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<document id="BC94A7CE97B966B43A2EFA5B6F1651DD" ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FFB5FFA466BAFB3761AE772D" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Zinophora : Chamberlin 1927" docType="treatment" docVersion="4" lastPageNumber="290" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="289" updateTime="1699501985278" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7F291E1001C58CE043BF4AF2B5468DF4">A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
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<subSubSection id="C326655AFFB5FFAA66BAFB37636872A5" box="[421,827,1204,1230]" pageId="38" pageNumber="289" type="nomenclature">
<paragraph id="8B8336D1FFB5FFAA66BAFB37636872A5" blockId="38.[421,827,1204,1230]" box="[421,827,1204,1230]" pageId="38" pageNumber="289">
<heading id="D0CB81BDFFB5FFAA66BAFB37636872A5" box="[421,827,1204,1230]" centered="true" fontSize="36" level="2" pageId="38" pageNumber="289" reason="6">
Genus
<taxonomicName id="4C3C4D52FFB5FFAA66ECFB37636872A5" ID-CoL="7Q6YR" authority="Chamberlin, 1927" authorityName=": Chamberlin" authorityYear="1927" box="[499,827,1204,1230]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB5FFAA66ECFB37623A72A5" box="[499,617,1204,1230]" italics="true" pageId="38" pageNumber="289">Zinophora</emphasis>
Chamberlin, 1927
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326655AFFB5FFAA678FFB5963B17353" pageId="38" pageNumber="289" type="reference_group">
<paragraph id="8B8336D1FFB5FFAA678FFB59613C736C" blockId="38.[144,1103,1242,1360]" pageId="38" pageNumber="289">
<taxonomicName id="4C3C4D52FFB5FFAA678FFB59619B7284" ID-CoL="7Q6YR" authority=": Chamberlin 1927: 211" authorityName=": Chamberlin" authorityPageNumber="211" authorityYear="1927" box="[144,456,1242,1264]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB5FFAA678FFB5960A37284" box="[144,240,1242,1263]" italics="true" pageId="38" pageNumber="289">Zinophora</emphasis>
: Chamberlin 1927: 211
</taxonomicName>
; Schubart 1966: 164; Demange 1983: 579; Hamer 1998: 55; Redman
<emphasis id="B948EAC3FFB5FFAA6320FB58615B736C" italics="true" pageId="38" pageNumber="289">et al.</emphasis>
2003: 230.
</paragraph>
<paragraph id="8B8336D1FFB5FFAA678FFA8961E9734B" blockId="38.[144,1103,1242,1360]" box="[144,442,1290,1312]" pageId="38" pageNumber="289">
<taxonomicName id="4C3C4D52FFB5FFAA678FFA8961E5734B" ID-CoL="7PVG4" authority=": Attems 1928: 376" authorityName="Attems" authorityPageNumber="376" authorityYear="1928" box="[144,438,1290,1312]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB5FFAA678FFA8961557374" box="[144,262,1290,1311]" italics="true" pageId="38" pageNumber="289">Poratophilus</emphasis>
: Attems 1928: 376
</taxonomicName>
.
</paragraph>
<paragraph id="8B8336D1FFB5FFAA678FFAA163B17353" blockId="38.[144,1103,1242,1360]" box="[144,994,1314,1336]" pageId="38" pageNumber="289">
<emphasis id="B948EAC3FFB5FFAA678FFAA16157735C" box="[144,260,1314,1335]" italics="true" pageId="38" pageNumber="289">Philoporatia</emphasis>
: Attems 1928: 376; Lawrence 1965: 50. Synonymised by Demange (1983: 579).
</paragraph>
</subSubSection>
<subSubSection id="C326655AFFB5FFA4678FFAB961AE772D" lastPageId="40" lastPageNumber="291" pageId="38" pageNumber="289" type="description">
<paragraph id="8B8336D1FFB5FFAA678FFAB96200733B" blockId="38.[144,1103,1242,1360]" box="[144,595,1338,1360]" pageId="38" pageNumber="289">
<taxonomicName id="4C3C4D52FFB5FFAA678FFAB961DC7324" box="[144,399,1338,1359]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="289" phylum="Arthropoda" rank="subGenus" subGenus="Philoporatia">
<emphasis id="B948EAC3FFB5FFAA678FFAB961DC7324" box="[144,399,1338,1359]" italics="true" pageId="38" pageNumber="289">Poratophilus (Philoporatia)</emphasis>
</taxonomicName>
: Schubart 1966: 156.
</paragraph>
<paragraph id="8B8336D1FFB5FFAA678FFAE363B37311" blockId="38.[144,1104,1376,1677]" box="[144,992,1376,1402]" pageId="38" pageNumber="289">
<typeStatus id="54878873FFB5FFAA678FFAE3609B7311" box="[144,200,1376,1402]" pageId="38" pageNumber="289">Type</typeStatus>
species:
<taxonomicName id="4C3C4D52FFB5FFAA6631FAE362987311" authority="Chamberlin, 1927" authorityName="Chamberlin" authorityYear="1927" box="[302,715,1376,1402]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="289" phylum="Arthropoda" rank="species" species="munda">
<emphasis id="B948EAC3FFB5FFAA6631FAE361A47311" box="[302,503,1376,1402]" italics="true" pageId="38" pageNumber="289">Zinophora munda</emphasis>
Chamberlin, 1927
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8B8336D1FFB5FFAA678FFA0A644B7028" blockId="38.[144,1104,1376,1677]" pageId="38" pageNumber="289">
Diagnosis: Telopodite with one or two simple femoral processes, where two spines present these of similar size. Apical elements comprising three components: the pectinophore, thumb and second lamella (
<figureCitation id="13072A54FFB5FFAA655EFA4A62C77388" box="[577,660,1481,1507]" captionStart="Figs 7174" captionStartId="39.[181,220,1558,1580]" captionTargetBox="[185,1105,472,1535]" captionTargetId="figure-111@39.[181,1141,465,1541]" captionTargetPageId="39" captionText="Figs 7174.Zinophora sabulosaAttems, ♂: (71) gonopod, oral view; (72) gonopod, aboral view; (73) head and first segments, lateral view; (74) epiproct, lateral view.Abbreviations:(col) collum,(fp) femoral process, (g) gonocoel, (ll) lateral lamella of gonocoxite,(lp) lateral process of telocoxite, (ml) median lamella of gonocoxite, (mp) median process of telocoxite, (pe) pectinophore, (s) solenomere, (sl) second lamella, (sp) spinous process, (t) thumb. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 71</figureCitation>
,
<emphasis id="B948EAC3FFB5FFAA65BEFA4962FC7388" box="[673,687,1482,1507]" italics="true" pageId="38" pageNumber="289">p</emphasis>
,
<emphasis id="B948EAC3FFB5FFAA65A1FA4962957388" box="[702,710,1482,1507]" italics="true" pageId="38" pageNumber="289">t</emphasis>
,
<emphasis id="B948EAC3FFB5FFAA65CAFA4A62BB7388" box="[725,744,1481,1507]" italics="true" pageId="38" pageNumber="289">sl</emphasis>
). Thumb narrow and falcate, or a saucer-shaped, laminate plate broadening apically, curving away from other apical elements. Distal ends of telocoxites complexly lobed (
<figureCitation id="13072A54FFB5FFAA65F0F98A63117048" box="[751,834,1545,1571]" captionStart="Figs 7174" captionStartId="39.[181,220,1558,1580]" captionTargetBox="[185,1105,472,1535]" captionTargetId="figure-111@39.[181,1141,465,1541]" captionTargetPageId="39" captionText="Figs 7174.Zinophora sabulosaAttems, ♂: (71) gonopod, oral view; (72) gonopod, aboral view; (73) head and first segments, lateral view; (74) epiproct, lateral view.Abbreviations:(col) collum,(fp) femoral process, (g) gonocoel, (ll) lateral lamella of gonocoxite,(lp) lateral process of telocoxite, (ml) median lamella of gonocoxite, (mp) median process of telocoxite, (pe) pectinophore, (s) solenomere, (sl) second lamella, (sp) spinous process, (t) thumb. Scale bar = 1 mm." pageId="38" pageNumber="289">Fig. 71</figureCitation>
). Prefemora of first legs not touching medially and syncoxosternum without suture (Redman
<emphasis id="B948EAC3FFB5FFAA648FF9A9639B7028" box="[912,968,1577,1603]" italics="true" pageId="38" pageNumber="289">et al.</emphasis>
2003).
</paragraph>
<paragraph id="8B8336D1FFB5FFAA678FF9D1645E70E7" blockId="38.[144,1104,1376,1677]" pageId="38" pageNumber="289">
Distribution: Most diverse in
<collectingCountry id="F32B7641FFB5FFAA66C6F9D162397007" box="[473,618,1618,1644]" name="South Africa" pageId="38" pageNumber="289">South Africa</collectingCountry>
, where the genus is widely distributed; also occurs in
<collectingCountry id="F32B7641FFB5FFAA67E3F9F161C070E7" box="[252,403,1650,1676]" name="Mozambique" pageId="38" pageNumber="289">Mozambique</collectingCountry>
,
<collectingCountry id="F32B7641FFB5FFAA6681F9F1624A70E7" box="[414,537,1650,1676]" name="Zimbabwe" pageId="38" pageNumber="289">Zimbabwe</collectingCountry>
,
<collectingCountry id="F32B7641FFB5FFAA653BF9F1622970E7" box="[548,634,1650,1676]" name="Zambia" pageId="38" pageNumber="289">Zambia</collectingCountry>
and
<collectingCountry id="F32B7641FFB5FFAA65AFF9F1634370E7" box="[688,784,1650,1676]" name="Namibia" pageId="38" pageNumber="289">Namibia</collectingCountry>
(Redman
<emphasis id="B948EAC3FFB5FFAA649AF9F063EE70E7" box="[901,957,1650,1676]" italics="true" pageId="38" pageNumber="289">et al.</emphasis>
2003).
</paragraph>
<paragraph id="8B8336D1FFB4FFA467AAFF4F61AE772D" blockId="39.[181,1142,204,422]" lastBlockId="40.[144,1104,204,326]" lastPageId="40" lastPageNumber="291" pageId="39" pageNumber="290">
Remarks: Attems (1928) assigned five new species to the genus
<taxonomicName id="4C3C4D52FFB4FFAB6466FF4F60A3776D" authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="39" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB4FFAB6466FF4F6458768D" box="[889,1035,204,230]" italics="true" pageId="39" pageNumber="290">Poratophilus</emphasis>
Silvestri, 1897
</taxonomicName>
, but stated that the illustrations of the
<typeStatus id="54878873FFB4FFAB65B7FF6F628B776D" box="[680,728,236,262]" pageId="39" pageNumber="290">type</typeStatus>
species,
<taxonomicName id="4C3C4D52FFB4FFAB645DFF6F6422776D" authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[834,1137,236,262]" class="Diplopoda" family="Spirostreptidae" genus="Kartinikus" kingdom="Animalia" order="Spirostreptida" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B948EAC3FFB4FFAB645DFF6F6392776D" box="[834,961,236,262]" italics="true" pageId="39" pageNumber="290">P. australis</emphasis>
Silvestri, 1897
</taxonomicName>
, were so poor that it was not possible to be certain about this designation, or of the three species described by
<bibRefCitation id="EFAD4B20FFB4FFAB66BBFEAF6275772D" author="CARL, J." box="[420,550,300,326]" pageId="39" pageNumber="290" pagination="383 - 409" refId="ref24085" refString="CARL, J. 1917. Spirostreptides nouveaux ou peu connus du Musee de Geneve. Revue Suisse Zoologie 25: 383 - 409." type="journal article" year="1917">Carl (1917)</bibRefCitation>
in
<taxonomicName id="4C3C4D52FFB4FFAB6555FEAF628F772D" authorityName="Silvestri" authorityYear="1897" box="[586,732,300,326]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="39" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB4FFAB6555FEAF628F772D" box="[586,732,300,326]" italics="true" pageId="39" pageNumber="290">Poratophilus</emphasis>
</taxonomicName>
. Attems (1928) suggested that if the five species described were found not to be congeneric with
<emphasis id="B948EAC3FFB4FFAB647AFECF639D770D" box="[869,974,332,358]" italics="true" pageId="39" pageNumber="290">
<taxonomicName id="4C3C4D52FFB4FFAB647AFECF6399770D" authorityName="Silvestri" authorityYear="1897" box="[869,970,332,358]" class="Diplopoda" family="Spirostreptidae" genus="Kartinikus" kingdom="Animalia" order="Spirostreptida" pageId="39" pageNumber="290" phylum="Arthropoda" rank="species" species="australis">australis</taxonomicName>
,
</emphasis>
then the name
<emphasis id="B948EAC3FFB4FFAB67AAFEEF611777ED" box="[181,324,364,390]" italics="true" pageId="39" pageNumber="290">Philoporatia</emphasis>
should be used. He did not provide any diagnosis for this genus. Schubart (1966) used the name
<emphasis id="B948EAC3FFB4FFAB66B9FE0F626777CD" box="[422,564,396,422]" italics="true" pageId="39" pageNumber="290">Philoporatia</emphasis>
as a subgenus of
<taxonomicName id="4C3C4D52FFB4FFAB65EFFE0F63D277CD" authorityName="Silvestri" authorityYear="1897" box="[752,897,396,422]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="39" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFB4FFAB65EFFE0F63D277CD" box="[752,897,396,422]" italics="true" pageId="39" pageNumber="290">Poratophilus</emphasis>
</taxonomicName>
, and Demange (1983) later synonymised
<emphasis id="B948EAC3FFBBFFA4666FFF4F6250768D" box="[368,515,204,230]" italics="true" pageId="40" pageNumber="291">Philoporatia</emphasis>
with
<taxonomicName id="4C3C4D52FFBBFFA46554FF4F6297768D" authorityName=": Chamberlin" authorityYear="1927" box="[587,708,204,230]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFBBFFA46554FF4F6297768D" box="[587,708,204,230]" italics="true" pageId="40" pageNumber="291">Zinophora</emphasis>
</taxonomicName>
, and placed Attemss (1928) and
<bibRefCitation id="EFAD4B20FFBBFFA4678FFF6F6176776D" author="CARL, J." box="[144,293,236,262]" pageId="40" pageNumber="291" pagination="383 - 409" refId="ref24085" refString="CARL, J. 1917. Spirostreptides nouveaux ou peu connus du Musee de Geneve. Revue Suisse Zoologie 25: 383 - 409." type="journal article" year="1917">Carls (1917)</bibRefCitation>
three species into the latter genus. Hoffman (1994) later verified
<taxonomicName id="4C3C4D52FFBBFFA46311FF6F60B9774D" authorityName="Silvestri" authorityYear="1897" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC3FFBBFFA46311FF6F60B9774D" italics="true" pageId="40" pageNumber="291">Poratophilus</emphasis>
</taxonomicName>
as a valid genus and described the South African
<taxonomicName id="4C3C4D52FFBBFFA46436FE8F641F774D" authority="Hoffman, 1994" authorityName="Hoffman" authorityYear="1994" box="[809,1100,268,294]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="gorteri">
<emphasis id="B948EAC3FFBBFFA46436FE8F63C7774E" box="[809,916,268,293]" italics="true" pageId="40" pageNumber="291">P. gorteri</emphasis>
Hoffman, 1994
</taxonomicName>
, as a second species in the genus.
</paragraph>
</subSubSection>
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