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<document id="997F8DC2080951D8C8AA9C88E84BBE3A" ID-DOI="10.5281/zenodo.3699463" ID-GBIF-Dataset="8a1b577e-5fce-489a-9777-32c9564a26b2" ID-ISSN="1942-1354" ID-Zenodo-Dep="3699463" ID-ZooBank="938220AB-7950-4850-88DC-3C3D8AB72C89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583500003806" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="DB4387E1713AFFAAFF410ABB0B9FFCA3" docLanguage="en" docName="InsectaMundi.0624.1-13.pdf.imf" docOrigin="Insecta Mundi 2018 (624)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nigidius helleri Boileau 1905" docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="277AFF997137FFA7FF810A620E3FFFA9" masterDocTitle="Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="12" updateTime="1698725587556" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="89EB8E15069E94CCEDEC3408AF36C393">Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
</mods:titleInfo>
<mods:name id="F3B6AC5C0D1A479A32D026303DA5F6DB" type="personal">
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<mods:roleTerm id="F3E05104E85306A5FB1FF2B6F4C3E8DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69817BFD7D8283170C078BFEC6E67854">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="A3FACD31D07E2BE793C7AAF9AEA9F41B">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
<mods:nameIdentifier id="00572136CF048297981A8F91850912E3" type="email">mjpaulsen@unl.edu</mods:nameIdentifier>
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<mods:title id="6D1A435669023FE0C9689818080FBAA9">Insecta Mundi</mods:title>
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<mods:part id="8837ACD76CA0C2F4A2366F434042AC34">
<mods:date id="745D38A5209F884BE455314FE0E88FFD">2018</mods:date>
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<mods:number id="07F772686EF16D69AF0A9B344ED690C8">2018-04-27</mods:number>
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<mods:detail id="0843D8BB84C58CF5DCFD733C3041EBEE" type="volume">
<mods:number id="BDDFC55FE94DD7EB70E6DA008FFAB6D6">2018</mods:number>
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<mods:number id="38C88914F61317F6A5477697D6B16725">624</mods:number>
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<mods:classification id="2ED471D099152B9B5DE247BA7F238CBF">journal article</mods:classification>
<mods:identifier id="358C86983AE3DE51621A6D2DF061E045" type="DOI">10.5281/zenodo.3699463</mods:identifier>
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<mods:identifier id="0DB234D928FFDE3E030355E6A8F060D3" type="ISSN">1942-1354</mods:identifier>
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<treatment id="DB4387E1713AFFAAFF410ABB0B9FFCA3" ID-DOI="http://doi.org/10.5281/zenodo.3704536" ID-GBIF-Taxon="162463117" ID-Zenodo-Dep="3704536" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3" httpUri="http://treatment.plazi.org/id/DB4387E1713AFFAAFF410ABB0B9FFCA3" lastPageNumber="12" pageId="13" pageNumber="12">
<subSubSection id="1BF0657C713AFFAAFF410ABB0C44FF58" box="[192,635,217,241]" pageId="13" pageNumber="12" type="nomenclature">
<paragraph id="535536F7713AFFAAFF410ABB0C44FF58" blockId="13.[192,635,217,241]" box="[192,635,217,241]" pageId="13" pageNumber="12">
<heading id="081D819B713AFFAAFF410ABB0C44FF58" bold="true" box="[192,635,217,241]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
<treatmentCitationGroup id="73FA11D9713AFFAAFF410ABB0C44FF58" box="[192,635,217,241]" pageId="13" pageNumber="12">
<treatmentCitation id="D24B10E6713AFFAAFF410ABB0C44FF58" author="Boileau" box="[192,635,217,241]" page="47" pageId="13" pageNumber="12" year="1905">
<taxonomicName id="94EA4D74713AFFAAFF410ABB0C44FF58" ID-CoL="6RY7S" authority="Boileau 1905: 47" authorityName="Boileau" authorityPageNumber="47" authorityYear="1905" box="[192,635,217,241]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="helleri">
<emphasis id="619EEAE5713AFFAAFF410ABB0C44FF58" bold="true" box="[192,635,217,241]" pageId="13" pageNumber="12">
<emphasis id="619EEAE5713AFFAAFF410ABB0FA7FF58" bold="true" box="[192,408,217,241]" italics="true" pageId="13" pageNumber="12">Nigidius helleri</emphasis>
<bibRefCitation id="377B4B06713AFFAAFE210ABB0C44FF58" author="Boileau" box="[416,635,217,241]" firstAuthor="Boileau" pageId="13" pageNumber="12" pagination="47 - 48" refId="ref6538" refString="Boileau, M. H. 1905. Description de Coleopteres nouveaux. Le Naturaliste 27 (431): 47 - 48." type="journal article" year="1905">Boileau 1905: 47</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713AFFAAFF410B780C70FDAF" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="535536F7713AFFAAFF410B780F01FE3D" blockId="13.[192,1440,282,404]" pageId="13" pageNumber="12">
<emphasis id="619EEAE5713AFFAAFF410B780FB9FE9B" bold="true" box="[192,390,282,306]" pageId="13" pageNumber="12">Type material.</emphasis>
<materialsCitation id="E3823CAA713AFFAAFE0A0B780F01FE3D" ID-GBIF-Occurrence="2574936446" collectionCode="MNHN" location="Palembang" pageId="13" pageNumber="12" specimenCode="EC4652" specimenCount="2" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="8C518855713AFFAAFE0A0B780FC7FE9B" box="[395,504,282,306]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
female (
<figureCitation id="CBD12A72713AFFAAFDDC0B780C9EFE9B" box="[605,673,282,306]" captionStart="Figures 6–7" captionStartId="7.[192,287,1471,1493]" captionTargetBox="[233,1354,216,1395]" captionTargetId="figure@7.[220,1364,210,1415]" captionTargetPageId="7" captionText="Figures 6–7. Nigidius helleri Boileau. 6) Major male, dorsal habitus. 7) Holotype male with labels. Photo courtesy of Antoine Mantilleri, MNHN." figureDoi="http://doi.org/10.5281/zenodo.3699473" httpUri="https://zenodo.org/record/3699473/files/figure.png" pageId="13" pageNumber="12">Fig. 7</figureCitation>
),
<collectionCode id="35FBAE32713AFFAAFD340B790D2DFE9B" box="[693,786,283,306]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="12">MNHN</collectionCode>
, labeled: a) “Palembang /
<collectingRegion id="912EF815713AFFAAFBC00B780A91FE9B" box="[1089,1198,282,306]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="12">Sumatra</collectingRegion>
”; b) “TYPE / [H.B.]”; c) handwritten, “Helleri / type”; d) “ex coll. / Boileau”; e) on red paper, “TYPE”; f) on red paper, “HOLOTYPE”; g) “
<typeStatus id="8C518855713AFFAAFECB0B390FDAFEDA" box="[330,485,347,371]" pageId="13" pageNumber="12" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="94EA4D74713AFFAAFE7B0B390C5BFEDA" box="[506,612,347,371]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713AFFAAFE7B0B390C5BFEDA" box="[506,612,347,371]" italics="true" pageId="13" pageNumber="12">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="94EA4D74713AFFAAFDFB0B390D10FEDA" authority="Boileau" authorityName="Boileau" box="[634,815,347,371]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="helleri">
<emphasis id="619EEAE5713AFFAAFDFB0B390CF7FEDA" box="[634,712,347,371]" italics="true" pageId="13" pageNumber="12">helleri</emphasis>
Boileau
</taxonomicName>
”; h) “
<collectionCode id="35FBAE32713AFFAAFCEE0B390DF5FEDA" box="[879,970,347,371]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="12">MNHN</collectionCode>
/
<specimenCode id="034C9E8C713AFFAAFC610B390A7BFEDA" box="[992,1092,347,371]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="12">EC4652</specimenCode>
”. Type locality: “
<location id="5635602C713AFFAAFA910B390BA3FEDA" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFA910B390BA3FEDA" box="[1296,1436,347,371]" name="Palembang" pageId="13" pageNumber="12" stateProvince="Sumatera Utara">Palembang</location>
,
<collectingRegion id="912EF815713AFFAAFF410B1E0F11FE3D" box="[192,302,380,404]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="12">Sumatra</collectingRegion>
”.
</materialsCitation>
</paragraph>
<paragraph id="535536F7713AFFAAFF410BCE0C70FDAF" blockId="13.[192,1440,428,518]" pageId="13" pageNumber="12">
<emphasis id="619EEAE5713AFFAAFF410BCE0C56FE6D" bold="true" box="[192,617,428,452]" pageId="13" pageNumber="12">Additional material examined.</emphasis>
<materialsCitation id="E3823CAA713AFFAAFDF30BCE0B03FE6C" ID-GBIF-Occurrence="2574936448" box="[626,1340,428,453]" collectionCode="SLTC" country="Indonesia" location="Tebo / Teluk Kayu Putih" pageId="13" pageNumber="12" specimenCount="1" stateProvince="JAMBI">
<emphasis id="619EEAE5713AFFAAFDF30BCE0D23FE6D" bold="true" box="[626,796,428,452]" pageId="13" pageNumber="12">
<collectingCountry id="2BFD7667713AFFAAFDF30BCE0D23FE6D" box="[626,796,428,452]" name="Indonesia" pageId="13" pageNumber="12">INDONESIA</collectingCountry>
</emphasis>
:
<collectingRegion id="912EF815713AFFAAFCAC0BCF0DB5FE6C" box="[813,906,429,453]" country="Indonesia" name="Jambi" pageId="13" pageNumber="12">JAMBI</collectingRegion>
:
<location id="5635602C713AFFAAFC160BCE0A85FE6D" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFC160BCE0A85FE6D" box="[919,1210,428,453]" country="Indonesia" name="Tebo / Teluk Kayu Putih" pageId="13" pageNumber="12" stateProvince="JAMBI">Tebo/Teluk Kayu Putih</location>
(3
<collectionCode id="35FBAE32713AFFAAFB650BCF0B0FFE6C" box="[1252,1328,429,453]" pageId="13" pageNumber="12">SLTC</collectionCode>
).
</materialsCitation>
<materialsCitation id="E3823CAA713AFFAAFAC40BCF0CD5FE4C" ID-GBIF-Occurrence="2574936431" collectionCode="SLTC" location="Annai Valley / Lapai" pageId="13" pageNumber="12" specimenCount="1" stateProvince="TERA BARAT">
<collectingRegion id="912EF815713AFFAAFAC40BCF0F4BFE4C" country="Indonesia" name="TERA BARAT" pageId="13" pageNumber="12">SUMATERA BARAT</collectingRegion>
:
<location id="5635602C713AFFAAFEFE0BAF0C52FE4C" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFEFE0BAF0C52FE4C" box="[383,621,461,485]" name="Annai Valley / Lapai" pageId="13" pageNumber="12" stateProvince="TERA BARAT">Annai Valley/Lapai</location>
(2
<collectionCode id="35FBAE32713AFFAAFD120BAF0CE1FE4C" box="[659,734,461,485]" pageId="13" pageNumber="12">SLTC</collectionCode>
);
</materialsCitation>
<materialsCitation id="E3823CAA713AFFAAFD700BAC0A15FE4C" ID-GBIF-Occurrence="2574936464" box="[753,1066,461,485]" collectionCode="SLTC" location="Mt. Singgalang" pageId="13" pageNumber="12" specimenCount="1">
<location id="5635602C713AFFAAFD700BAC0D92FE4C" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFD700BAC0D92FE4C" box="[753,941,461,485]" name="Mt. Singgalang" pageId="13" pageNumber="12">Mt. Singgalang</location>
(2
<collectionCode id="35FBAE32713AFFAAFC520BAF0A21FE4C" box="[979,1054,461,485]" pageId="13" pageNumber="12">SLTC</collectionCode>
);
</materialsCitation>
<materialsCitation id="E3823CAA713AFFAAFBB00BAC0B03FE4C" ID-GBIF-Occurrence="2574936436" box="[1073,1340,461,485]" collectionCode="MJPC" location="Mt. Talang" pageId="13" pageNumber="12" specimenCount="1">
<location id="5635602C713AFFAAFBB00BAC0A86FE4C" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFBB00BAC0A86FE4C" box="[1073,1209,461,485]" name="Mt. Talang" pageId="13" pageNumber="12">Mt. Talang</location>
(9
<collectionCode id="35FBAE32713AFFAAFB5E0BAF0B10FE4C" box="[1247,1327,461,485]" collectionName="MJPC" pageId="13" pageNumber="12">MJPC</collectionCode>
);
</materialsCitation>
<materialsCitation id="E3823CAA713AFFAAFAC20BAF0F74FDAF" ID-GBIF-Occurrence="2574936441" collectionCode="MJPC" location="Padang" pageId="13" pageNumber="12" specimenCount="1">
<location id="5635602C713AFFAAFAC20BAF0B9FFE4C" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFAC20BAF0B9FFE4C" box="[1347,1440,461,485]" name="Padang" pageId="13" pageNumber="12">Padang</location>
(10
<collectionCode id="35FBAE32713AFFAAFF6F0B8C0F01FDAF" box="[238,318,494,518]" collectionName="MJPC" pageId="13" pageNumber="12">MJPC</collectionCode>
);
</materialsCitation>
<materialsCitation id="E3823CAA713AFFAAFED30B8C0C70FDAF" ID-GBIF-Occurrence="2574936472" box="[338,591,494,518]" collectionCode="MJPC" location="Pariaman" pageId="13" pageNumber="12" specimenCount="1">
<location id="5635602C713AFFAAFED30B8C0FF3FDAF" LSID="urn:lsid:plazi:treatment:DB4387E1713AFFAAFF410ABB0B9FFCA3:5635602C713AFFAAFED30B8C0FF3FDAF" box="[338,460,494,518]" name="Pariaman" pageId="13" pageNumber="12">Pariaman</location>
(
<specimenCount id="45ECFD7E713AFFAAFE5F0B8C0FD5FDAF" box="[478,490,494,518]" pageId="13" pageNumber="12" type="generic">1</specimenCount>
<collectionCode id="35FBAE32713AFFAAFE730B8C0C7CFDAF" box="[498,579,494,518]" collectionName="MJPC" pageId="13" pageNumber="12">MJPC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713AFFAAFF41087C0A64FD31" pageId="13" pageNumber="12" type="diagnosis">
<paragraph id="535536F7713AFFAAFF41087C0A64FD31" blockId="13.[192,1440,542,664]" pageId="13" pageNumber="12">
<emphasis id="619EEAE5713AFFAAFF41087C0F73FD9F" bold="true" box="[192,332,542,566]" pageId="13" pageNumber="12">Diagnosis.</emphasis>
The Sumatran member of the obesus group is recognizable by its more homogeneous pronotal punctures that, although variable in size, are regularly distributed and never fine enough to give the appearance of impunctate areas (
<figureCitation id="CBD12A72713AFFAAFDD5083D0CA5FDD1" box="[596,666,607,632]" captionStart="Figures 8–9" captionStartId="8.[192,286,617,639]" captionTargetBox="[214,1406,223,582]" captionTargetId="figure@8.[192,1440,210,592]" captionTargetPageId="8" captionText="Figures 8–9. Pronotal punctation of obesus group species. 8) Surface more uniformly punctate, N. helleri. 9) Surface variable with almost impunctate and coarsely punctate areas, N. gravelyi Paulsen, n. sp." figureDoi="http://doi.org/10.5281/zenodo.3699475" httpUri="https://zenodo.org/record/3699475/files/figure.png" pageId="13" pageNumber="12">Fig. 8</figureCitation>
). In addition, the anterior lateral margin of the pronotum is the least produced in
<taxonomicName id="94EA4D74713AFFAAFE1808E20C33FD31" box="[409,524,640,664]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="helleri">
<emphasis id="619EEAE5713AFFAAFE1808E20C33FD31" box="[409,524,640,664]" italics="true" pageId="13" pageNumber="12">N. helleri</emphasis>
</taxonomicName>
, and the seta of the pronotum are most distinct.
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713AFFAAFF4108D30B9FFCA3" pageId="13" pageNumber="12" type="discussion">
<paragraph id="535536F7713AFFAAFF4108D30B9FFCA3" blockId="13.[192,1440,688,778]" pageId="13" pageNumber="12">
<emphasis id="619EEAE5713AFFAAFF4108D30F7FFD60" bold="true" box="[192,320,689,713]" pageId="13" pageNumber="12">Remarks.</emphasis>
This name has until now been misapplied to different species in the cornutus group of
<taxonomicName id="94EA4D74713AFFAAFAB608D20B9FFD61" box="[1335,1440,688,712]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713AFFAAFAB608D20B9FFD61" box="[1335,1440,688,712]" italics="true" pageId="13" pageNumber="12">Nigidius</emphasis>
</taxonomicName>
from
<collectingRegion id="912EF815713AFFAAFF7E08B00F06FD40" box="[255,313,722,745]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="12">Java</collectingRegion>
and
<collectingRegion id="912EF815713AFFAAFEF008B30FE0FD40" box="[369,479,721,745]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="12">Sumatra</collectingRegion>
, and this will be addressed fully in a subsequent paper on that group. Specimens examined are from February, April, August, and December, indicating that adults are present year-round.
</paragraph>
</subSubSection>
</treatment>
</document>
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