036,1939]" captionTargetId-2="figure@19.[151,1436,233,1866]" captionTargetId-3="figure@21.[151,1436,339,1320]" captionTargetId-4="figure@22.[151,1436,799,1851]" captionTargetId-5="figure@23.[151,1436,826,1969]" captionTargetId-6="figure@25.[151,1436,303,1040]" captionTargetId-7="figure@26.[151,1436,193,1237]" captionTargetId-8="figure@27.[164,1423,1417,1941]" captionTargetId-9="figure@29.[151,1436,267,1225]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-10="30" captionTargetPageId-11="31" captionTargetPageId-12="33" captionTargetPageId-13="35" captionTargetPageId-14="36" captionTargetPageId-15="37" captionTargetPageId-16="39" captionTargetPageId-17="40" captionTargetPageId-18="41" captionTargetPageId-2="19" captionTargetPageId-3="21" captionTargetPageId-4="22" captionTargetPageId-5="23" captionTargetPageId-6="25" captionTargetPageId-7="26" captionTargetPageId-8="27" captionTargetPageId-9="29" captionText-0="FIGURE 6. Morphology of Parachorius asymmetricus. (A) aedeagus, left lateral view (holotype). (B) aedeagus, left lateral view (paratype). (C) aedeagus, right lateral view (holotype). (D) aedeagus, dorsal view (holotype). (E) habitus (male, holotype). (F) right metafemur (male, holotype). (G) protibia (putative female). (H) protibia (male, holotype)." captionText-1="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." captionText-10="FIGURE 16. Morphology of Parachorius maruyamai according to Masumoto et al. (2012). (A) habitus (male, holotype). (B) head and pronotum (dorsal view, scale 1 mm). (C) left metafemur (male). (D) aedeagus." captionText-11="FIGURE 17. Morphology of Parachorius newthayerae. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype)." captionText-12="FIGURE 18. Morphology of Parachorius nudus. (A, C) aedeagus, right lateral view (holotype). (B, D) aedeagus, left lateral view (holotype). (E) protibia (male, holotype). (F) habitus (male, holotype)." captionText-13="FIGURE 19. Morphology of Parachorius peninsularis. (A) aedeagus, lateral view (lectotype). (B) aedeagus, lateral view (Malay, Kedah). (C) aedeagus, lateral view (Sumatra). (D) aedeagus, lateral view (Thailand, Thung Yai). (E) habitus (male, lectotype). (F – H) right metafemur (male). (I) aedeagus, lateral view." captionText-14="FIGURE 20. Morphology of Parachorius pseudojavanus. (A) aedeagus lateral view (male, holotype). (B) aedeagus lateral view (male, paratype). (C) habitus (male, holotype). (D) right metafemur (male). (E) protibia (male). (F) aedeagus lateral view." captionText-15="FIGURE 21. Morphology of Parachorius schuelkei. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype)." captionText-16="FIGURE 22. Morphology of Parachorius semsanganus. (A) aedeagus, lateral view (paratype). (B) aedeagus, dorsal view (paratype). (C) aedeagus, apical view (paratype). (D) aedeagus lateral view. (E) right metafemur (male, paratype). (F) habitus (male, holotype)." captionText-17="FIGURE 23. Morphology of Parachorius solodovnikovi. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Thailand). (C) aedeagus, lateral view (paratype, Vietnam). (D) aedeagus, lateral view (E) habitus (male, holotype). (F) left metafemur (male, paratype, Vietnam) (G) left metafemur (male, holotype). (H) left protibia (male, holotype, original image flipped)." captionText-18="FIGURE 24. Morphology of Parachorius thomsoni. (A) aedeagus, lateral view (P. thomsoni holotype, male) (B) aedeagus, lateral view (P. lannathai, holotype, male). (C – D) aedeagus, lateral view. (E) left metafemur (P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (P. lannathai, holotype, male). (H) habitus (P. thomsoni, holotype, male)." captionText-2="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." captionText-3="FIGURE 9. Morphology of Parachorius fungorum. (A – C, G) aedeagus, lateral view. (D) habitus (male). (E-F) left metafemur (male)." captionText-4="FIGURE 10. Morphology of Parachorius globosus. (A – C) aedeagus. (D) habitus (male). (E) habitus (female, holotype). (F) disc of left elytron. (G) protibia (female)" captionText-5="FIGURE 11. Morphology of Parachorius gotoi. (A, C) aedeagus. (B) left metafemur (male). (D) protibia (male). (E) habitus (male)." captionText-6="FIGURE 12. Morphology of Parachorius hookeri. (A) aedeagus, lateral view (holotype). (B) aedeagus lateral view. (C) left metafemur (male, holotype). (D) habitus (male, holotype)" captionText-7="FIGURE 13. Morphology of Parachorius humeralis. (A – B) aedeagus (Darjeeling). (C) aedeagus. (D) habitus (male, holotype). (E) habitus (male). (F – H) right metafemur (male). (I) protibia (female). (J) protibia (male)." captionText-8="FIGURE 14. Morphology of Parachorius javanus. (A, E) aedeagus lateral view (male, holotype). (B) habitus (male, holotype). (C – D) protibia (male, holotype)" captionText-9="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri-0="https://zenodo.org/record/1001927/files/figure.png" httpUri-1="https://zenodo.org/record/1001929/files/figure.png" httpUri-10="https://zenodo.org/record/1001949/files/figure.png" httpUri-11="https://zenodo.org/record/1001951/files/figure.png" httpUri-12="https://zenodo.org/record/1001953/files/figure.png" httpUri-13="https://zenodo.org/record/1001955/files/figure.png" httpUri-14="https://zenodo.org/record/1001957/files/figure.png" httpUri-15="https://zenodo.org/record/1001959/files/figure.png" httpUri-16="https://zenodo.org/record/1001961/files/figure.png" httpUri-17="https://zenodo.org/record/1001963/files/figure.png" httpUri-18="https://zenodo.org/record/1001965/files/figure.png" httpUri-2="https://zenodo.org/record/1001931/files/figure.png" httpUri-3="https://zenodo.org/record/1001935/files/figure.png" httpUri-4="https://zenodo.org/record/1001937/files/figure.png" httpUri-5="https://zenodo.org/record/1001939/files/figure.png" httpUri-6="https://zenodo.org/record/1001941/files/figure.png" httpUri-7="https://zenodo.org/record/1001943/files/figure.png" httpUri-8="https://zenodo.org/record/1001945/files/figure.png" httpUri-9="https://zenodo.org/record/1001947/files/figure.png" pageId="13" pageNumber="114">Figs. 6–24</figureCitation>
and their distribution maps are given in 
<figureCitation id="13532A72AE50B63DD9FC45B5FA5A0163" box="[1269,1409,762,787]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStartId-0="42.[151,250,1911,1933]" captionStartId-1="43.[151,250,1905,1927]" captionStartId-2="44.[151,250,1770,1792]" captionTargetBox-0="[151,1433,273,1824]" captionTargetBox-1="[151,1436,265,1825]" captionTargetBox-2="[151,1436,193,1748]" captionTargetId-0="figure@42.[151,1436,273,1824]" captionTargetId-1="figure@43.[151,1436,265,1825]" captionTargetId-2="figure@44.[151,1436,193,1748]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionTargetPageId-2="44" captionText-0="FIGURE 25. Distribution maps of Parachorius species. (A) Parachorius asymmetricus. (B) Parachorius bolavensis. (C) Parachorius fukiensis. (D) Parachorius fungorum. (E) Parachorius globosus. (F) Parachorius gotoi. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." captionText-1="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." captionText-2="FIGURE 27. Distribution maps of Parachorius species. (A) Parachorius peninsularis. (B) Parachorius pseudojavanus. (C) Parachorius schuelkei. (D) Parachorius semsanganus. (E) Parachorius solodovnikovi. (F) Parachorius thomsoni. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study. Literature data — localities retrieved from literature." httpUri-0="https://zenodo.org/record/1001967/files/figure.png" httpUri-1="https://zenodo.org/record/1001969/files/figure.png" httpUri-2="https://zenodo.org/record/1001971/files/figure.png" pageId="13" pageNumber="114">Figs. 25–27.</figureCitation>
</paragraph>
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<keyLead id="309995C2AE50B63DDD9E440DFA470127" box="[151,1436,834,855]" pageId="13" pageNumber="114">
1 Clypeus with two teeth, outer margin of each tooth not notched basally (
<figureCitation id="13532A72AE50B63DDE9F440DFBD90127" box="[918,1026,834,855]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="13" pageNumber="114">Fig. 4F–G</figureCitation>
).................................2
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- Clypeus with two teeth, outer margin of each tooth distinctly or slightly notched basally (
<figureCitation id="13532A72AE50B63DD9314410FB7A0104" box="[1080,1185,863,884]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="13" pageNumber="114">Fig. 4D–E</figureCitation>
)..................10
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2 Clypeus with two distinctly produced teeth (
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).........................................................3
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</paragraph>
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- Clypeus with two less produced teeth (
<figureCitation id="13532A72AE50B63DDF5D44D5FD7A01DF" box="[596,673,922,943]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="13" pageNumber="114">Fig. 4F</figureCitation>
............................................................... 
<figureCitation id="13532A72AE50B63DD88344D5FA4001DF" box="[1418,1435,922,943]" captionStart="FIGURE 5" captionStartId="12.[151,250,1891,1913]" captionTargetBox="[162,1421,892,1854]" captionTargetId="figure@12.[151,1436,877,1870]" captionTargetPageId="12" captionText="FIGURE 5. Diversity of aedeagal parameres in Parachorius. (A) Parachorius fungorum. (B) Parachorius hookeri. (C) Parachorius schuelkei. (D) Parachorius thomsoni. (E) Parachorius globosus. (F) Parachorius newthayerae. (G) Parachorius bolavensis. (H) Parachorius fukiensis. (I) Parachorius nudus. (J) Parachorius peninsularis. (K) Parachorius pseudojavanus. (L) Parachorius javanus. (M) Parachorius solodovnikovi. (N) Parachorius longipenis. (O) Parachorius humeralis. (P) Parachorius semsanganus. (Q) Parachorius gotoi." httpUri="https://zenodo.org/record/1001925/files/figure.png" pageId="13" pageNumber="114">5</figureCitation>
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3 Body black; parameres widely rounded; head triangularly shaped (
<figureCitation id="13532A72AE50B63DDE5644F8FC7D01BC" box="[863,934,951,972]" captionStart="FIGURE 21" captionStartId="37.[151,250,1909,1931]" captionTargetBox="[151,1436,973,1878]" captionTargetId="figure@37.[151,1436,973,1888]" captionTargetPageId="37" captionText="FIGURE 21. Morphology of Parachorius schuelkei. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype)." httpUri="https://zenodo.org/record/1001959/files/figure.png" pageId="13" pageNumber="114">Fig. 21</figureCitation>
); size 
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; southern China........................................................................................ 
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Tarasov
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<emphasis id="B91CEAE5AE50B63DD82F449BFA40019A" bold="true" box="[1318,1435,980,1002]" pageId="13" pageNumber="114">new species</emphasis>
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<paragraph id="8BD736F7AE50B63DDD9E44BDFA400654" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
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- Body reddish brown, rarely dark brown; parameres either claw shaped or apically acute but not widely rounded; head more rounded and not as distinctly triangular as in 
<taxonomicName id="4C684D74AE50B63DDF9B4340FCD90654" ID-CoL="6TNLX" authorityName="S. Tarasov" authorityYear="2017" box="[658,770,1039,1060]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="B91CEAE5AE50B63DDF9B4340FCD90654" box="[658,770,1039,1060]" italics="true" pageId="13" pageNumber="114">P. schuelkei</emphasis>
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......................................................4
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4 Protibia with first tooth somewhat modified in male—slightly thickened and spatulate; parameres notched basally and claw shaped in lateral view (
<figureCitation id="13532A72AE50B63DDCB64305FDDD062F" box="[447,518,1098,1119]" captionStart="FIGURE 14" captionStartId="27.[151,250,1980,2002]" captionTargetBox="[164,1415,1417,1938]" captionTargetId="figure@27.[164,1423,1417,1941]" captionTargetPageId="27" captionText="FIGURE 14. Morphology of Parachorius javanus. (A, E) aedeagus lateral view (male, holotype). (B) habitus (male, holotype). (C – D) protibia (male, holotype)" httpUri="https://zenodo.org/record/1001945/files/figure.png" pageId="13" pageNumber="114">Fig. 14</figureCitation>
); length 
<quantity id="4C909B12AE50B63DDF524305FD7C062F" box="[603,679,1098,1119]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="13" pageNumber="114" unit="mm" value="6.7">6.7 mm</quantity>
; known only from the 
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; 
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........ 
<taxonomicName id="4C684D74AE50B63DD9714305FA47062F" ID-CoL="6TNM6" authority="(Boucomont, 1914)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[1144,1436,1098,1119]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B91CEAE5AE50B63DD9714305FB01062F" box="[1144,1242,1098,1119]" italics="true" pageId="13" pageNumber="114">P. javanus</emphasis>
(
<bibRefCitation id="EFF94B06AE50B63DD9EF4305FA4E062F" author="Boucomont" box="[1254,1429,1098,1119]" pageId="13" pageNumber="114" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
)
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</paragraph>
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- Protibia first tooth not modified in males; parameres medially narrowed (in lateral view), apically acute (
<figureCitation id="13532A72AE50B63DD9D34328FAF9060C" box="[1242,1314,1127,1148]" captionStart="FIGURE 20" captionStartId="36.[151,250,1708,1730]" captionTargetBox="[151,1420,733,1677]" captionTargetId="figure@36.[151,1436,733,1687]" captionTargetPageId="36" captionText="FIGURE 20. Morphology of Parachorius pseudojavanus. (A) aedeagus lateral view (male, holotype). (B) aedeagus lateral view (male, paratype). (C) habitus (male, holotype). (D) right metafemur (male). (E) protibia (male). (F) aedeagus lateral view." httpUri="https://zenodo.org/record/1001957/files/figure.png" pageId="13" pageNumber="114">Fig. 20</figureCitation>
); length 
<quantity id="4C909B12AE50B63DD87E4328FED206EA" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="13" pageNumber="114" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
; 
<collectingRegion id="49ACF815AE50B63DDC1A43CAFEE506EA" box="[275,318,1157,1178]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="114">Java</collectingRegion>
............................................................. 
<taxonomicName id="4C684D74AE50B63DD92E43CAFAC406EA" ID-CoL="75NM8" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1063,1311,1157,1178]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="pseudojavanus" status="sp. nov.">
<emphasis id="B91CEAE5AE50B63DD92E43CAFB1406EA" box="[1063,1231,1157,1178]" italics="true" pageId="13" pageNumber="114">P. pseudojavanus</emphasis>
Tarasov
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<emphasis id="B91CEAE5AE50B63DD82F43CBFA4006EA" bold="true" box="[1318,1435,1156,1178]" pageId="13" pageNumber="114">new species</emphasis>
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5 Metatibia strongly curved and notched medially (
<figureCitation id="13532A72AE50B63DDFCD43EDFCFE06C7" box="[708,805,1186,1207]" captionStart="FIGURE 16" captionStartId="30.[151,250,803,825]" captionTargetBox="[346,1241,307,778]" captionTargetId="figure@30.[340,1247,303,781]" captionTargetPageId="30" captionText="FIGURE 16. Morphology of Parachorius maruyamai according to Masumoto et al. (2012). (A) habitus (male, holotype). (B) head and pronotum (dorsal view, scale 1 mm). (C) left metafemur (male). (D) aedeagus." httpUri="https://zenodo.org/record/1001949/files/figure.png" pageId="13" pageNumber="114">Figs. 16C</figureCitation>
, 
<figureCitation id="13532A72AE50B63DDE3843EDFC8506C7" box="[817,862,1186,1207]" captionStart="FIGURE 17" captionStartId="31.[151,250,1575,1597]" captionTargetBox="[151,1416,301,1538]" captionTargetId="figure@31.[151,1436,301,1553]" captionTargetPageId="31" captionText="FIGURE 17. Morphology of Parachorius newthayerae. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype)." httpUri="https://zenodo.org/record/1001951/files/figure.png" pageId="13" pageNumber="114">17D</figureCitation>
); frontal angles of pronotum from moderately to distinctly sharp (
<figureCitation id="13532A72AE50B63DDC2B43F0FE5906A4" box="[290,386,1215,1236]" captionStart="FIGURE 16" captionStartId="30.[151,250,803,825]" captionTargetBox="[346,1241,307,778]" captionTargetId="figure@30.[340,1247,303,781]" captionTargetPageId="30" captionText="FIGURE 16. Morphology of Parachorius maruyamai according to Masumoto et al. (2012). (A) habitus (male, holotype). (B) head and pronotum (dorsal view, scale 1 mm). (C) left metafemur (male). (D) aedeagus." httpUri="https://zenodo.org/record/1001949/files/figure.png" pageId="13" pageNumber="114">Figs. 16B</figureCitation>
, 
<figureCitation id="13532A72AE50B63DDC8543F0FE3206A4" box="[396,489,1215,1236]" captionStart="FIGURE 17" captionStartId="31.[151,250,1575,1597]" captionTargetBox="[151,1416,301,1538]" captionTargetId="figure@31.[151,1436,301,1553]" captionTargetPageId="31" captionText="FIGURE 17. Morphology of Parachorius newthayerae. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype)." httpUri="https://zenodo.org/record/1001951/files/figure.png" pageId="13" pageNumber="114">17C, E–F</figureCitation>
).............................................................................6
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</paragraph>
<paragraph id="8BD736F7AE50B63DDD9E4392FA470682" blockId="13.[151,1436,834,2028]" box="[151,1436,1245,1266]" pageId="13" pageNumber="114">
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- Metatibia slightly sinuate (
<figureCitation id="13532A72AE50B63DDCEB4392FDE30682" box="[482,568,1245,1266]" captionStart="FIGURE 24" captionStartId="41.[151,250,1951,1973]" captionTargetBox="[168,1425,1036,1927]" captionTargetId="figure@41.[168,1432,1036,1939]" captionTargetPageId="41" captionText="FIGURE 24. Morphology of Parachorius thomsoni. (A) aedeagus, lateral view (P. thomsoni holotype, male) (B) aedeagus, lateral view (P. lannathai, holotype, male). (C – D) aedeagus, lateral view. (E) left metafemur (P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (P. lannathai, holotype, male). (H) habitus (P. thomsoni, holotype, male)." httpUri="https://zenodo.org/record/1001965/files/figure.png" pageId="13" pageNumber="114">Fig. 24E</figureCitation>
); frontal angles of pronotum rounded (
<figureCitation id="13532A72AE50B63DDEA24392FBDE0682" box="[939,1029,1245,1266]" captionStart="FIGURE 24" captionStartId="41.[151,250,1951,1973]" captionTargetBox="[168,1425,1036,1927]" captionTargetId="figure@41.[168,1432,1036,1939]" captionTargetPageId="41" captionText="FIGURE 24. Morphology of Parachorius thomsoni. (A) aedeagus, lateral view (P. thomsoni holotype, male) (B) aedeagus, lateral view (P. lannathai, holotype, male). (C – D) aedeagus, lateral view. (E) left metafemur (P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (P. lannathai, holotype, male). (H) habitus (P. thomsoni, holotype, male)." httpUri="https://zenodo.org/record/1001965/files/figure.png" pageId="13" pageNumber="114">Fig. 24G</figureCitation>
)................................7
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<paragraph id="8BD736F7AE50B63DDD9E43B5FA40075D" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
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6 Middle part of metafemur dilated posteriorly and produced as a large, rounded angle; pronotum distinctly angulate laterally with frontal angles moderately sharp (
<figureCitation id="13532A72AE50B63DDF444258FD4F075C" box="[589,660,1303,1324]" captionStart="FIGURE 17" captionStartId="31.[151,250,1575,1597]" captionTargetBox="[151,1416,301,1538]" captionTargetId="figure@31.[151,1436,301,1553]" captionTargetPageId="31" captionText="FIGURE 17. Morphology of Parachorius newthayerae. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype)." httpUri="https://zenodo.org/record/1001951/files/figure.png" pageId="13" pageNumber="114">Fig. 17</figureCitation>
); length 7.8–8.0 mm; southern Laos...... 
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<emphasis id="B91CEAE5AE50B63DD9354258FB14075C" box="[1084,1231,1303,1324]" italics="true" pageId="13" pageNumber="114">P. newthayerae</emphasis>
Tarasov
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<emphasis id="B91CEAE5AE50B63DD82F4258FA40075D" bold="true" box="[1318,1435,1303,1325]" pageId="13" pageNumber="114">new species</emphasis>
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<paragraph id="8BD736F7AE50B63DDD9E427AFA4707F4" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
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- Metafemoral posterior margin simple, metafemur not dilated; pronotum presumably (as none of the specimens were studied) not angulate laterally with frontal angles distinctly sharp (
<figureCitation id="13532A72AE50B63DDE02421DFC890717" box="[779,850,1362,1383]" captionStart="FIGURE 16" captionStartId="30.[151,250,803,825]" captionTargetBox="[346,1241,307,778]" captionTargetId="figure@30.[340,1247,303,781]" captionTargetPageId="30" captionText="FIGURE 16. Morphology of Parachorius maruyamai according to Masumoto et al. (2012). (A) habitus (male, holotype). (B) head and pronotum (dorsal view, scale 1 mm). (C) left metafemur (male). (D) aedeagus." httpUri="https://zenodo.org/record/1001949/files/figure.png" pageId="13" pageNumber="114">Fig. 16</figureCitation>
); length 
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; central Thailand.......................................................................... 
<taxonomicName id="4C684D74AE50B63DDE8E4220FB6D07F4" ID-CoL="75NLV" authority="Masumoto, Ochi" authorityName="Masumoto, Ochi" box="[903,1206,1391,1412]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="maruyamai">
<emphasis id="B91CEAE5AE50B63DDE8E4220FBD707F4" box="[903,1036,1391,1412]" italics="true" pageId="13" pageNumber="114">P. maruyamai</emphasis>
Masumoto, Ochi
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, &amp; Sakchoowong, 2012
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7 Elytra covered with dense contiguous and rugose punctures (
<figureCitation id="13532A72AE50B63DDE2E42C2FCA607D2" box="[807,893,1421,1442]" captionStart="FIGURE 10" captionStartId="22.[151,250,1898,1920]" captionTargetBox="[151,1428,799,1834]" captionTargetId="figure@22.[151,1436,799,1851]" captionTargetPageId="22" captionText="FIGURE 10. Morphology of Parachorius globosus. (A – C) aedeagus. (D) habitus (male). (E) habitus (female, holotype). (F) disc of left elytron. (G) protibia (female)" httpUri="https://zenodo.org/record/1001937/files/figure.png" pageId="13" pageNumber="114">Fig. 10F</figureCitation>
); pronotum almost as long as elytra; wings reduced (
<figureCitation id="13532A72AE50B63DD87C42C2FF2707CF" captionStart="FIGURE 10" captionStartId="22.[151,250,1898,1920]" captionTargetBox="[151,1428,799,1834]" captionTargetId="figure@22.[151,1436,799,1851]" captionTargetPageId="22" captionText="FIGURE 10. Morphology of Parachorius globosus. (A – C) aedeagus. (D) habitus (male). (E) habitus (female, holotype). (F) disc of left elytron. (G) protibia (female)" httpUri="https://zenodo.org/record/1001937/files/figure.png" pageId="13" pageNumber="114">Fig. 10</figureCitation>
); 
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; northeastern India....................................................... 
<taxonomicName id="4C684D74AE50B63DD9A642E5FA4007CF" ID-CoL="6TNLT" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1199,1435,1450,1471]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B91CEAE5AE50B63DD9A642E5FAC707CF" box="[1199,1308,1450,1471]" italics="true" pageId="13" pageNumber="114">P. globosus</emphasis>
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</keyLead>
</paragraph>
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<keyLead id="309995C2AE50B63DDD9E4288FA40078A" pageId="13" pageNumber="114">
- Elytra covered with sparse (not contiguous) punctures (
<figureCitation id="13532A72AE50B63DDFFB4288FC9307AC" box="[754,840,1479,1500]" captionStart="FIGURE 24" captionStartId="41.[151,250,1951,1973]" captionTargetBox="[168,1425,1036,1927]" captionTargetId="figure@41.[168,1432,1036,1939]" captionTargetPageId="41" captionText="FIGURE 24. Morphology of Parachorius thomsoni. (A) aedeagus, lateral view (P. thomsoni holotype, male) (B) aedeagus, lateral view (P. lannathai, holotype, male). (C – D) aedeagus, lateral view. (E) left metafemur (P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (P. lannathai, holotype, male). (H) habitus (P. thomsoni, holotype, male)." httpUri="https://zenodo.org/record/1001965/files/figure.png" pageId="13" pageNumber="114">Fig. 24F</figureCitation>
); pronotum notably shorter than elytra; wings normally developed............................................................................................... 8
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</keyStep>
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<paragraph id="8BD736F7AE50B63DDD9E414DFA470422" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
<keyLead id="309995C2AE50B63DDD9E414DFA470422" pageId="13" pageNumber="114">
8 Metafemoral posterior margin with two distinct denticles, apical denticle is produced and spur-like; metatibial inner margin excavated apically; metafemoral posterior margin not serrate; parameres of aedeagus simple, apex slightly rounded, triangularly shaped (
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); length 6.5–9.0 mm; eastern 
<collectingCountry id="F37F7667AE50B63DDFCE4172FD200422" box="[711,763,1597,1618]" name="India" pageId="13" pageNumber="114">India</collectingCountry>
, 
<collectingCountry id="F37F7667AE50B63DDE0C4172FCBF0422" box="[773,868,1597,1618]" name="Myanmar" pageId="13" pageNumber="114">Myanmar</collectingCountry>
, 
<collectingCountry id="F37F7667AE50B63DDE644172FC1E0422" box="[877,965,1597,1618]" name="Thailand" pageId="13" pageNumber="114">Thailand</collectingCountry>
, 
<collectingCountry id="F37F7667AE50B63DDEC74172FC260422" box="[974,1021,1597,1618]" name="Laos" pageId="13" pageNumber="114">Laos</collectingCountry>
............. 
<taxonomicName id="4C684D74AE50B63DD9A14172FA470422" ID-CoL="4CPV4" authority="Harold, 1873" authorityName="Harold" authorityYear="1873" box="[1192,1436,1597,1618]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="thomsoni">
<emphasis id="B91CEAE5AE50B63DD9A14172FACC0422" box="[1192,1303,1597,1618]" italics="true" pageId="13" pageNumber="114">P. thomsoni</emphasis>
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</paragraph>
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<keyLead id="309995C2AE50B63DDD9E4115FA4704FC" pageId="13" pageNumber="114">- Metafemoral posterior margin without pair of distinct denticles, simple or dilated, either serrate or not; metatibial inner margin not excavated apically; parameres of aedeagus simple, apex widely rounded and inferiorly angulate....................9</keyLead>
</paragraph>
</keyStep>
<keyStep id="309C2E52AE50B63DDD9E41DAFA40056F" pageId="13" pageNumber="114">
<paragraph id="8BD736F7AE50B63DDD9E41DAFA470494" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
<keyLead id="309995C2AE50B63DDD9E41DAFA470494" pageId="13" pageNumber="114">
9 Metafemoral posterior margin serrate (sometimes slightly) and varies from simple, not modified, to dilated; metatibial inner margin produced in small angle apically (
<figureCitation id="13532A72AE50B63DDF6641FDFD7104B7" box="[623,682,1714,1735]" captionStart="FIGURE 9" captionStartId="21.[151,250,1325,1347]" captionTargetBox="[151,1436,339,1299]" captionTargetId="figure@21.[151,1436,339,1320]" captionTargetPageId="21" captionText="FIGURE 9. Morphology of Parachorius fungorum. (A – C, G) aedeagus, lateral view. (D) habitus (male). (E-F) left metafemur (male)." httpUri="https://zenodo.org/record/1001935/files/figure.png" pageId="13" pageNumber="114">Fig. 9</figureCitation>
); length 6.5–9.0 mm; southern 
<collectingCountry id="F37F7667AE50B63DDEDF41FDFBCA04B7" box="[982,1041,1714,1735]" name="China" pageId="13" pageNumber="114">China</collectingCountry>
, 
<collectingCountry id="F37F7667AE50B63DD91541FDFBA004B7" box="[1052,1147,1714,1735]" name="Myanmar" pageId="13" pageNumber="114">Myanmar</collectingCountry>
, Indochina.............................................................................. 
<taxonomicName id="4C684D74AE50B63DDEDF4180FA470494" ID-CoL="75NM4" authority="Kryzhanovsky &amp; Medvedev, 1966" authorityName="Kryzhanovsky &amp; Medvedev" authorityYear="1966" box="[982,1436,1743,1764]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="fungorum">
<emphasis id="B91CEAE5AE50B63DDEDF4180FB900494" box="[982,1099,1743,1764]" italics="true" pageId="13" pageNumber="114">P. fungorum</emphasis>
<bibRefCitation id="EFF94B06AE50B63DD95B4180FA470494" author="Kryzhanovsky" box="[1106,1436,1743,1764]" pageId="13" pageNumber="114" refString="Kryzhanovsky, O. L. &amp; Medvedev, S. I. (1966) Dung beetles of the subfamily Coprinae in Yunnan province of South-Western China. Part 1. Entomologicheskoe Obozrenie, Revue d'entomologie de l'URSS, 45, 387 - 400." type="journal article" year="1966">Kryzhanovsky &amp; Medvedev, 1966</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BD736F7AE50B63DDD9E41A2FA40056F" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
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- Metafemoral posterior margin simple, not serrate, not dilated; metatibial inner margin straight, not produced apically (
<figureCitation id="13532A72AE50B63DD84141A2FA4B0572" box="[1352,1424,1773,1794]" captionStart="FIGURE 12" captionStartId="25.[151,250,1061,1083]" captionTargetBox="[151,1426,303,1030]" captionTargetId="figure@25.[151,1436,303,1040]" captionTargetPageId="25" captionText="FIGURE 12. Morphology of Parachorius hookeri. (A) aedeagus, lateral view (holotype). (B) aedeagus lateral view. (C) left metafemur (male, holotype). (D) habitus (male, holotype)" httpUri="https://zenodo.org/record/1001941/files/figure.png" pageId="13" pageNumber="114">Fig. 12</figureCitation>
); size 6.0 mm; northern India........................................................... 
<taxonomicName id="4C684D74AE50B63DD9B24045FA40056F" ID-CoL="75NLT" authority="Arrow, 1931" authorityName="Arrow M. E. Bacchus" authorityYear="1931" box="[1211,1435,1802,1823]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="hookeri">
<emphasis id="B91CEAE5AE50B63DD9B24045FAC7056F" box="[1211,1308,1802,1823]" italics="true" pageId="13" pageNumber="114">P. hookeri</emphasis>
<bibRefCitation id="EFF94B06AE50B63DD82A4045FA40056F" author="Arrow" box="[1315,1435,1802,1823]" pageId="13" pageNumber="114" refString="Arrow, G. J. (1931) Coleoptera Lamellicornia part. III. (Coprinae). Taylor and Francis, London, 428 pp., 13 pls." type="book" year="1931">Arrow, 1931</bibRefCitation>
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<paragraph id="8BD736F7AE50B63DDD9E4068FA40054C" blockId="13.[151,1436,834,2028]" box="[151,1435,1831,1852]" pageId="13" pageNumber="114">
<keyLead id="309995C2AE50B63DDD9E4068FA40054C" box="[151,1435,1831,1852]" pageId="13" pageNumber="114">
10 Outer margin of each clypeal tooth slightly notched basally (
<figureCitation id="13532A72AE50B63DDE2F4068FCAA054C" box="[806,881,1831,1852]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="13" pageNumber="114">Fig. 4E</figureCitation>
)............................................ 11
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE50B63DDD9E400AFA40052A" blockId="13.[151,1436,834,2028]" box="[151,1435,1861,1882]" pageId="13" pageNumber="114">
<keyLead id="309995C2AE50B63DDD9E400AFA40052A" box="[151,1435,1861,1882]" pageId="13" pageNumber="114">
- Outer margin of each clypeal tooth distinctly notched basally (
<figureCitation id="13532A72AE50B63DDE3E400AFC59052A" box="[823,898,1861,1882]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="13" pageNumber="114">Fig. 4D</figureCitation>
)......................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309C2E52AE50B63DDD9E402DFA40059C" pageId="13" pageNumber="114">
<paragraph id="8BD736F7AE50B63DDD9E402DFA4705E4" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
<keyLead id="309995C2AE50B63DDD9E402DFA4705E4" pageId="13" pageNumber="114">
11 Inner margin of metatibia denticulate with relatively small teeth; protibia with apical tooth wide, nearly merging with second tooth; parameres elongate and rounded apically; length 
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(
<figureCitation id="13532A72AE50B63DDE5B4030FC4305E4" box="[850,920,1919,1940]" captionStart="FIGURE 11" captionStartId="23.[151,250,1984,2006]" captionTargetBox="[151,1410,826,1961]" captionTargetId="figure@23.[151,1436,826,1969]" captionTargetPageId="23" captionText="FIGURE 11. Morphology of Parachorius gotoi. (A, C) aedeagus. (B) left metafemur (male). (D) protibia (male). (E) habitus (male)." httpUri="https://zenodo.org/record/1001939/files/figure.png" pageId="13" pageNumber="114">Fig. 11</figureCitation>
); Taiwan............. 
<taxonomicName id="4C684D74AE50B63DD9934030FA4705E4" ID-CoL="6TNM5" authority="(Masumoto, 1986)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Masumoto" baseAuthorityYear="1986" box="[1178,1436,1919,1940]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="gotoi">
<emphasis id="B91CEAE5AE50B63DD9934030FB3805E4" box="[1178,1251,1919,1940]" italics="true" pageId="13" pageNumber="114">P. gotoi</emphasis>
(
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)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE50B63DDD9E40D2FA40059C" blockId="13.[151,1436,834,2028]" pageId="13" pageNumber="114">
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- Inner margin of metatibia denticulate with large teeth (sometimes denticulation obliterated); protibia with apical and second teeth distinctly separated from each other; parameres bear membranous lobes and bent upward apically; length 
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(
<figureCitation id="13532A72AE50B63DDDE14098FEEB059C" box="[232,304,2007,2028]" captionStart="FIGURE 22" captionStartId="39.[151,250,1152,1174]" captionTargetBox="[151,1430,195,1127]" captionTargetId="figure@39.[151,1436,195,1131]" captionTargetPageId="39" captionText="FIGURE 22. Morphology of Parachorius semsanganus. (A) aedeagus, lateral view (paratype). (B) aedeagus, dorsal view (paratype). (C) aedeagus, apical view (paratype). (D) aedeagus lateral view. (E) right metafemur (male, paratype). (F) habitus (male, holotype)." httpUri="https://zenodo.org/record/1001961/files/figure.png" pageId="13" pageNumber="114">Fig. 22</figureCitation>
); northern Laos................................................. 
<taxonomicName id="4C684D74AE50B63DD92D4098FA40059C" ID-CoL="6TNM9" authority="Tarasov &amp; Keith, 2011" authorityName="Tarasov &amp; Keith" authorityYear="2011" box="[1060,1435,2007,2028]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="semsanganus">
<emphasis id="B91CEAE5AE50B63DD92D4098FB67059C" box="[1060,1212,2007,2028]" italics="true" pageId="13" pageNumber="114">P. semsanganus</emphasis>
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</paragraph>
</keyStep>
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<paragraph id="8BD736F7AE53B63EDD9E47D8FA4002BA" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
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12 Parameres asymmetrical; humeral area of elytron usually with yellow spot, sometimes yellow spot absent and elytra uniformly colored (
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); length 
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; India.................................... 
<taxonomicName id="4C684D74AE53B63ED93A47FAFAC402BA" ID-CoL="6TNLS" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1075,1311,181,202]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
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Tarasov
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<emphasis id="B91CEAE5AE53B63ED82C47FBFA4002BA" bold="true" box="[1317,1435,180,202]" pageId="14" pageNumber="115">new species</emphasis>
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<paragraph id="8BD736F7AE53B63EDD9E479DFA400297" blockId="14.[151,1436,151,1145]" box="[151,1435,210,231]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E479DFA400297" box="[151,1435,210,231]" pageId="14" pageNumber="115">- Parameres symmetrical; humeral area of elytron with or without yellow spot.....................................13</keyLead>
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<keyLead id="309995C2AE53B63EDD9E47A0FA400374" box="[151,1435,239,260]" pageId="14" pageNumber="115">
13 Parameres with horn-like upper lobes dorsally (
<figureCitation id="13532A72AE53B63EDFA147A0FCCE0374" box="[680,789,239,260]" captionStart="FIGURE 5" captionStartId="12.[151,250,1891,1913]" captionTargetBox="[162,1421,892,1854]" captionTargetId="figure@12.[151,1436,877,1870]" captionTargetPageId="12" captionText="FIGURE 5. Diversity of aedeagal parameres in Parachorius. (A) Parachorius fungorum. (B) Parachorius hookeri. (C) Parachorius schuelkei. (D) Parachorius thomsoni. (E) Parachorius globosus. (F) Parachorius newthayerae. (G) Parachorius bolavensis. (H) Parachorius fukiensis. (I) Parachorius nudus. (J) Parachorius peninsularis. (K) Parachorius pseudojavanus. (L) Parachorius javanus. (M) Parachorius solodovnikovi. (N) Parachorius longipenis. (O) Parachorius humeralis. (P) Parachorius semsanganus. (Q) Parachorius gotoi." httpUri="https://zenodo.org/record/1001925/files/figure.png" pageId="14" pageNumber="115">Fig. 5M–O</figureCitation>
)...................................................14
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE53B63EDD9E4642FA400352" blockId="14.[151,1436,151,1145]" box="[151,1435,269,290]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E4642FA400352" box="[151,1435,269,290]" pageId="14" pageNumber="115">
- Parameres normal, without horn-like upper lobes (
<figureCitation id="13532A72AE53B63EDFCD4642FCF70352" box="[708,812,269,290]" captionStart="FIGURE 5" captionStartId="12.[151,250,1891,1913]" captionTargetBox="[162,1421,892,1854]" captionTargetId="figure@12.[151,1436,877,1870]" captionTargetPageId="12" captionText="FIGURE 5. Diversity of aedeagal parameres in Parachorius. (A) Parachorius fungorum. (B) Parachorius hookeri. (C) Parachorius schuelkei. (D) Parachorius thomsoni. (E) Parachorius globosus. (F) Parachorius newthayerae. (G) Parachorius bolavensis. (H) Parachorius fukiensis. (I) Parachorius nudus. (J) Parachorius peninsularis. (K) Parachorius pseudojavanus. (L) Parachorius javanus. (M) Parachorius solodovnikovi. (N) Parachorius longipenis. (O) Parachorius humeralis. (P) Parachorius semsanganus. (Q) Parachorius gotoi." httpUri="https://zenodo.org/record/1001925/files/figure.png" pageId="14" pageNumber="115">Fig. 5G–H</figureCitation>
).................................................16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309C2E52AE53B63EDD9E4665FA4003C4" pageId="14" pageNumber="115">
<paragraph id="8BD736F7AE53B63EDD9E4665FA47030A" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<materialsCitation id="3B003CAAAE53B63EDD9E4665FA47030A" ID-GBIF-Occurrence="1658411508" country="India" location="Humeral area" pageId="14" pageNumber="115" specimenCount="1">
<keyLead id="309995C2AE53B63EDD9E4665FA47030A" pageId="14" pageNumber="115">
14 
<location id="8EB7602CAE53B63EDDE84665FEB9034F" LSID="urn:lsid:plazi:treatment:03C187E1AE50B63EDD9E45DFFA470609:8EB7602CAE53B63EDDE84665FEB9034F" box="[225,354,298,319]" country="India" name="Humeral area" pageId="14" pageNumber="115">Humeral area</location>
of elytron usually (but not always) with yellow spot; inner margin of metatibia not denticulate; metafemoral posterior margin usually greatly dilated apically, less often dilation reduced; aedeagus as in 
<figureCitation id="13532A72AE53B63ED9394608FBA7032C" box="[1072,1148,327,348]" captionStart="FIGURE 5" captionStartId="12.[151,250,1891,1913]" captionTargetBox="[162,1421,892,1854]" captionTargetId="figure@12.[151,1436,877,1870]" captionTargetPageId="12" captionText="FIGURE 5. Diversity of aedeagal parameres in Parachorius. (A) Parachorius fungorum. (B) Parachorius hookeri. (C) Parachorius schuelkei. (D) Parachorius thomsoni. (E) Parachorius globosus. (F) Parachorius newthayerae. (G) Parachorius bolavensis. (H) Parachorius fukiensis. (I) Parachorius nudus. (J) Parachorius peninsularis. (K) Parachorius pseudojavanus. (L) Parachorius javanus. (M) Parachorius solodovnikovi. (N) Parachorius longipenis. (O) Parachorius humeralis. (P) Parachorius semsanganus. (Q) Parachorius gotoi." httpUri="https://zenodo.org/record/1001925/files/figure.png" pageId="14" pageNumber="115">Fig. 5N</figureCitation>
, length 3.8–5.0 mm (
<figureCitation id="13532A72AE53B63ED8414608FA56032C" box="[1352,1421,327,348]" captionStart="FIGURE 13" captionStartId="26.[151,250,1259,1281]" captionTargetBox="[151,1416,193,1211]" captionTargetId="figure@26.[151,1436,193,1237]" captionTargetPageId="26" captionText="FIGURE 13. Morphology of Parachorius humeralis. (A – B) aedeagus (Darjeeling). (C) aedeagus. (D) habitus (male, holotype). (E) habitus (male). (F – H) right metafemur (male). (I) protibia (female). (J) protibia (male)." httpUri="https://zenodo.org/record/1001943/files/figure.png" pageId="14" pageNumber="115">Fig. 13</figureCitation>
); northeastern 
<collectingCountry id="F37F7667AE53B63EDC54462AFE55030A" box="[349,398,357,378]" name="India" pageId="14" pageNumber="115">India</collectingCountry>
................................................................ 
<taxonomicName id="4C684D74AE53B63ED99C462AFA47030A" ID-CoL="75NM5" authority="(Arrow, 1907)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[1173,1436,357,378]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B91CEAE5AE53B63ED99C462AFAD6030A" box="[1173,1293,357,378]" italics="true" pageId="14" pageNumber="115">P. humeralis</emphasis>
(
<bibRefCitation id="EFF94B06AE53B63ED810462AFA4E030A" author="Arrow" box="[1305,1429,357,378]" pageId="14" pageNumber="115" refString="Arrow, G. J. (1907) Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 2). The Annals and Magazine of Natural History, including Zoology, Botany and Geology, 7 (19), 416 - 439." type="journal article" year="1907">Arrow, 1907</bibRefCitation>
)
</taxonomicName>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8BD736F7AE53B63EDD9E46CDFA4003C4" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E46CDFA4003C4" pageId="14" pageNumber="115">- Humeral area of elytron without yellow spot, elytra uniformly colored; inner margin of metatibia denticulate with relatively small teeth; metafemoral posterior margin either simple or dilated and produced in a slight angle..................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="309C2E52AE53B63EDD9E46F2FA4000D0" pageId="14" pageNumber="115">
<paragraph id="8BD736F7AE53B63EDD9E46F2FA40005A" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E46F2FA40005A" pageId="14" pageNumber="115">
15 Parameres longer than in next species (
<figureCitation id="13532A72AE53B63EDF6846F2FD0203A2" box="[609,729,445,466]" captionStart="FIGURE 15" captionStartId="29.[151,250,1247,1269]" captionTargetBox="[151,1436,267,1223]" captionTargetId="figure@29.[151,1436,267,1225]" captionTargetPageId="29" captionText="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri="https://zenodo.org/record/1001947/files/figure.png" pageId="14" pageNumber="115">Fig. 15A–D</figureCitation>
); protibia with apical tooth spatulate, bent downward and notably larger than second tooth; first tooth wide, nearly merging with second tooth; metafemoral posterior margin simple, not modified or dilated and produced in slight angle in basal half (
<figureCitation id="13532A72AE53B63EDFC946B8FCDC007C" box="[704,775,503,524]" captionStart="FIGURE 15" captionStartId="29.[151,250,1247,1269]" captionTargetBox="[151,1436,267,1223]" captionTargetId="figure@29.[151,1436,267,1225]" captionTargetPageId="29" captionText="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri="https://zenodo.org/record/1001947/files/figure.png" pageId="14" pageNumber="115">Fig. 15</figureCitation>
); length 3.8–5.0 mm; Indochina...................................................................................................... 
<taxonomicName id="4C684D74AE53B63ED95A455AFAC4005A" ID-CoL="6TNM7" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1107,1311,533,554]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="longipenis" status="sp. nov.">
<emphasis id="B91CEAE5AE53B63ED95A455AFB15005A" box="[1107,1230,533,554]" italics="true" pageId="14" pageNumber="115">P. longipenis</emphasis>
Tarasov
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE53B63ED82F455BFA40005A" box="[1318,1435,532,554]" pageId="14" pageNumber="115" rank="species">
<emphasis id="B91CEAE5AE53B63ED82F455BFA40005A" bold="true" box="[1318,1435,532,554]" pageId="14" pageNumber="115">new species</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE53B63EDD9E457DFA4000D0" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E457DFA4000D0" pageId="14" pageNumber="115">
- Parameres shorter than in previous species (
<figureCitation id="13532A72AE53B63EDF9B457DFCD20037" box="[658,777,562,583]" captionStart="FIGURE 23" captionStartId="40.[151,250,1887,1909]" captionTargetBox="[151,1423,840,1815]" captionTargetId="figure@40.[151,1436,840,1830]" captionTargetPageId="40" captionText="FIGURE 23. Morphology of Parachorius solodovnikovi. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Thailand). (C) aedeagus, lateral view (paratype, Vietnam). (D) aedeagus, lateral view (E) habitus (male, holotype). (F) left metafemur (male, paratype, Vietnam) (G) left metafemur (male, holotype). (H) left protibia (male, holotype, original image flipped)." httpUri="https://zenodo.org/record/1001963/files/figure.png" pageId="14" pageNumber="115">Fig. 23A–C</figureCitation>
); protibia with apical tooth approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; metafemoral posterior margin simple, not modified, sometimes slightly dilated but not produced in angle; length 3.9–5.0 mm (
<figureCitation id="13532A72AE53B63EDE7A4522FC6000F2" box="[883,955,621,642]" captionStart="FIGURE 23" captionStartId="40.[151,250,1887,1909]" captionTargetBox="[151,1423,840,1815]" captionTargetId="figure@40.[151,1436,840,1830]" captionTargetPageId="40" captionText="FIGURE 23. Morphology of Parachorius solodovnikovi. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Thailand). (C) aedeagus, lateral view (paratype, Vietnam). (D) aedeagus, lateral view (E) habitus (male, holotype). (F) left metafemur (male, paratype, Vietnam) (G) left metafemur (male, holotype). (H) left protibia (male, holotype, original image flipped)." httpUri="https://zenodo.org/record/1001963/files/figure.png" pageId="14" pageNumber="115">Fig. 23</figureCitation>
); Indochina..................................................................................................... 
<taxonomicName id="4C684D74AE53B63ED92445C5FAC400EF" ID-CoL="4CPV3" authority="Tarasov" authorityName="Tarasov" authorityYear="2017" box="[1069,1311,650,671]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="solodovnikovi" status="sp. nov.">
<emphasis id="B91CEAE5AE53B63ED92445C5FB1500EF" box="[1069,1230,650,671]" italics="true" pageId="14" pageNumber="115">P. solodovnikovi</emphasis>
Tarasov
</taxonomicName>
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<emphasis id="B91CEAE5AE53B63ED82C45C5FA4000D0" bold="true" box="[1317,1435,650,672]" pageId="14" pageNumber="115">new species</emphasis>
</taxonomicNameLabel>
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</paragraph>
</keyStep>
<keyStep id="309C2E52AE53B63EDD9E45E8FA400164" pageId="14" pageNumber="115">
<paragraph id="8BD736F7AE53B63EDD9E45E8FA4000AA" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E45E8FA4000AA" pageId="14" pageNumber="115">
16 Apical spur of protibia equal or longer than first protibial tooth and first three protarsomeres combined (
<figureCitation id="13532A72AE53B63ED9CC45E8FAD400CC" box="[1221,1295,679,700]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="14" pageNumber="115">Fig. 7E</figureCitation>
); inner margin of metatibia not denticulate (
<figureCitation id="13532A72AE53B63EDCFC458AFD9B00AA" box="[501,576,709,730]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="14" pageNumber="115">Fig. 7G</figureCitation>
)....................................................................17
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE53B63EDD9E45ADFA400164" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E45ADFA400164" pageId="14" pageNumber="115">
- Apical spur of protibia shorter than first protibial tooth and first three protarsomeres combined (
<figureCitation id="13532A72AE53B63ED99F45ADFB350087" box="[1174,1262,738,759]" captionStart="FIGURE 18" captionStartId="33.[151,250,1485,1507]" captionTargetBox="[151,1431,628,1457]" captionTargetId="figure@33.[151,1436,628,1464]" captionTargetPageId="33" captionText="FIGURE 18. Morphology of Parachorius nudus. (A, C) aedeagus, right lateral view (holotype). (B, D) aedeagus, left lateral view (holotype). (E) protibia (male, holotype). (F) habitus (male, holotype)." httpUri="https://zenodo.org/record/1001953/files/figure.png" pageId="14" pageNumber="115">Fig. 18E</figureCitation>
); inner margin of metatibiae with relatively small teeth (
<figureCitation id="13532A72AE53B63EDF5845B0FD190164" box="[593,706,767,788]" captionStart="FIGURE 19" captionStartId="35.[151,250,1502,1524]" captionTargetBox="[151,1436,473,1484]" captionTargetId="figure@35.[151,1436,473,1491]" captionTargetPageId="35" captionText="FIGURE 19. Morphology of Parachorius peninsularis. (A) aedeagus, lateral view (lectotype). (B) aedeagus, lateral view (Malay, Kedah). (C) aedeagus, lateral view (Sumatra). (D) aedeagus, lateral view (Thailand, Thung Yai). (E) habitus (male, lectotype). (F – H) right metafemur (male). (I) aedeagus, lateral view." httpUri="https://zenodo.org/record/1001955/files/figure.png" pageId="14" pageNumber="115">Fig. 19F–H</figureCitation>
).........................................................18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309C2E52AE53B63EDD9E4452FA4701D7" pageId="14" pageNumber="115">
<paragraph id="8BD736F7AE53B63EDD9E4452FA400120" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E4452FA400120" pageId="14" pageNumber="115">
17 Parameres widened basally (lateral view), apex acute; metafemoral posterior margin dilated near middle; dilation widely rimmed (
<figureCitation id="13532A72AE53B63EDC314475FEA8013F" box="[312,371,826,847]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="14" pageNumber="115">Fig. 7</figureCitation>
); length 
<quantity id="4C909B12AE53B63EDCC14475FDE5013F" box="[456,574,826,847]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="14" pageNumber="115" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
; southern Laos............................... 
<taxonomicName id="4C684D74AE53B63ED95B4475FAC4013F" ID-CoL="6TNM4" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1106,1311,826,847]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
<emphasis id="B91CEAE5AE53B63ED95B4475FB15013F" box="[1106,1230,826,847]" italics="true" pageId="14" pageNumber="115">P. bolavensis</emphasis>
Tarasov
</taxonomicName>
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<emphasis id="B91CEAE5AE53B63ED82F4475FA400120" bold="true" box="[1318,1435,826,848]" pageId="14" pageNumber="115">new species</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD736F7AE53B63EDD9E4418FA4701D7" blockId="14.[151,1436,151,1145]" pageId="14" pageNumber="115">
<keyLead id="309995C2AE53B63EDD9E4418FA4701D7" pageId="14" pageNumber="115">
- Parameres emarginate superiorly (
<emphasis id="B91CEAE5AE53B63EDF254418FD91011C" box="[556,586,855,876]" italics="true" pageId="14" pageNumber="115">i.e.</emphasis>
, dorsally in lateral view, 
<figureCitation id="13532A72AE53B63EDE494418FC72011C" box="[832,937,855,876]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="14" pageNumber="115">Fig. 8A–C</figureCitation>
), apex rounded; metafemoral posterior margin simple or dilated in apical third, but not rimmed (
<figureCitation id="13532A72AE53B63EDF9E443AFD0A01FA" box="[663,721,885,906]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="14" pageNumber="115">Fig. 8</figureCitation>
); length 
<quantity id="4C909B12AE53B63EDE2F443AFC4701FA" box="[806,924,885,906]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.6" metricValueMin="3.5" pageId="14" pageNumber="115" unit="mm" value="4.05" valueMax="4.6" valueMin="3.5">3.5–4.6 mm</quantity>
; southeastern 
<collectingCountry id="F37F7667AE53B63ED92D443AFBBB01FA" box="[1060,1120,885,906]" name="China" pageId="14" pageNumber="115">China</collectingCountry>
, 
<collectingCountry id="F37F7667AE53B63ED960443AFB4101FA" box="[1129,1178,885,906]" name="Laos" pageId="14" pageNumber="115">Laos</collectingCountry>
, 
<collectingCountry id="F37F7667AE53B63ED9AD443AFB2E01FA" box="[1188,1269,885,906]" name="Vietnam" pageId="14" pageNumber="115">Vietnam</collectingCountry>
........................................................................................... 
<taxonomicName id="4C684D74AE53B63ED98844DDFA4701D7" ID-CoL="75NLS" authority="(Balthasar, 1960)" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[1153,1436,914,935]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE53B63ED98844DDFB2B01D7" box="[1153,1264,914,935]" italics="true" pageId="14" pageNumber="115">P. fukiensis</emphasis>
(
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)
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18 Protibia with apical tooth spatulate, bent downward and notably larger than second tooth; first tooth wide, appearing bifurcate as it merges with denticles located between first and second teeth; metafemoral posterior margin simple, not dilated; aedeagus as in 
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; length 
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(
<figureCitation id="13532A72AE53B63EDF0B44A3FD930671" box="[514,584,1004,1025]" captionStart="FIGURE 18" captionStartId="33.[151,250,1485,1507]" captionTargetBox="[151,1431,628,1457]" captionTargetId="figure@33.[151,1436,628,1464]" captionTargetPageId="33" captionText="FIGURE 18. Morphology of Parachorius nudus. (A, C) aedeagus, right lateral view (holotype). (B, D) aedeagus, left lateral view (holotype). (E) protibia (male, holotype). (F) habitus (male, holotype)." httpUri="https://zenodo.org/record/1001953/files/figure.png" pageId="14" pageNumber="115">Fig. 18</figureCitation>
); Cambodia.......................................... 
<taxonomicName id="4C684D74AE53B63ED9C844A3FA470671" ID-CoL="6TNM8" authority="(Sharp, 1875)" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[1217,1436,1004,1025]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="nudus">
<emphasis id="B91CEAE5AE53B63ED9C844A3FAC90671" box="[1217,1298,1004,1025]" italics="true" pageId="14" pageNumber="115">P. nudus</emphasis>
(
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- Protibia with apical tooth approximately twice as wide as second tooth, first and second teeth distinctly separated from each other; metafemoral posterior margin simple, not modified or slightly dilated in apical forth and produced in rounded angle preapically; aedeagus as in 
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Fig. 
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<date id="FFD61037AE53B63EDF1D4308FDF1062C" box="[532,554,1095,1116]" pageId="14" pageNumber="115">5I</date>
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; length 4.0– 
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(
<figureCitation id="13532A72AE53B63EDFFE4308FCE5062C" box="[759,830,1095,1116]" captionStart="FIGURE 19" captionStartId="35.[151,250,1502,1524]" captionTargetBox="[151,1436,473,1484]" captionTargetId="figure@35.[151,1436,473,1491]" captionTargetPageId="35" captionText="FIGURE 19. Morphology of Parachorius peninsularis. (A) aedeagus, lateral view (lectotype). (B) aedeagus, lateral view (Malay, Kedah). (C) aedeagus, lateral view (Sumatra). (D) aedeagus, lateral view (Thailand, Thung Yai). (E) habitus (male, lectotype). (F – H) right metafemur (male). (I) aedeagus, lateral view." httpUri="https://zenodo.org/record/1001955/files/figure.png" pageId="14" pageNumber="115">Fig. 19</figureCitation>
); 
<collectingCountry id="F37F7667AE53B63EDE594308FC73062C" box="[848,936,1095,1116]" name="Thailand" pageId="14" pageNumber="115">Thailand</collectingCountry>
, 
<location id="8EB7602CAE53B63EDEB84308FB8E062C" LSID="urn:lsid:plazi:treatment:03C187E1AE50B63EDD9E45DFFA470609:8EB7602CAE53B63EDEB84308FB8E062C" box="[945,1109,1095,1116]" country="Thailand" name="Malay Peninsula" pageId="14" pageNumber="115" stateProvince="Sumatra">Malay Peninsula</location>
, 
<collectingRegion id="49ACF815AE53B63ED9574308FB75062C" box="[1118,1198,1095,1116]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="115">Sumatra</collectingRegion>
................................................................................................ 
<taxonomicName id="4C684D74AE53B63ED977432BFA470609" ID-CoL="75NLW" authority="(Arrow, 1907)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[1150,1436,1124,1145]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91CEAE5AE53B63ED977432BFAD70609" box="[1150,1292,1124,1145]" italics="true" pageId="14" pageNumber="115">P. peninsularis</emphasis>
(
<bibRefCitation id="EFF94B06AE53B63ED810432BFA4E0609" author="Arrow" box="[1305,1429,1124,1145]" pageId="14" pageNumber="115" refString="Arrow, G. J. (1907) Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 2). The Annals and Magazine of Natural History, including Zoology, Botany and Geology, 7 (19), 416 - 439." type="journal article" year="1907">Arrow, 1907</bibRefCitation>
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