treatments-xml/data/03/95/87/039587ADBF3DFFA0FF7F439F20F2FC3C.xml
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<document id="11B26FD934719042E733E697CA0C971D" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin &amp; Jia, Xinyang" docDate="2023" docId="039587ADBF3DFFA0FF7F439F20F2FC3C" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Youtrechus zijunae Tian &amp; Huang &amp; Jia 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="6" updateTime="1699475196432" updateUser="ExternalLinkService">
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<mods:title id="AC0788EB8092469F3D4D7BCF2E831633">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="9367F8000BACCE3BD7ACF601032207E1">Tian, Mingyi</mods:namePart>
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<mods:namePart id="15086B7A38CCA14524B535F4D8C5F52D">Huang, Sunbin</mods:namePart>
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<mods:namePart id="29A2E41879DA608D4A1ECA4485E64ABA">Jia, Xinyang</mods:namePart>
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<paragraph id="8B8336BBBF3DFFACFF7F439F23A8FCE0" blockId="5.[151,456,762,825]" box="[151,456,762,789]" pageId="5" pageNumber="6">
<heading id="D0CB81D7BF3DFFACFF7F439F23A8FCE0" bold="true" box="[151,456,762,789]" fontSize="11" level="2" pageId="5" pageNumber="6" reason="3">
<emphasis id="B948EAA9BF3DFFACFF7F439F23A8FCE0" bold="true" box="[151,456,762,789]" pageId="5" pageNumber="6">
<taxonomicName id="4C3C4D38BF3DFFACFF7F439F231EFCE0" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[151,382,762,789]" class="Insecta" family="Carabidae" genus="Youtrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zijunae" status="sp. nov.">
<emphasis id="B948EAA9BF3DFFACFF7F439F231EFCE0" bold="true" box="[151,382,762,789]" italics="true" pageId="5" pageNumber="6">Youtrechus zijunae</emphasis>
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<taxonomicNameLabel id="A27B57D2BF3DFFACFE6D439E23A8FCE0" box="[389,456,763,789]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3266530BF3DFFACFF7F427A2326FCCC" box="[151,326,799,825]" pageId="5" pageNumber="6" type="description">
<paragraph id="8B8336BBBF3DFFACFF7F427A2326FCCC" blockId="5.[151,456,762,825]" box="[151,326,799,825]" pageId="5" pageNumber="6">
<figureCitation id="13072A3EBF3DFFACFF7F427A236FFCCC" box="[151,271,799,825]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="5" pageNumber="6">Figures 1n</figureCitation>
,
<figureCitation id="13072A3EBF3DFFACFEF4427A2326FCCC" box="[284,326,799,825]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="6.[151,250,1943,1967]" captionStartId-1="7.[151,250,1943,1967]" captionStartId-2="8.[151,250,1943,1967]" captionTargetBox-0="[263,1317,195,1905]" captionTargetBox-1="[165,1412,182,1920]" captionTargetBox-2="[162,1426,182,1920]" captionTargetId-0="figure-28@6.[258,1330,181,1920]" captionTargetId-1="figure-20@7.[162,1425,181,1920]" captionTargetId-2="figure-28@8.[162,1426,181,1920]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 2. Habitus of Youtrechus zijunae n. gen., n. sp., male, holotype." captionText-1="FIGURE 3. Youtrechus zijunae n. gen., n. sp.A. head (ventral); B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." captionText-2="FIGURE 4. Feixian Dong, the type locality of Youtrechus zijunae n. gen., n. sp. A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera Eutrichodesmus and Skleroprotopus; F. a running individual of Youtrechus zijunae n. gen., n. sp. inside the cave." figureDoi-0="http://doi.org/10.5281/zenodo.7642404" figureDoi-1="http://doi.org/10.5281/zenodo.7642406" figureDoi-2="http://doi.org/10.5281/zenodo.7642408" httpUri-0="https://zenodo.org/record/7642404/files/figure.png" httpUri-1="https://zenodo.org/record/7642406/files/figure.png" httpUri-2="https://zenodo.org/record/7642408/files/figure.png" pageId="5" pageNumber="6">24</figureCitation>
</paragraph>
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<subSubSection id="C3266530BF3DFFACFF7F42032001FC3C" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8B8336BBBF3DFFACFF7F42032001FC3C" blockId="5.[151,1437,870,2013]" pageId="5" pageNumber="6">
<emphasis id="B948EAA9BF3DFFACFF7F42032366FC75" bold="true" box="[151,262,870,896]" pageId="5" pageNumber="6">Material.</emphasis>
<materialsCitation id="3B543CE6BF3DFFACFEE742022742FC50" ID-GBIF-Occurrence="4030741303" collectingDate="2019-07-13" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Mengzhen Chen &amp; Zijun Ma &amp; Jingli Cheng" county="Feixian Dong" elevation="160" latitude="29.73" location="Xianning" longLatPrecision="735" longitude="114.24" municipality="Xian'an District" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<typeStatus id="54878819BF3DFFACFEE742022317FC74" box="[271,375,871,897]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3AFD32BF3DFFACFE68420223D9FC74" box="[384,441,871,897]" pageId="5" pageNumber="6" type="male">male</specimenCount>
,
<geoCoordinate id="EE08507CBF3DFFACFE2E4202204DFC74" box="[454,557,871,897]" degrees="29.73" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="555" value="29.73">29.73 °N</geoCoordinate>
/
<geoCoordinate id="EE08507CBF3DFFACFDAF420220DAFC74" box="[583,698,871,897]" degrees="114.24" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="555" value="114.24">114.24 °E</geoCoordinate>
,
<quantity id="4CC49B5EBF3DFFACFD2042022173FC74" box="[712,787,871,897]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="6" unit="m" value="160.0">
<elevation id="0011D188BF3DFFACFD2042022173FC74" box="[712,787,871,897]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="6" unit="m" value="160.0">160 m</elevation>
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, a limestone cave called
<collectingCounty id="62E24E37BF3DFFACFBDB420226B1FC74" box="[1075,1233,871,897]" pageId="5" pageNumber="6">Feixian Dong</collectingCounty>
,
<collectingMunicipality id="6BE7ACC1BF3DFFACFB36420227F8FC74" box="[1246,1432,871,897]" pageId="5" pageNumber="6">Xianan District</collectingMunicipality>
,
<location id="8EE36060BF3DFFACFF7F42EE2362FC50" LSID="urn:lsid:plazi:treatment:039587ADBF3DFFA0FF7F439F20F2FC3C:8EE36060BF3DFFACFF7F42EE2362FC50" box="[151,258,907,933]" county="Feixian Dong" latitude="29.73" longLatPrecision="735" longitude="114.24" municipality="Xian'an District" name="Xianning" pageId="5" pageNumber="6" stateProvince="Hubei">Xianning</location>
,
<collectingRegion id="49F8F859BF3DFFACFEE542EE2334FC50" box="[269,340,907,933]" country="China" name="Hubei" pageId="5" pageNumber="6">Hubei</collectingRegion>
,
<date id="FF82107BBF3DFFACFEB742EE2396FC50" box="[351,502,907,933]" pageId="5" pageNumber="6" value="2019-07-13">
<collectingDate id="EFC6E993BF3DFFACFEB742EE2396FC50" box="[351,502,907,933]" pageId="5" pageNumber="6" value="2019-07-13">2019. VII. 13</collectingDate>
</date>
,
<collectorName id="26C9536DBF3DFFACFDE942EE20EFFC50" box="[513,655,907,933]" pageId="5" pageNumber="6">Mingyi Tian</collectorName>
,
<collectorName id="26C9536DBF3DFFACFD7142EE2134FC50" box="[665,852,907,933]" pageId="5" pageNumber="6">Mengzhen Chen</collectorName>
,
<collectorName id="26C9536DBF3DFFACFCB642EE21A6FC50" box="[862,966,907,933]" pageId="5" pageNumber="6">Zijun Ma</collectorName>
&amp;
<collectorName id="26C9536DBF3DFFACFC0142EE2617FC50" box="[1001,1143,907,933]" pageId="5" pageNumber="6">Jingli Cheng</collectorName>
leg., in
<collectionCode id="ED2DAE7EBF3DFFACFB3842EE277EFC50" box="[1232,1310,907,933]" collectionName="SCAU" pageId="5" pageNumber="6">SCAU</collectionCode>
;
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<materialsCitation id="3B543CE6BF3DFFACFAC142EE2001FC3C" ID-GBIF-Occurrence="4030741322" collectingDate="2019-07-13" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Mengzhen Chen &amp; Zijun Ma &amp; Jingli Cheng" county="Feixian Dong" elevation="160" latitude="29.73" location="Xianning" longLatPrecision="735" longitude="114.24" municipality="Xian'an District" pageId="5" pageNumber="6" specimenCount="13" specimenCount-female="3" specimenCount-male="10" stateProvince="Hubei" typeStatus="paratype">
<typeStatus id="54878819BF3DFFACFAC142EE27F9FC50" box="[1321,1433,907,933]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
:
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and
<specimenCount id="9D3AFD32BF3DFFACFEDA42CA23C1FC3C" box="[306,417,943,969]" pageId="5" pageNumber="6" type="female">3 females</specimenCount>
, idem, in
<collectionCode id="ED2DAE7EBF3DFFACFDE742CA203DFC3C" box="[527,605,943,969]" collectionName="SCAU" pageId="5" pageNumber="6">SCAU</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3266530BF3DFFACFF2F42B72291FBC0" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B8336BBBF3DFFACFF2F42B72291FBC0" blockId="5.[151,1437,870,2013]" pageId="5" pageNumber="6">
<emphasis id="B948EAA9BF3DFFACFF2F42B72322FC19" bold="true" box="[199,322,978,1004]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Semi-aphenopsian and medium to large sized for cave trechines, with moderately elongated body and appendages, eyeless and depigmented; fore part (head plus prothorax) evidently shorter than elytra, surface smooth.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF3DFFA0FF2F455B262EFD5C" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6" type="description">
<paragraph id="8B8336BBBF3DFFACFF2F455B264DFBAC" blockId="5.[151,1437,870,2013]" box="[199,1069,1086,1113]" pageId="5" pageNumber="6">
<emphasis id="B948EAA9BF3DFFACFF2F455B2339FBAD" bold="true" box="[199,345,1086,1112]" pageId="5" pageNumber="6">Description.</emphasis>
Length:
<quantity id="4CC49B5EBF3DFFACFE56455B2028FBAC" box="[446,584,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="6.9" metricValueMin="6.5" pageId="5" pageNumber="6" unit="mm" value="6.7" valueMax="6.9" valueMin="6.5">6.56.9 mm</quantity>
; width:
<quantity id="4CC49B5EBF3DFFACFD4A455A2144FBAC" box="[674,804,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="5" pageNumber="6" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1mm</quantity>
. Habitus as in
<figureCitation id="13072A3EBF3DFFACFC20455A264AFBAC" box="[968,1066,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1317,195,1905]" captionTargetId="figure-28@6.[258,1330,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Habitus of Youtrechus zijunae n. gen., n. sp., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7642404" httpUri="https://zenodo.org/record/7642404/files/figure.png" pageId="5" pageNumber="6">Figure 2</figureCitation>
.
</paragraph>
<paragraph id="8B8336BBBF3DFFACFF2F450626F0FB30" blockId="5.[151,1437,870,2013]" pageId="5" pageNumber="6">Body yellowish brown, but antennae, mouthparts palps and tarsi pale; moderately shiny and whole body glabrous though genae with a few setae; microsculptural engraved meshes isodiametric on head, transverse striate on pronotum and elytra; fore body much shorter than elytra, (HLm+PL) / EL = 0. 800.82.</paragraph>
<paragraph id="8B8336BBBF3DFFACFF2F45AA2719F980" blockId="5.[151,1437,870,2013]" pageId="5" pageNumber="6">
Head stout, slightly longer than wide, HLm/HW = 1.581.64, HLl/HW = 1.161.22; genae fairly developed, slightly expanded, widest at about 1/3 of head from neck to apical margin of labrum; frons rather flat, vertex convex, frontal furrows widely and strongly divergent posteriad; clypeus transverse, quadrisetose; labrum transverse, frontal margin slightly emarginate, 6-setose; two pairs of supraorbital setiferous pores present at about middle and 1/4 of head from labrum; labial suture clearly visible, mentum bisetose on either side of tooth, mental tooth short and blunt at tip, basal fovea widely concave; submentum 6-setose; ligula thick and short, bisetose at apex; palps moderately elongated, 2
<superScript id="7C499BF3BF3DFFACFEF744C3234FFA41" attach="left" box="[287,303,1446,1460]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
labial palpomere 1.3 times as long as 3
<superScript id="7C499BF3BF3DFFACFD0D44C32093FA41" attach="none" box="[741,755,1446,1460]" fontSize="6" pageId="5" pageNumber="6">rd</superScript>
, the former bisetose on inner margin, with a seta medially on outer margin and another on out subapex, the latter glabrous; 3
<superScript id="7C499BF3BF3DFFACFCB144AF2107FA2D" attach="left" box="[857,871,1482,1496]" fontSize="6" pageId="5" pageNumber="6">rd</superScript>
maxillary palpomere 1.1 times as long as 4
<superScript id="7C499BF3BF3DFFACFAB844AF273DFA2D" attach="none" box="[1360,1373,1482,1496]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
, both glabrous; suborbital pore just between base and submentum (
<figureCitation id="13072A3EBF3DFFACFCBD448A21AFF9FC" box="[853,975,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[165,1412,182,1920]" captionTargetId="figure-20@7.[162,1425,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Youtrechus zijunae n. gen., n. sp.A. head (ventral); B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642406" httpUri="https://zenodo.org/record/7642406/files/figure.png" pageId="5" pageNumber="6">Figure 3A</figureCitation>
). Antennae thin and rather long, but not extended beyond elytral apices, longer in male; the 1
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antennomere stouter, as long as the 2
<superScript id="7C499BF3BF3DFFACFB6A477726F2F9D5" attach="none" box="[1154,1170,1554,1568]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
, the 3
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5
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longest, the 11
<superScript id="7C499BF3BF3DFFACFF5A475322DFF9B1" attach="left" box="[178,191,1590,1604]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
slightly longer than 10
<superScript id="7C499BF3BF3DFFACFE55475323AAF9B1" attach="none" box="[445,458,1590,1604]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
, the comparative length of each antennomere from the 1
<superScript id="7C499BF3BF3DFFACFBD147522624F9B0" attach="left" box="[1081,1092,1591,1605]" fontSize="6" pageId="5" pageNumber="6">st</superScript>
to 11
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in
<typeStatus id="54878819BF3DFFACFB4647522770F9A4" box="[1198,1296,1591,1617]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
as: 1
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(1.00) / 2
<superScript id="7C499BF3BF3DFFACFF5C473F22A4F99D" attach="left" box="[180,196,1626,1640]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
(1.17) / 3
<superScript id="7C499BF3BF3DFFACFEDB473F2321F99D" attach="left" box="[307,321,1626,1640]" fontSize="6" pageId="5" pageNumber="6">rd</superScript>
(1.83) / 4
<superScript id="7C499BF3BF3DFFACFE44473F23D9F99D" attach="left" box="[428,441,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.78) / 5
<superScript id="7C499BF3BF3DFFACFDCF473F2054F99D" attach="left" box="[551,564,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.89) / 6
<superScript id="7C499BF3BF3DFFACFD4A473F20CFF99D" attach="left" box="[674,687,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.67) / 7
<superScript id="7C499BF3BF3DFFACFCF5473F214AF99D" attach="left" box="[797,810,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.61) / 8
<superScript id="7C499BF3BF3DFFACFC70473F21C5F99D" attach="left" box="[920,933,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.44) / 9
<superScript id="7C499BF3BF3DFFACFBF9473F267EF99D" attach="left" box="[1041,1054,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.39) / 10
<superScript id="7C499BF3BF3DFFACFB72473F26C7F99D" attach="left" box="[1178,1191,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
(1.28) /11
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(1.56).
</paragraph>
<paragraph id="8B8336BBBF3DFFACFF2F471A277DF8B8" blockId="5.[151,1437,870,2013]" pageId="5" pageNumber="6">Pronotum quadrate, slightly longer than wide, PL/PW =1.071.11; wider than head, PW/HW = 1.211.23, slightly longer than head excluding mandibles, PL/HLl = 1.031.05; widest a little before middle, gently narrowed anteriorly and posteriorly, latero-margins finely bordered; front slightly concave, the base is prominent, with the presence of secondary angles, strongly sinuate on sides, making a sharp, though rectangular, hind angle on each side; anterior latero-marginal setae at about apical 1/8, posterior ones just on hind angles; disc moderately convex; median line well-defined, basal transversal impression and foveae well-marked. Scutellum fairly small.</paragraph>
<paragraph id="8B8336BBBF3DFFA0FF2F463227F2FEB4" blockId="5.[151,1437,870,2013]" lastBlockId="9.[151,1437,150,969]" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
Elytra elongated ovate, much longer than wide,EL/EW=1.641.66; much wider than pronotum,EW/PnW=1.75 1.76; widest at about 3/7 of elytra from base, lateral sides smooth, finely bordered throughout, marginal gutters wellmarked; slightly depressed medially near base, disc moderately convex; punctures of striae invisible, intervals more or less subequal in width except 1
<superScript id="7C499BF3BF3DFFACFDE246A62075F824" attach="left" box="[522,533,1987,2001]" fontSize="6" pageId="5" pageNumber="6">st</superScript>
one. Chaetotaxy (
<figureCitation id="13072A3EBF3DFFACFD3446A62131F828" box="[732,849,1987,2013]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[165,1412,182,1920]" captionTargetId="figure-20@7.[162,1425,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Youtrechus zijunae n. gen., n. sp.A. head (ventral); B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642406" httpUri="https://zenodo.org/record/7642406/files/figure.png" pageId="5" pageNumber="6">Figure 3B</figureCitation>
): anterior and medial dorsal pores located on 3
<superScript id="7C499BF3BF3DFFACFABA46A72700F825" attach="left" box="[1362,1376,1986,2000]" fontSize="6" pageId="5" pageNumber="6">rd</superScript>
stria, at about 2/13 and 4/13 of elytra from base respectively; posterior dorsal pore located on 5
<superScript id="7C499BF3BF31FFA0FB9541F326EAFF51" attach="left" box="[1149,1162,150,164]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
interval at about 6/13 of elytra from base; the preapical pore located at the apical anastomosis of 2
<superScript id="7C499BF3BF31FFA0FC2D41DF21B5FF3D" attach="left" box="[965,981,186,200]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
and 3
<superScript id="7C499BF3BF31FFA0FBF041DF2646FF3D" attach="left" box="[1048,1062,186,200]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
striae, and at about apical 2/13 of elytra, much closer to elytral suture than to apical margin; basal pore present; 1
<superScript id="7C499BF3BF31FFA0FC1741BA266AFF18" attach="left" box="[1023,1034,223,237]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
and 2
<superScript id="7C499BF3BF31FFA0FBA441BB263CFF19" attach="left" box="[1100,1116,222,236]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
marginal umbilicate close to marginal gutter; 4
<superScript id="7C499BF3BF31FFA0FEB54067230AFEE5" attach="left" box="[349,362,258,272]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
pore distant from 3
<superScript id="7C499BF3BF31FFA0FDA94067202FFEE5" attach="none" box="[577,591,258,272]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
; 5
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and 6
<superScript id="7C499BF3BF31FFA0FD51406720A6FEE5" attach="left" box="[697,710,258,272]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
pores located at about middle of elytra, close to each other; 7
<superScript id="7C499BF3BF31FFA0FA8040672715FEE5" attach="none" box="[1384,1397,258,272]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
, 8
<superScript id="7C499BF3BF31FFA0FA78406727FDFEE5" attach="none" box="[1424,1437,258,272]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
and apical pores distant from marginal gutter; the anguloapical one present, subequal to elytral margin and suture.
</paragraph>
<caption id="DF436633BF3EFFAFFF7F46F221E6F845" ID-DOI="http://doi.org/10.5281/zenodo.7642404" ID-Zenodo-Dep="7642404" box="[151,902,1943,1968]" httpUri="https://zenodo.org/record/7642404/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1943,1967]" targetBox="[263,1317,195,1905]" targetPageId="6">
<paragraph id="8B8336BBBF3EFFAFFF7F46F221E6F845" blockId="6.[151,902,1943,1968]" box="[151,902,1943,1968]" pageId="6" pageNumber="7">
<emphasis id="B948EAA9BF3EFFAFFF7F46F22373F85A" bold="true" box="[151,275,1943,1967]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
Habitus of
<taxonomicName id="4C3C4D38BF3EFFAFFE6246FD2028F85A" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[394,584,1944,1968]" class="Insecta" family="Carabidae" genus="Youtrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zijunae" status="n. gen., n. sp.">
<emphasis id="B948EAA9BF3EFFAFFE6246FD2028F85A" box="[394,584,1944,1968]" italics="true" pageId="6" pageNumber="7">Youtrechus zijunae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF3EFFAFFDA746FD20BBF845" bold="true" box="[591,731,1944,1968]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A27B57D2BF3EFFAFFDA746FD20BBF845" box="[591,731,1944,1968]" pageId="6" pageNumber="7" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</emphasis>
, male, holotype.
</paragraph>
</caption>
<caption id="DF436633BF3FFFAEFF7F46F22009F821" ID-DOI="http://doi.org/10.5281/zenodo.7642406" ID-Zenodo-Dep="7642406" httpUri="https://zenodo.org/record/7642406/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1943,1967]" targetBox="[165,1412,182,1920]" targetPageId="7">
<paragraph id="8B8336BBBF3FFFAEFF7F46F22009F821" blockId="7.[151,1437,1943,2004]" pageId="7" pageNumber="8">
<emphasis id="B948EAA9BF3FFFAEFF7F46F22372F85A" bold="true" box="[151,274,1943,1967]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
<taxonomicName id="4C3C4D38BF3FFFAEFEFF46FD23B4F85A" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[279,468,1944,1968]" class="Insecta" family="Carabidae" genus="Youtrechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="zijunae" status="n. gen., n. sp.">
<emphasis id="B948EAA9BF3FFFAEFEFF46FD23B4F85A" box="[279,468,1944,1968]" italics="true" pageId="7" pageNumber="8">Youtrechus zijunae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF3FFFAEFE3246FD2004F845" bold="true" box="[474,612,1944,1968]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A27B57D2BF3FFFAEFE3246FD2004F845" box="[474,612,1944,1968]" pageId="7" pageNumber="8" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</emphasis>
A. head (ventral); B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view.
</paragraph>
</caption>
<caption id="DF436633BF30FFA1FF7F46F22175F802" ID-DOI="http://doi.org/10.5281/zenodo.7642408" ID-Zenodo-Dep="7642408" httpUri="https://zenodo.org/record/7642408/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1943,1967]" targetBox="[162,1426,182,1920]" targetPageId="8">
<paragraph id="8B8336BBBF30FFA1FF7F46F22175F802" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="9">
<emphasis id="B948EAA9BF30FFA1FF7F46F22375F85A" bold="true" box="[151,277,1943,1967]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Feixian Dong, the type locality of
<taxonomicName id="4C3C4D38BF30FFA1FD6246FD212BF85A" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[650,843,1944,1968]" class="Insecta" family="Carabidae" genus="Youtrechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zijunae" status="n. gen., n. sp.">
<emphasis id="B948EAA9BF30FFA1FD6246FD212BF85A" box="[650,843,1944,1968]" italics="true" pageId="8" pageNumber="9">Youtrechus zijunae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF30FFA1FCBC46FD2188F845" bold="true" box="[852,1000,1944,1968]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A27B57D2BF30FFA1FCBC46FD2188F845" box="[852,1000,1944,1968]" pageId="8" pageNumber="9" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</emphasis>
A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera
<taxonomicName id="4C3C4D38BF30FFA1FC7C46D92657F821" authorityName="Silvestri" authorityYear="1910" box="[916,1079,1980,2004]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAA9BF30FFA1FC7C46D92657F821" box="[916,1079,1980,2004]" italics="true" pageId="8" pageNumber="9">Eutrichodesmus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4D38BF30FFA1FB8546D92766F821" authorityName="Attems" authorityYear="1901" box="[1133,1286,1980,2004]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAA9BF30FFA1FB8546D92766F821" box="[1133,1286,1980,2004]" italics="true" pageId="8" pageNumber="9">Skleroprotopus</emphasis>
</taxonomicName>
; F. a running individual of
<taxonomicName id="4C3C4D38BF30FFA1FEC8468523BEF802" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[288,478,2016,2040]" class="Insecta" family="Carabidae" genus="Youtrechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zijunae" status="n. gen., n. sp.">
<emphasis id="B948EAA9BF30FFA1FEC8468523BEF802" box="[288,478,2016,2040]" italics="true" pageId="8" pageNumber="9">Youtrechus zijunae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF30FFA1FE0C46852011F80D" bold="true" box="[484,625,2016,2040]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A27B57D2BF30FFA1FE0C46852011F80D" box="[484,625,2016,2040]" pageId="8" pageNumber="9" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</emphasis>
inside the cave.
</paragraph>
</caption>
<paragraph id="8B8336BBBF31FFA0FF2F402E26D1FE58" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="10">
Legs thin and slender, bearing short pubescence; the 1
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and 2
<superScript id="7C499BF3BF31FFA0FC74402F21CCFEAD" attach="left" box="[924,940,330,344]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
protarsomeres widened in male, distinctly denticulate inwardly; the 1
<superScript id="7C499BF3BF31FFA0FE2C400A23AFFE88" attach="left" box="[452,463,367,381]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
protarsomere distinctly widened in female; the 1
<superScript id="7C499BF3BF31FFA0FC12400A2665FE88" attach="left" box="[1018,1029,367,381]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
tarsomere much shorter than 2
<superScript id="7C499BF3BF31FFA0FA8B400B2713FE89" attach="none" box="[1379,1395,366,380]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
4
<superScript id="7C499BF3BF31FFA0FA78400B27FDFE89" attach="right" box="[1424,1437,366,380]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
tarsomeres together in fore leg, whereas as long as those in middle and hind legs respectively.
</paragraph>
<paragraph id="8B8336BBBF31FFA0FF2F40D22179FE00" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="10">Ventrites smooth, IVVI each with two pairs of paramedial setae, together with several pubescence in middle portion, VII bisetose apically in male, 6-setose in female.</paragraph>
<paragraph id="8B8336BBBF31FFA0FF2F409A262EFD5C" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="10">
Male genitalia (
<figureCitation id="13072A3EBF31FFA0FE95409A207CFDEC" box="[381,540,511,537]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[165,1412,182,1920]" captionTargetId="figure-20@7.[162,1425,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Youtrechus zijunae n. gen., n. sp.A. head (ventral); B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642406" httpUri="https://zenodo.org/record/7642406/files/figure.png" pageId="9" pageNumber="10">Figure 3C, D</figureCitation>
): The median lobe of aedeagus quite large, distinctly arcuate near base; base large, about 1/3 as long as the median lobe; the apical half nearly straight, slightly narrowed towards apex, which is blunt and slightly bent at tip; endophallus armed with a large and long copulatory piece which is about 3/8 as long as the median lobe; in dorsal view the apical lobe much wider than long, broadly rounded at apex; parameres less developed, much shorter than the median lobe, each bearing three long setae at apex.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF31FFA0FF2F43D62376FD04" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8B8336BBBF31FFA0FF2F43D62376FD04" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="10">
<emphasis id="B948EAA9BF31FFA0FF2F43D6232EFD38" bold="true" box="[199,334,691,717]" pageId="9" pageNumber="10">Etymology.</emphasis>
Dedicated to Ms. Zijun Ma, a member of SCAU caving team, who is one of the collectors of the
<typeStatus id="54878819BF31FFA0FF7F43B222A7FD04" box="[151,199,727,753]" pageId="9" pageNumber="10">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF31FFA0FF2F439F20F2FC3C" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B8336BBBF31FFA0FF2F439F26B8FCE0" blockId="9.[151,1437,150,969]" box="[199,1240,762,789]" pageId="9" pageNumber="10">
<emphasis id="B948EAA9BF31FFA0FF2F439F2301FCE1" bold="true" box="[199,353,762,788]" pageId="9" pageNumber="10">Distribution.</emphasis>
<collectingCountry id="F32B762BBF31FFA0FE8F439E23CAFCE0" box="[359,426,763,789]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
(
<collectingRegion id="49F8F859BF31FFA0FE51439E2063FCE0" box="[441,515,763,789]" country="China" name="Hubei" pageId="9" pageNumber="10">Hubei</collectingRegion>
: Xianning). Known only from the cave Feixian Dong (
<figureCitation id="13072A3EBF31FFA0FB80439E26ABFCE0" box="[1128,1227,763,789]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[162,1426,182,1920]" captionTargetId="figure-28@8.[162,1426,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Feixian Dong, the type locality of Youtrechus zijunae n. gen., n. sp. A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera Eutrichodesmus and Skleroprotopus; F. a running individual of Youtrechus zijunae n. gen., n. sp. inside the cave." figureDoi="http://doi.org/10.5281/zenodo.7642408" httpUri="https://zenodo.org/record/7642408/files/figure.png" pageId="9" pageNumber="10">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="8B8336BBBF31FFA0FF2F427A20F2FC3C" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="10">
Located in Huaping Shan, Tingsi, southern Xianning City, Feixian Dong is
<quantity id="4CC49B5EBF31FFA0FBCF427B2617FCCC" box="[1063,1143,798,825]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="km" value="1.5">1.5 km</quantity>
long. This obsolete show cave retains natural conditions in some parts along the main passage which are suitable for cave animals (
<figureCitation id="13072A3EBF31FFA0FAB9422622BFFC74" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[162,1426,182,1920]" captionTargetId="figure-28@8.[162,1426,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Feixian Dong, the type locality of Youtrechus zijunae n. gen., n. sp. A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera Eutrichodesmus and Skleroprotopus; F. a running individual of Youtrechus zijunae n. gen., n. sp. inside the cave." figureDoi="http://doi.org/10.5281/zenodo.7642408" httpUri="https://zenodo.org/record/7642408/files/figure.png" pageId="9" pageNumber="10">Figure 4AC</figureCitation>
). The beetles were collected from several sites inside the cave, on the ground or under stones, from
<quantity id="4CC49B5EBF31FFA0FACF420222CDFC50" metricMagnitude="2" metricUnit="m" metricValue="4.8" metricValueMax="8.0" metricValueMin="1.6" pageId="9" pageNumber="10" unit="m" value="480.0" valueMax="800.0" valueMin="160.0">160 to 800 m</quantity>
from the entrance (
<figureCitation id="13072A3EBF31FFA0FE6E42EE2399FC50" box="[390,505,907,933]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[162,1426,182,1920]" captionTargetId="figure-28@8.[162,1426,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Feixian Dong, the type locality of Youtrechus zijunae n. gen., n. sp. A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera Eutrichodesmus and Skleroprotopus; F. a running individual of Youtrechus zijunae n. gen., n. sp. inside the cave." figureDoi="http://doi.org/10.5281/zenodo.7642408" httpUri="https://zenodo.org/record/7642408/files/figure.png" pageId="9" pageNumber="10">Figure 4F</figureCitation>
). Other animals observed in Feixian Dong were bats, millipedes of
<taxonomicName id="4C3C4D38BF31FFA0FB0042EE22D7FC3C" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B948EAA9BF31FFA0FB0042EE27FCFC50" box="[1256,1436,907,933]" italics="true" pageId="9" pageNumber="10">Eutrichodesmus</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C3C4D38BF31FFA0FF0642CA23DEFC3C" box="[238,446,943,969]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B948EAA9BF31FFA0FF0642CA23F7FC3C" box="[238,407,943,969]" italics="true" pageId="9" pageNumber="10">Skleroprotopus</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13072A3EBF31FFA0FE2442CA20E5FC3C" box="[460,645,943,969]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[162,1426,182,1920]" captionTargetId="figure-28@8.[162,1426,181,1920]" captionTargetPageId="8" captionText="FIGURE 4. Feixian Dong, the type locality of Youtrechus zijunae n. gen., n. sp. A. main entrance; B. a window; C. main passage; D. Bats on roof of a chamber; E &amp; G. millipedes of the genera Eutrichodesmus and Skleroprotopus; F. a running individual of Youtrechus zijunae n. gen., n. sp. inside the cave." figureDoi="http://doi.org/10.5281/zenodo.7642408" httpUri="https://zenodo.org/record/7642408/files/figure.png" pageId="9" pageNumber="10">Figure 4D, E, G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>