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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FFB7FFAA66BAFA3C60BE72E5" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Harpagophora arida Redman 2003" docType="treatment" docVersion="2" lastPageNumber="289" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="287" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vohland, Katrin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamer, Michelle</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>251</mods:start>
<mods:end>251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5" httpUri="http://treatment.plazi.org/id/039587C7FFB7FFAA66BAFA3C60BE72E5" lastPageId="38" lastPageNumber="289" pageId="36" pageNumber="287">
<subSubSection box="[421,827,1471,1497]" pageId="36" pageNumber="287" type="nomenclature">
<paragraph blockId="36.[144,1104,1471,1677]" box="[421,827,1471,1497]" pageId="36" pageNumber="287">
<heading box="[421,827,1471,1497]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="287" reason="3">
<taxonomicName ID-CoL="3JSC6" authority="Redman, 2003" authorityName="Redman" authorityYear="2003" box="[421,827,1471,1497]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="arida">
<emphasis box="[421,657,1471,1497]" italics="true" pageId="36" pageNumber="287">Harpagophora arida</emphasis>
Redman, 2003
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[559,688,1511,1538]" pageId="36" pageNumber="287" type="description">
<paragraph blockId="36.[144,1104,1471,1677]" box="[559,688,1511,1538]" pageId="36" pageNumber="287">
<figureCitation box="[559,688,1511,1538]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="36" pageNumber="287">Figs 65–70</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[144,732,1549,1572]" pageId="36" pageNumber="287" type="reference_group">
<paragraph blockId="36.[144,1104,1471,1677]" box="[144,732,1549,1572]" pageId="36" pageNumber="287">
<taxonomicName authority=": Redman et al. 2003: 215 - 217" authorityName="Redman" authorityPageNumber="215 - 217" authorityYear="2003" box="[144,620,1549,1572]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="arida">
<emphasis box="[144,334,1550,1571]" italics="true" pageId="36" pageNumber="287">Harpagophora arida</emphasis>
: Redman
<emphasis box="[428,468,1550,1571]" italics="true" pageId="36" pageNumber="287">et al</emphasis>
. 2003: 215–217
</taxonomicName>
, figs 20–26.
</paragraph>
</subSubSection>
<subSubSection lastPageId="38" lastPageNumber="289" pageId="36" pageNumber="287" type="diagnosis">
<paragraph blockId="36.[144,1104,1471,1677]" lastBlockId="37.[181,1142,203,438]" lastPageId="37" lastPageNumber="288" pageId="36" pageNumber="287">
Diagnosis: Gonopod teloxocite apically broadly rounded, in oral view with small sclerotised, basally directed projection (
<figureCitation box="[535,615,1618,1644]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="36" pageNumber="287">Fig. 66</figureCitation>
,
<emphasis box="[625,633,1619,1644]" italics="true" pageId="36" pageNumber="287">t</emphasis>
). Gonopod telopodite with a larger straight femoral spine having a slightly furcated tip (
<figureCitation box="[638,720,1650,1677]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="36" pageNumber="287">Fig. 65</figureCitation>
,
<emphasis box="[732,768,1650,1676]" italics="true" pageId="36" pageNumber="287">fp1</emphasis>
), and a shorter strong spine at the telopodite knee (
<figureCitation box="[405,485,203,230]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="37" pageNumber="288">Fig. 65</figureCitation>
,
<emphasis box="[496,531,204,230]" italics="true" pageId="37" pageNumber="288">fp2</emphasis>
), as well as a smaller third spine at flexure of telopodite (
<figureCitation box="[190,270,235,262]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="37" pageNumber="288">Fig. 65</figureCitation>
,
<emphasis box="[281,306,236,261]" italics="true" pageId="37" pageNumber="288">as</emphasis>
). Pectinophore elongate, ribbon-like, curling distally, with proximal margin serrated (
<figureCitation box="[285,368,267,294]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="37" pageNumber="288">Fig. 65</figureCitation>
,
<emphasis box="[380,406,268,293]" italics="true" pageId="37" pageNumber="288">pe</emphasis>
). Spine branch short, narrow, and apically bifid (
<figureCitation box="[955,1037,267,294]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="37" pageNumber="288">Fig. 65</figureCitation>
,
<emphasis box="[1049,1074,268,294]" italics="true" pageId="37" pageNumber="288">sb</emphasis>
).
</paragraph>
<paragraph blockId="37.[181,1142,203,438]" box="[181,321,308,334]" pageId="37" pageNumber="288">Description:</paragraph>
<paragraph blockId="37.[181,1142,203,438]" pageId="37" pageNumber="288">
<emphasis box="[181,226,348,374]" italics="true" pageId="37" pageNumber="288">Size</emphasis>
: Relatively stout. In males, body width anteriorly 8.4 mm, posteriorly 8.5 mm, length about
<quantity box="[333,418,380,406]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="288" unit="mm" value="90.0">90 mm</quantity>
; in females
<quantity box="[560,661,380,406]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="37" pageNumber="288" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
,
<quantity box="[675,787,380,406]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="37" pageNumber="288" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
and
<quantity box="[846,987,380,406]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" metricValueMax="10.1" metricValueMin="9.3" pageId="37" pageNumber="288" unit="mm" value="97.0" valueMax="101.0" valueMin="93.0">93–101 mm</quantity>
, respectively. There are 56–58 body segments.
</paragraph>
<caption pageId="37" pageNumber="288" startId="37.[181,220,1534,1556]" targetBox="[193,1116,490,1513]" targetPageId="37" targetType="figure">
<paragraph blockId="37.[181,1143,1534,1676]" pageId="37" pageNumber="288">
Figs 65–70.
<taxonomicName authority="Redman" authorityName="Redman" box="[292,560,1534,1556]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="288" phylum="Arthropoda" rank="species" species="arida">
<emphasis box="[292,480,1534,1555]" italics="true" pageId="37" pageNumber="288">Harpagophora arida</emphasis>
Redman
</taxonomicName>
, ♂: (65) telopodite, oral view; (66) telocoxite, oral view; (67) gnathochilarium; (68) head and first segments, lateral view; (69) epiproct, lateral view; (70) leg 2 of male. Abbreviations: (as) additional small spine on telopodite, (c) cardo, (col) collum, (cx) coxa, (fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<emphasis box="[144,223,204,230]" italics="true" pageId="38" pageNumber="289">Colour</emphasis>
: In life, dark red-brown to black, appearing slightly striated due to lighter prozonites and venter. Frons and first antennomeres chestnut brown. Head, collum and segment 2 dark brown, further segments nearly black, anal segment dark brown. Legs and distal antennomeres dark brown. Sternites and anterior part of prozonites yellowish, ventrally and dorsally reaching suture between meta- and prozonite.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<emphasis box="[144,205,364,390]" italics="true" pageId="38" pageNumber="289">Head</emphasis>
: Frons hairless, slightly wrinkled. Epicranial groove shallow, ending at distinct interocular groove. There are 6 supralabral pits. Eyes with suture from hind edge. Antenna reaching 2
<superScript attach="left" box="[330,346,427,441]" fontSize="6" pageId="38" pageNumber="289">nd</superScript>
segment; antennomere 2&gt;3&gt;4&gt;5=6&gt;2&gt;7. Gnathochilarium having mentum (
<figureCitation box="[249,330,459,486]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 67</figureCitation>
,
<emphasis box="[340,360,460,485]" italics="true" pageId="38" pageNumber="289">m</emphasis>
) and prebasilare (
<figureCitation box="[558,638,459,486]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 67</figureCitation>
,
<emphasis box="[649,677,460,486]" italics="true" pageId="38" pageNumber="289">pb</emphasis>
) merged; stipes (
<figureCitation box="[866,946,459,486]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 67</figureCitation>
,
<emphasis box="[957,976,460,485]" italics="true" pageId="38" pageNumber="289">st</emphasis>
) connected with cardo (
<figureCitation box="[278,360,491,518]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 67</figureCitation>
,
<emphasis box="[372,384,492,517]" italics="true" pageId="38" pageNumber="289">c</emphasis>
); hypostoma with median suture (
<figureCitation box="[769,851,491,518]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 67</figureCitation>
,
<emphasis box="[863,877,492,518]" italics="true" pageId="38" pageNumber="289">h</emphasis>
).
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<emphasis box="[144,226,524,550]" italics="true" pageId="38" pageNumber="289">Collum</emphasis>
: Subquadratic, slightly protruded backwards, with a strong lateral ridge, channeled anteriorly (
<figureCitation box="[335,420,556,582]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 68</figureCitation>
).
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<emphasis box="[144,387,588,614]" italics="true" pageId="38" pageNumber="289">Tergites and sternites</emphasis>
: Ozopores from 6
<superScript attach="left" box="[598,611,587,601]" fontSize="6" pageId="38" pageNumber="289">th</superScript>
segment onwards. Prozonites and sternites with transverse striation, metazonites below ozopores with marked longitudinal stripes. Limbus finely dentate. Epiproct (
<figureCitation box="[521,605,651,678]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 69</figureCitation>
) with very small, slightly upwards­directed projection, hypoproct broadly rounded.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<emphasis box="[144,197,716,741]" italics="true" pageId="38" pageNumber="289">Legs</emphasis>
: Syncoxosternite of 1
<superScript attach="left" box="[443,454,716,730]" fontSize="6" pageId="38" pageNumber="289">st</superScript>
pair of legs with distinct suture. Pads present on tibia and postfemur, distal pad always larger than interior one, smaller on posterior legs, absent on legs of last 3 or 4 diplosegments.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" box="[144,452,812,838]" pageId="38" pageNumber="289">
<emphasis box="[144,259,812,838]" italics="true" pageId="38" pageNumber="289">Gonopods</emphasis>
: As in diagnosis.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
<materialsCitation collectingDate="2001-04-01" collectionCode="ZMHB" collectorName="M. Uhlig" country="Namibia" county="River Canyon" latitude="-27.682777" location="Karasburg district" longLatPrecision="20" longitude="17.813055" municipality="Fish River" pageId="38" pageNumber="289" specimenCode="MYR 13730" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<typeStatus box="[144,232,850,871]" pageId="38" pageNumber="289">Paratypes</typeStatus>
(examined):
<specimenCount box="[356,384,850,871]" pageId="38" pageNumber="289" type="male">1♂</specimenCount>
<specimenCount box="[392,421,850,871]" pageId="38" pageNumber="289" type="female">3♀</specimenCount>
<collectingCountry box="[428,531,850,871]" name="Namibia" pageId="38" pageNumber="289">NAMIBIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5:8EE3600AFFB5FFAA6501FCD162977503" box="[542,708,850,872]" country="Namibia" county="River Canyon" latitude="-27.682777" longLatPrecision="20" longitude="17.813055" municipality="Fish River" name="Karasburg district" pageId="38" pageNumber="289">Karasburg district</location>
,
<location LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5:8EE3600AFFB5FFAA65CFFCD1635F7503" box="[720,780,850,872]" country="Namibia" county="River Canyon" latitude="-27.682777" longLatPrecision="20" longitude="17.813055" municipality="Fish River" name="Karios" pageId="38" pageNumber="289">Karios</location>
8,
<location LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5:8EE3600AFFB5FFAA6433FCD1638D750C" box="[812,990,850,872]" country="Namibia" county="River Canyon" latitude="-27.682777" longLatPrecision="20" longitude="17.813055" municipality="Fish River" name="Fish River Canyon" pageId="38" pageNumber="289">
<collectingMunicipality box="[812,909,850,872]" pageId="38" pageNumber="289">
Fish
<collectingCounty box="[859,909,850,872]" pageId="38" pageNumber="289">River</collectingCounty>
</collectingMunicipality>
Canyon
</location>
,
<geoCoordinate box="[1001,1104,850,872]" degrees="27" direction="south" minutes="40" orientation="latitude" pageId="38" pageNumber="289" precision="15" seconds="58" value="-27.682777">27°40'58&quot;S</geoCoordinate>
<geoCoordinate box="[144,250,874,895]" degrees="17" direction="east" minutes="48" orientation="longitude" pageId="38" pageNumber="289" precision="15" seconds="47" value="17.813055">17°48'47&quot;E</geoCoordinate>
,
<date box="[259,345,874,896]" pageId="38" pageNumber="289" value="2001-04-01">
<collectingDate box="[259,345,874,896]" pageId="38" pageNumber="289" value="2001-04-01">1.iv.2001</collectingDate>
</date>
,
<collectorName box="[355,440,874,896]" pageId="38" pageNumber="289">M. Uhlig</collectorName>
, BIOTA 522, plot 19 (
<collectionCode box="[642,708,874,895]" pageId="38" pageNumber="289">ZMHB</collectionCode>
/
<specimenCode box="[714,833,874,895]" collectionCode="MYR" pageId="38" pageNumber="289">MYR 13730</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
Other material examined:
<materialsCitation collectingDate="1972-01-15" collectingDateMax="1972-01-18" collectingDateMin="1972-01-15" collectionCode="NMSA" country="Namibia" county="Luderitz" latitude="-26.7" location="Plateau'" longLatPrecision="1240" longitude="15.266666" municipality="Aar" pageId="38" pageNumber="289" specimenCode="NMSA 19707" specimenCount="1" specimenCount-male="1">
<collectingCountry box="[377,480,909,930]" name="Namibia" pageId="38" pageNumber="289">NAMIBIA</collectingCountry>
:
<specimenCount box="[487,515,909,930]" pageId="38" pageNumber="289" type="male">1♂</specimenCount>
‘
<location LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5:8EE3600AFFB5FFAA6514FC0E620875C8" box="[523,603,909,931]" country="Namibia" county="Luderitz" latitude="-26.7" longLatPrecision="1240" longitude="15.266666" municipality="Aar" name="Plateau'" pageId="38" pageNumber="289">Plateau’</location>
,
<collectingMunicipality box="[610,645,909,930]" pageId="38" pageNumber="289">Aar</collectingMunicipality>
,
<collectingCounty box="[653,730,909,931]" pageId="38" pageNumber="289">Lüderitz</collectingCounty>
[
<geoCoordinate box="[739,812,908,930]" degrees="26" direction="south" minutes="42" orientation="latitude" pageId="38" pageNumber="289" precision="925" value="-26.7">26°42'S</geoCoordinate>
<geoCoordinate box="[817,893,908,930]" degrees="15" direction="east" minutes="16" orientation="longitude" pageId="38" pageNumber="289" precision="925" value="15.266666">15°16'E</geoCoordinate>
],
<date box="[907,1027,908,930]" pageId="38" pageNumber="289" value="1972-01-15" valueMax="1972-01-18" valueMin="1972-01-15">
<collectingDate box="[907,1027,908,930]" pageId="38" pageNumber="289" value="1972-01-15" valueMax="1972-01-18" valueMin="1972-01-15">15–18.i.1972</collectingDate>
</date>
(
<specimenCode collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="38" pageNumber="289" type="Museum">NMSA 19707</specimenCode>
)
</materialsCitation>
;
<materialsCitation box="[220,992,932,955]" collectingDate="1972-01-14" collectingDateMax="1972-01-15" collectingDateMin="1972-01-14" collectionCode="NMNW" country="Namibia" county="River Canyon" latitude="-26.833334" location="Fish River Canyon" longLatPrecision="1239" longitude="17.283333" municipality="Fish River" pageId="38" pageNumber="289" specimenCode="SMN 22138" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount box="[220,248,933,954]" pageId="38" pageNumber="289" type="male">1♂</specimenCount>
<specimenCount box="[254,282,933,954]" pageId="38" pageNumber="289" type="female">1♀</specimenCount>
<location LSID="urn:lsid:plazi:treatment:039587C7FFB7FFAA66BAFA3C60BE72E5:8EE3600AFFB5FFAA663FFC26619F75D1" box="[288,460,933,955]" country="Namibia" county="River Canyon" latitude="-26.833334" longLatPrecision="1239" longitude="17.283333" municipality="Fish River" name="Fish River Canyon" pageId="38" pageNumber="289">
<collectingMunicipality box="[288,383,933,955]" pageId="38" pageNumber="289">
Fish
<collectingCounty box="[333,383,933,955]" pageId="38" pageNumber="289">River</collectingCounty>
</collectingMunicipality>
Canyon
</location>
[
<geoCoordinate box="[470,543,932,954]" degrees="26" direction="south" minutes="50" orientation="latitude" pageId="38" pageNumber="289" precision="925" value="-26.833334">26°50'S</geoCoordinate>
<geoCoordinate box="[549,625,933,954]" degrees="17" direction="east" minutes="17" orientation="longitude" pageId="38" pageNumber="289" precision="925" value="17.283333">17°17'E</geoCoordinate>
],
<date box="[640,761,932,954]" pageId="38" pageNumber="289" value="1972-01-14" valueMax="1972-01-15" valueMin="1972-01-14">
<collectingDate box="[640,761,932,954]" pageId="38" pageNumber="289" value="1972-01-14" valueMax="1972-01-15" valueMin="1972-01-14">14–15.i.1972</collectingDate>
</date>
(
<specimenCode box="[774,888,933,954]" collectionCode="SMN" country="China" lsid="urn:lsid:biocol.org:col:14058" name="Simao District National Medical and Pharmaceutical Institute" pageId="38" pageNumber="289" type="Herbarium">SMN 22138</specimenCode>
) (
<collectionCode box="[906,986,933,954]" country="Namibia" httpUri="http://grbio.org/cool/y6s1-e7mm" name="National Museum of Namibia" pageId="38" pageNumber="289">NMNW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="38.[144,1104,204,1092]" pageId="38" pageNumber="289">
Distribution: Distributed in southern
<collectingCountry box="[540,637,970,996]" name="Namibia" pageId="38" pageNumber="289">Namibia</collectingCountry>
, in the Fish River Canyon area (
<collectingRegion box="[976,1045,970,996]" country="Namibia" name="//Karas" pageId="38" pageNumber="289">Karas</collectingRegion>
). The
<typeStatus box="[144,242,1002,1028]" pageId="38" pageNumber="289">holotype</typeStatus>
(Albany Museum, Grahamstown,
<collectingCountry box="[630,775,1002,1028]" name="South Africa" pageId="38" pageNumber="289">South Africa</collectingCountry>
, uncatalogued) was collected from Ais-Ais [
<geoCoordinate box="[306,435,1034,1061]" degrees="27" direction="south" minutes="55" orientation="latitude" pageId="38" pageNumber="289" precision="15" seconds="20" value="-27.922222">27°55'20&quot;S</geoCoordinate>
<geoCoordinate box="[442,574,1034,1060]" degrees="17" direction="east" minutes="29" orientation="longitude" pageId="38" pageNumber="289" precision="15" seconds="23" value="17.489723">17°29'23&quot;E</geoCoordinate>
,
<quantity box="[586,659,1034,1061]" metricMagnitude="2" metricUnit="m" metricValue="2.42" pageId="38" pageNumber="289" unit="m" value="242.0">242 m</quantity>
] (Redman
<emphasis box="[784,833,1034,1060]" italics="true" pageId="38" pageNumber="289">et al</emphasis>
. 2003), which is part of the Fish River Canyon.
</paragraph>
<paragraph blockId="38.[144,1104,1108,1166]" pageId="38" pageNumber="289">Ecology: The animals were seen in large numbers after a rain event by M. Uhlig (pers. comm.).</paragraph>
</subSubSection>
</treatment>
</document>
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