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<document id="9080801760C86F8E28B9D9D16B3D1170" ID-DOI="10.5281/zenodo.10667690" ID-ISSN="1407-8619" ID-Zenodo-Dep="10667690" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708024575242" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Vitali, Francesco" docDate="2020" docId="03FCEC6BFFA9BA259AC5FE7EFCE813D8" docLanguage="en" docName="BalticJColeopterol.20.1.1-6.pdf" docOrigin="Baltic Journal of Coleopterology 20 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.1:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="1" docTitle="Kuskaella petriolii Fanti &amp; Vitali 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="FFC59413FFA8BA219A26FF89FFFF170C" masterDocTitle="A new species of the enigmatic genus Kuskaella Fanti &amp; Kupryjanowicz, 2017 (Coleoptera, Cantharidae) from Baltic amber" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1708024901013" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="6CAD0B999B3F3BF8FF8CD504AB56B1A4">A new species of the enigmatic genus Kuskaella Fanti &amp; Kupryjanowicz, 2017 (Coleoptera, Cantharidae) from Baltic amber</mods:title>
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<mods:namePart id="4AD042E42949091E2C21F36EB9BA68CE">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="F85632A959A7B1D55253D2BD31D91F55">Vitali, Francesco</mods:namePart>
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<mods:title id="C7DEDB56FA5C0AA23B3A6030F8FDF778">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="7ABE2989580343B842CAAA5D57425B58">2020</mods:date>
<mods:detail id="F56085C63ABF3F6EA8EEA6DAC70B0FE3" type="pubDate">
<mods:number id="BDED60C97BC067EC377F34E8A1A4E173">2020-08-31</mods:number>
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<mods:detail id="19D10CAFF0392FF1ABB7D6B3A5BDDC4B" type="volume">
<mods:number id="DCB0DC3D4CD5C52CDB6D659B995859BA">20</mods:number>
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<mods:identifier id="51B6B992C561F576C228041C51595766" type="DOI">10.5281/zenodo.10667690</mods:identifier>
<mods:identifier id="3018E2B1B0B627902942BC0AFC205FEC" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03FCEC6BFFA9BA259AC5FE7EFCE813D8" LSID="urn:lsid:plazi:treatment:03FCEC6BFFA9BA259AC5FE7EFCE813D8" httpUri="http://treatment.plazi.org/id/03FCEC6BFFA9BA259AC5FE7EFCE813D8" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
<subSubSection id="C34F0EF6FFA9BA209AC5FE7EFDF81503" box="[227,519,503,527]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BEA5D7DFFA9BA209AC5FE7EFDF81503" blockId="1.[227,519,503,561]" box="[227,519,503,527]" pageId="1" pageNumber="2">
<heading id="D0A2EA11FFA9BA209AC5FE7EFDF81503" bold="true" box="[227,519,503,527]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="4">
<emphasis id="B921816FFFA9BA209AC5FE7EFDF81503" bold="true" box="[227,519,503,527]" pageId="1" pageNumber="2">
<taxonomicName id="4C5526FEFFA9BA209AC5FE7EFE511503" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[227,430,503,527]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFA9BA209AC5FE7EFE511503" bold="true" box="[227,430,503,527]" italics="true" pageId="1" pageNumber="2">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFA9BA209B92FE7EFDF81503" box="[436,519,503,527]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFA9BA209AC5FD90FEAA153D" box="[227,341,537,561]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BEA5D7DFFA9BA209AC5FD90FEAA153D" blockId="1.[227,519,503,561]" box="[227,341,537,561]" pageId="1" pageNumber="2">
(
<figureCitation id="136E41F8FFA9BA209ACCFD90FEB3153D" box="[234,332,537,561]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[303,343,854,878]" captionStartId-1="2.[303,341,1660,1684]" captionStartId-2="3.[227,266,986,1010]" captionText-0="Fig. 1. Kuskaella petriolii n. sp. in Baltic amber (BaA06RU). A: Holotype, habitus (lateral view), scale bar = 500 Pm; B: Holotype, habitus (frontal view), scale bar = 400 Pm; C: Holotype, detail of tarsomeres and claws, scale bar = 50 Pm; D: Holotype, detail of antenna, scale bar = 1.0 mm." captionText-1="Fig. 2. Kuskaella petriolii n. sp. in Baltic amber (BaA06RU).A: Holotype, detail of head and pronotum, scale bar = 100 Pm; B: Holotype, detail of elytra (lateral view), scale bar = 500 Pm; C: Holotype, detail of humeral zone, scale bar = 500 Pm; D: Holotype, detail of elytral apex, scale bar = 400 Pm." captionText-2="Fig. 3. Kuskaella petriolii n. sp. in Baltic amber (BaA06RU). Holotype, detail of pronotum, scale bar =100 Pm." pageId="1" pageNumber="2">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFA9BA209AC5FDD7FE601208" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BEA5D7DFFA9BA209AC5FDD7FEAC15B7" blockId="1.[227,736,606,699]" pageId="1" pageNumber="2">
<materialsCitation id="3B3D5720FFA9BA209AC5FDD7FEAC15B7" collectorName="W. Lewita &amp; F. Fanti" country="Poland" location="Gdansk" pageId="1" pageNumber="2" specimenCount="1" typeStatus="holotype">
<emphasis id="B921816FFFA9BA209AC5FDD7FEB0157A" bold="true" box="[227,335,606,630]" pageId="1" pageNumber="2">
<typeStatus id="54EEE3DFFFA9BA209AC5FDD7FEB4157A" box="[227,331,606,630]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, Baltic amber, ex coll.
<collectorName id="26A038ABFFA9BA209850FDD6FD1F157A" box="[630,736,606,630]" pageId="1" pageNumber="2">W. Lewita</collectorName>
(
<location id="8E8A0BA6FFA9BA209ACCFD09FEBC1594" LSID="urn:lsid:plazi:treatment:03FCEC6BFFA9BA259AC5FE7EFCE813D8:8E8A0BA6FFA9BA209ACCFD09FEBC1594" box="[234,323,640,664]" country="Poland" name="Gdansk" pageId="1" pageNumber="2">Gdańsk</location>
,
<collectingCountry id="F3421DEDFFA9BA209B77FD09FE5D1594" box="[337,418,640,664]" name="Poland" pageId="1" pageNumber="2">Poland</collectingCountry>
),
<collectorName id="26A038ABFFA9BA209B9FFD08FDED1594" box="[441,530,640,664]" pageId="1" pageNumber="2">F. Fanti</collectorName>
coll., access code BaA06RU.
</materialsCitation>
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FD6EFE121485" blockId="1.[227,737,743,905]" pageId="1" pageNumber="2">
The amber piece is nearly shaped like a half-moon and measures approximately 26 x
<quantity id="4CADF098FFA9BA209866FC83FD72142E" box="[576,653,778,802]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="2" unit="mm" value="10.0">10 mm</quantity>
. The inclusion is complete, except for the left antenna totally missing. The mandibles are covered by turbidity and not visible.
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FC3FFE9214C2" blockId="1.[227,365,950,974]" box="[227,365,950,974]" pageId="1" pageNumber="2">
<emphasis id="B921816FFFA9BA209AC5FC3FFE9214C2" bold="true" box="[227,365,950,974]" pageId="1" pageNumber="2">Type locality</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FC73FDB6131E" blockId="1.[227,585,1018,1043]" box="[227,585,1018,1043]" pageId="1" pageNumber="2">Gdańsk Bay - Sambian Peninsula.</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FBC9FE7E1354" blockId="1.[227,385,1088,1112]" box="[227,385,1088,1112]" pageId="1" pageNumber="2">
<emphasis id="B921816FFFA9BA209AC5FBC9FE7E1354" bold="true" box="[227,385,1088,1112]" pageId="1" pageNumber="2">Syninclusions</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FB0DFE601208" blockId="1.[227,737,1156,1284]" pageId="1" pageNumber="2">
One
<taxonomicName id="4C5526FEFFA9BA209B30FB0DFE881390" box="[278,375,1156,1180]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
(
<taxonomicName id="4C5526FEFFA9BA209BA3FB0DFDFD1390" box="[389,514,1156,1180]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
), two Dipterans, two Heteropteran neanids, an indeterminable disarticulated insect, air bubbles, botanical remains, and stellate hairs.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFA9BA209AC5FAB8FBD41657" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BEA5D7DFFA9BA209AC5FAB8FE2C1245" blockId="1.[227,467,1329,1353]" box="[227,467,1329,1353]" pageId="1" pageNumber="2">
<emphasis id="B921816FFFA9BA209AC5FAB8FE2C1245" bold="true" box="[227,467,1329,1353]" pageId="1" pageNumber="2">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209AC5FAFCFBD41657" blockId="1.[227,738,1397,1697]" lastBlockId="1.[774,1285,220,347]" pageId="1" pageNumber="2">
<taxonomicName id="4C5526FEFFA9BA209AC5FAFCFE4E1281" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[227,433,1397,1421]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFA9BA209AC5FAFCFE4E1281" box="[227,433,1397,1421]" italics="true" pageId="1" pageNumber="2">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFA9BA209B9FFAFFFE0A1281" box="[441,501,1398,1421]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
is easily distinguishable from
<taxonomicName id="4C5526FEFFA9BA209B6EFA11FD4912A3" authority="Fanti &amp; Damgaard, 2018" authorityName="Fanti &amp; Damgaard" authorityYear="2018" box="[328,694,1431,1455]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="bajerae">
<emphasis id="B921816FFFA9BA209B6EFA11FE4B12A3" box="[328,436,1431,1455]" italics="true" pageId="1" pageNumber="2">K. bajerae</emphasis>
<bibRefCitation id="EFC4208CFFA9BA209B9EFA1EFD4912A3" author="Fanti F. &amp; Damgaard A. L." box="[440,694,1431,1455]" pageId="1" pageNumber="2" pagination="1 - 32" refId="ref1746" refString="Fanti F., Damgaard A. L. 2018. Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 18 (1): 1 - 32." type="journal article" year="2018">Fanti &amp; Damgaard, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5526FEFFA9BA209AC5FA32FD3C12DE" authority="Fanti &amp; Kupryjanowicz, 2017" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2017" box="[227,707,1466,1490]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="macroptera">
<emphasis id="B921816FFFA9BA209AC5FA32FE7F12DE" box="[227,384,1467,1490]" italics="true" pageId="1" pageNumber="2">K. macroptera</emphasis>
<bibRefCitation id="EFC4208CFFA9BA209BA3FA33FD3C12DE" author="Fanti F. &amp; Kupryjanowicz J." box="[389,707,1466,1490]" pageId="1" pageNumber="2" pagination="785 - 788" refId="ref1788" refString="Fanti F., Kupryjanowicz J., 2017. A new soldier beetle from Eocene Baltic amber. Acta Palaeontologica Polonica, 62 (4): 785 - 788." type="journal article" year="2017">Fanti &amp; Kupryjanowicz, 2017</bibRefCitation>
</taxonomicName>
by its last maxillary palpomere without an elongated tip at the apex (with a more or less elongated point in the other species). Furthermore, it differs from both species in its pronotum, sub-rectangular with straight sides instead of apically narrowed from the basal half, as in
<taxonomicName id="4C5526FEFFA9BA209876F903FD4111AC" authorityName="Fanti &amp; Damgaard" authorityYear="2018" box="[592,702,1672,1697]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="bajerae">
<emphasis id="B921816FFFA9BA209876F903FD4111AC" box="[592,702,1672,1697]" italics="true" pageId="1" pageNumber="2">K. bajerae</emphasis>
</taxonomicName>
, or abruptly (male) or gradually (female) narrowed from the basal third, as in
<taxonomicName id="4C5526FEFFA9BA209E35FF76FB55161A" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2017" box="[1043,1194,255,278]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="macroptera">
<emphasis id="B921816FFFA9BA209E35FF76FB55161A" box="[1043,1194,255,278]" italics="true" pageId="1" pageNumber="2">K. macroptera</emphasis>
</taxonomicName>
. Finally,
<taxonomicName id="4C5526FEFFA9BA209920FEA9FC301634" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[774,975,288,312]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFA9BA209920FEA9FC301634" box="[774,975,288,312]" italics="true" pageId="1" pageNumber="2">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFA9BA2099F2FEA8FBF21634" box="[980,1037,289,312]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C5526FEFFA9BA209EB2FEA8FAFA1634" authorityName="Fanti &amp; Damgaard" authorityYear="2018" box="[1172,1285,288,312]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="bajerae">
<emphasis id="B921816FFFA9BA209EB2FEA8FAFA1634" box="[1172,1285,288,312]" italics="true" pageId="1" pageNumber="2">K. bajerae</emphasis>
</taxonomicName>
in its longer antennomeres.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFA9BA229920FE01FB16139A" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="8BEA5D7DFFA9BA209920FE01FC7716AC" blockId="1.[774,904,392,416]" box="[774,904,392,416]" pageId="1" pageNumber="2">
<emphasis id="B921816FFFA9BA209920FE01FC7716AC" bold="true" box="[774,904,392,416]" pageId="1" pageNumber="2">Description</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209920FE45FC241547" blockId="1.[774,1285,460,587]" pageId="1" pageNumber="2">
Adult, winged, slender. Male, based on the long antennae and the last sternite elongated and narrower than last tergite. Body length about
<quantity id="4CADF098FFA9BA209EC4FD98FCCD1547" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="2" unit="mm" value="5.8">5.8 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BEA5D7DFFA9BA209920FDF1FBA913BB" blockId="1.[774,1286,632,1207]" pageId="1" pageNumber="2">Head exposed, wide and broad, as wide as pronotum, slightly triangular behind eyes, covered with shallow dense punctation. Mandibles not visible. Maxillary palpomeres unequal; first segment very short, second elongated and globular; third small and short; last palpomere globular, apically thinner, without elongated point. Labial palps 3-segmented, last palpomere globular. Eyes large, roundish, prominent. Antennae 11-segmented, filiform, long, almost reaching the elytral apex; covered with short sparse setae; scape long, club-shaped; pedicel robust and more than two times shorter than scape; antennomeres III-IX very elongate, sub-equal, almost as long as scape; antennomere X very slightly shorter than previous ones; antennomere XI elongate with rounded apex.</paragraph>
<paragraph id="8BEA5D7DFFA9BA209920FB6DFB5012C6" blockId="1.[774,1285,1251,1482]" pageId="1" pageNumber="2">Pronotum transverse, sub-rectangular, slightly convex; apical margin straight and very slightly bordered; basal margin undulate; sides straight and bordered; surface wrinkled, covered with shallow dense punctation and with a small depression near the middle of the basal margin. Scutellum triangular with rounded apex.</paragraph>
<paragraph id="8BEA5D7DFFA9BA229920FA7EFE40139A" blockId="1.[774,1285,1527,1688]" lastBlockId="3.[227,736,1115,1174]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">Elytra long, slender, slightly wider than pronotal base and slightly surpassing the last abdominal segment, parallel-sided, rounded at apex, surface wrinkled and covered with short recumbent setae. Posterior wings as long as elytra and completely covered by them except for a small part near the elytral apex.</paragraph>
<caption id="DF2A0DF5FFAABA239B09FCDFFB0014A2" pageId="2" pageNumber="3" startId="2.[303,343,854,878]" targetType="figure">
<paragraph id="8BEA5D7DFFAABA239B09FCDFFB0014A2" blockId="2.[303,1359,854,942]" pageId="2" pageNumber="3">
Fig. 1.
<taxonomicName id="4C5526FEFFAABA239B5BFCDFFDB51462" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[381,586,854,878]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFAABA239B5BFCDFFDB51462" box="[381,586,854,878]" italics="true" pageId="2" pageNumber="3">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFAABA239876FCDEFD741462" box="[592,651,855,878]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
in Baltic amber (BaA06RU). A: Holotype, habitus (lateral view), scale bar = 500
<emphasis id="B921816FFFAABA239BF1FCFFFE191482" bold="true" box="[471,486,886,910]" pageId="2" pageNumber="3">P</emphasis>
m; B: Holotype, habitus (frontal view), scale bar = 400
<emphasis id="B921816FFFAABA239E10FCFFFBBA1482" bold="true" box="[1078,1093,886,910]" pageId="2" pageNumber="3">P</emphasis>
m; C: Holotype, detail of tarsomeres and claws, scale bar = 50
<emphasis id="B921816FFFAABA239893FC1FFD3B14A2" bold="true" box="[693,708,918,942]" pageId="2" pageNumber="3">P</emphasis>
m; D: Holotype, detail of antenna, scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF2A0DF5FFAABA239B09F9F5FB0A11D4" pageId="2" pageNumber="3" startId="2.[303,341,1660,1684]" targetType="figure">
<paragraph id="8BEA5D7DFFAABA239B09F9F5FB0A11D4" blockId="2.[303,1359,1660,1752]" pageId="2" pageNumber="3">
Fig. 2.
<taxonomicName id="4C5526FEFFAABA239B50F9F5FDC81198" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[374,567,1660,1684]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFAABA239B50F9F5FDC81198" box="[374,567,1660,1684]" italics="true" pageId="2" pageNumber="3">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFAABA23981AF9F4FD8D1198" box="[572,626,1661,1684]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
in Baltic amber (BaA06RU).A: Holotype, detail of head and pronotum, scale bar = 100
<emphasis id="B921816FFFAABA239BF7F917FE1F11BA" bold="true" box="[465,480,1694,1718]" pageId="2" pageNumber="3">P</emphasis>
m; B: Holotype, detail of elytra (lateral view), scale bar = 500
<emphasis id="B921816FFFAABA239E7AF917FB9411BA" bold="true" box="[1116,1131,1694,1718]" pageId="2" pageNumber="3">P</emphasis>
m; C: Holotype, detail of humeral zone, scale bar = 500
<emphasis id="B921816FFFAABA2398AEF949FD6811D4" bold="true" box="[648,663,1728,1752]" pageId="2" pageNumber="3">P</emphasis>
m; D: Holotype, detail of elytral apex, scale bar = 400
<emphasis id="B921816FFFAABA239EECF949FB2611D4" bold="true" box="[1226,1241,1728,1752]" pageId="2" pageNumber="3">P</emphasis>
m.
</paragraph>
</caption>
<caption id="DF2A0DF5FFABBA229AC5FC53FEB81318" pageId="3" pageNumber="4" startId="3.[227,266,986,1010]" targetType="figure">
<paragraph id="8BEA5D7DFFABBA229AC5FC53FEB81318" blockId="3.[227,1286,986,1044]" pageId="3" pageNumber="4">
Fig. 3.
<taxonomicName id="4C5526FEFFABBA229B0BFC53FE0B14FE" authority="Fanti &amp; Vitali, 2020" authorityName="Fanti &amp; Vitali" authorityYear="2020" box="[301,500,986,1010]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="petriolii" status="sp. nov.">
<emphasis id="B921816FFFABBA229B0BFC53FE0B14FE" box="[301,500,986,1010]" italics="true" pageId="3" pageNumber="4">Kuskaella petriolii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C14FFABBA229BDCFC52FDCC14FE" box="[506,563,987,1010]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
in Baltic amber (BaA06RU). Holotype, detail of pronotum, scale bar = 100
<emphasis id="B921816FFFABBA229B38FC75FED31318" bold="true" box="[286,300,1020,1044]" pageId="3" pageNumber="4">P</emphasis>
m.
</paragraph>
</caption>
<paragraph id="8BEA5D7DFFABBA229AC5FB4AFDE0124E" blockId="3.[227,737,1219,1346]" pageId="3" pageNumber="4">Sternum sub-quadrate and robust, abdominal segments strongly transverse, pubescence short, last sternite triangular and narrower than last tergite. Aedeagus not visible.</paragraph>
<paragraph id="8BEA5D7DFFABBA229AC5FAE6FB16139A" blockId="3.[227,737,1390,1724]" lastBlockId="3.[774,1285,1115,1175]" pageId="3" pageNumber="4">Legs short and robust; coxae extremely massive; trochanteres small, slightly elongated and roundish at apex; femora robust, enlarged, not curved; tibiae cylindrical; protibiae shorter than profemora; mesotibiae as long as mesofemora (or very slightly longer); metatibiae longer than metafemora; tarsal formula 5-5-5; first tarsomere elongated, slightly widened apically and approximately 1.6 times as long as second; second elongated; third shorter than previous; fourth bilobed; fifth elongated and flat; claws simple with a very small obtuse denticle at the base.</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFABBA229920FB4AFB5A1268" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BEA5D7DFFABBA229920FB4AFC8713D7" blockId="3.[774,888,1219,1243]" box="[774,888,1219,1243]" pageId="3" pageNumber="4">
<emphasis id="B921816FFFABBA229920FB4AFC8713D7" bold="true" box="[774,888,1219,1243]" pageId="3" pageNumber="4">Etymology</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFABBA229920FA8EFB5A1268" blockId="3.[774,1285,1287,1380]" pageId="3" pageNumber="4">
The new species is dedicated to our friend Andrea Petrioli (Asciano,
<collectingCountry id="F3421DEDFFABBA229E3DFAA3FBB0124E" box="[1051,1103,1322,1346]" name="Italy" pageId="3" pageNumber="4">Italy</collectingCountry>
), specialist of endogenous fauna, especially
<taxonomicName id="4C5526FEFFABBA229E09FAC5FB5E1268" authorityName="Latreille" authorityYear="1802" box="[1071,1185,1356,1380]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EF6FFABBA259920FA18FCE813D8" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BEA5D7DFFABBA229920FA18FC7D12A5" blockId="3.[774,898,1425,1449]" box="[774,898,1425,1449]" pageId="3" pageNumber="4">
<emphasis id="B921816FFFABBA229920FA18FC7D12A5" bold="true" box="[774,898,1425,1449]" pageId="3" pageNumber="4">Discussion</emphasis>
</paragraph>
<paragraph id="8BEA5D7DFFABBA229920FA5FFBE21174" blockId="3.[774,1285,1494,1656]" pageId="3" pageNumber="4">
Mouth characters of
<taxonomicName id="4C5526FEFFABBA2299C2FA5FFBAE12E2" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2017" box="[996,1105,1494,1518]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B921816FFFABBA2299C2FA5FFBAE12E2" box="[996,1105,1494,1518]" italics="true" pageId="3" pageNumber="4">Kuskaella</emphasis>
</taxonomicName>
provide a strong basis to classify this genus in the
<taxonomicName id="4C5526FEFFABBA229E51FA71FB00111C" authorityName="Kiesenwetter" authorityYear="1852" box="[1143,1279,1528,1552]" pageId="3" pageNumber="1" rank="subFamily" subFamily="Malthininae">Malthininae</taxonomicName>
, but the fact that the mandibles are not visible in all known specimens leaves it questionable to its exact systematic position.
</paragraph>
<paragraph id="8BEA5D7DFFACBA259B09FF5EFD1616D3" blockId="4.[303,813,214,479]" pageId="4" pageNumber="5">
In fact, though long elytra are only known in Malchinini
<bibRefCitation id="EFC4208CFFACBA259B9EFF71FD88161C" author="Brancucci M." box="[440,631,248,272]" pageId="4" pageNumber="5" pagination="215 - 388" refId="ref1580" refString="Brancucci M. 1980. Morphologie comparee, evolution et systematique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5: 215 - 388." type="journal article" year="1980">Brancucci, 1980</bibRefCitation>
, Mimoplatycini
<bibRefCitation id="EFC4208CFFACBA259B09FE95FE1A163F" author="Kazantsev S. V." box="[303,485,283,307]" pageId="4" pageNumber="5" pagination="283 - 291" refId="ref1870" refString="Kazantsev S. V. 2013. New taxa of Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) with synonymic and taxonomic notes. Russian Entomological Journal, 22 (4): 283 - 291." type="journal article" year="2013">Kazantsev, 2013</bibRefCitation>
and some extinct Malthinini Kiesenwetter, 1852 (
<bibRefCitation id="EFC4208CFFACBA259821FEB7FD4B165A" author="Brancucci M." box="[519,692,318,342]" pageId="4" pageNumber="5" pagination="215 - 388" refId="ref1580" refString="Brancucci M. 1980. Morphologie comparee, evolution et systematique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5: 215 - 388." type="journal article" year="1980">Brancucci 1980</bibRefCitation>
;
<bibRefCitation id="EFC4208CFFACBA25989BFEB6FE991674" author="Kazantsev S. V." pageId="4" pageNumber="5" pagination="283 - 291" refId="ref1870" refString="Kazantsev S. V. 2013. New taxa of Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) with synonymic and taxonomic notes. Russian Entomological Journal, 22 (4): 283 - 291." type="journal article" year="2013">Kazantsev 2013</bibRefCitation>
;
<bibRefCitation id="EFC4208CFFACBA259B49FEE9FD891674" author="Fanti F. &amp; Castiglione E." box="[367,630,352,376]" pageId="4" pageNumber="5" pagination="123 - 127" refId="ref1710" refString="Fanti F., Castiglione E. 2017. Description of a new genus and species of Cantharidae from Eocene Baltic amber (Insecta, Coleptera). Palaeodiversity, 10: 123 - 127." type="journal article" year="2017">Fanti &amp; Castiglione 2017</bibRefCitation>
), and the pronotal structure excludes Mimoplatycini, the absence of the mandibular characters does not allow discriminating between the remaining tribes.
</paragraph>
<paragraph id="8BEA5D7DFFACBA259B09FD84FE9115A1" blockId="4.[303,813,524,685]" pageId="4" pageNumber="5">
In addition,
<taxonomicName id="4C5526FEFFACBA259BF7FD85FDB51528" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2017" box="[465,586,524,548]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B921816FFFACBA259BF7FD85FDB51528" box="[465,586,524,548]" italics="true" pageId="4" pageNumber="5">Kuskaella</emphasis>
</taxonomicName>
shows a peculiar aedeagus with a very elongated lobe, which is rounded-globular at the apex and equipped with a horizontal appendix (
<bibRefCitation id="EFC4208CFFACBA25980DFDFAFE9C15A1" author="Fanti F. &amp; Kupryjanowicz J." pageId="4" pageNumber="5" pagination="785 - 788" refId="ref1788" refString="Fanti F., Kupryjanowicz J., 2017. A new soldier beetle from Eocene Baltic amber. Acta Palaeontologica Polonica, 62 (4): 785 - 788." type="journal article" year="2017">Fanti &amp; Kupryjanowicz 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA5D7DFFACBA259B09FD53FD2014EF" blockId="4.[303,813,730,995]" pageId="4" pageNumber="5">
<bibRefCitation id="EFC4208CFFACBA259B09FD53FE1615FE" author="Brancucci M." box="[303,489,730,754]" pageId="4" pageNumber="5" pagination="215 - 388" refId="ref1580" refString="Brancucci M. 1980. Morphologie comparee, evolution et systematique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5: 215 - 388." type="journal article" year="1980">Brancucci (1980)</bibRefCitation>
considered
<taxonomicName id="4C5526FEFFACBA259840FD53FD1515FE" authorityName="Kiesenwetter" authorityYear="1852" box="[614,746,730,754]" pageId="4" pageNumber="1" rank="subFamily" subFamily="Malthininae">Malthininae</taxonomicName>
as the most recently evolved subfamily within
<taxonomicName id="4C5526FEFFACBA259B09FC96FE47143B" authorityName="Imhoff" authorityYear="1856" box="[303,440,799,823]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
, but their subsequent discovery in the Cretaceous Burmese amber (
<bibRefCitation id="EFC4208CFFACBA25985EFCC8FCD71455" author="Hsiao Y. &amp; Slipinski A. &amp; Deng C. &amp; Pang H." box="[632,808,833,858]" pageId="4" pageNumber="5" pagination="119 - 123" refId="ref1820" refString="Hsiao Y., Slipinski A., Deng C., Pang H. 2016. A new genus and species of soldier beetle from Upper Cretaceous Burmese amber (Coleoptera, Cantharidae, Malthininae). Cretaceous Research, 69 (2017): 119 - 123." type="journal article" year="2016">
Hsiao
<emphasis id="B921816FFFACBA25989CFCCAFD111455" box="[698,750,833,858]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2016
</bibRefCitation>
; Fanti 2017), suggested that this subfamily (
<bibRefCitation id="EFC4208CFFACBA2598D5FCEDFE9A1492" author="Fanti F." pageId="4" pageNumber="5" pagination="65 - 68" refId="ref1673" refString="Fanti F. 2019. New fossil Malthodes Kiesenwetter, 1852 of the subgenus Libertimalthodes Kupryjanowicz &amp; Fanti, 2019 from Baltic amber (Coleoptera: Cantharidae). Palaeodiversity, 12: 65 - 68." type="journal article" year="2019">Fanti 2019</bibRefCitation>
;
<bibRefCitation id="EFC4208CFFACBA259B48FC0FFD681492" author="Kupryjanowicz J. &amp; Fanti F." box="[366,663,902,926]" pageId="4" pageNumber="5" refId="ref1910" refString="Kupryjanowicz J., Fanti F. 2019. New subgenus and three new species of soldier beetles from the" type="book" year="2019">Kupryjanowicz &amp; Fanti 2019</bibRefCitation>
; Parisi &amp;
<bibRefCitation id="EFC4208CFFACBA2598D1FC0FFE9814CC" author="Fanti F." pageId="4" pageNumber="5" pagination="65 - 68" refId="ref1673" refString="Fanti F. 2019. New fossil Malthodes Kiesenwetter, 1852 of the subgenus Libertimalthodes Kupryjanowicz &amp; Fanti, 2019 from Baltic amber (Coleoptera: Cantharidae). Palaeodiversity, 12: 65 - 68." type="journal article" year="2019">Fanti 2019</bibRefCitation>
) is however older than assumed, back-dating even more the origin of
<taxonomicName id="4C5526FEFFACBA259875FC42FD2414EF" authorityName="Imhoff" authorityYear="1856" box="[595,731,971,995]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
.
</paragraph>
<paragraph id="8BEA5D7DFFACBA259B09FB99FCE813D8" blockId="4.[303,813,1040,1236]" pageId="4" pageNumber="5">
According to
<bibRefCitation id="EFC4208CFFACBA259BEDFB99FCD71324" author="Fanti F. &amp; Kupryjanowicz J." box="[459,808,1040,1064]" pageId="4" pageNumber="5" pagination="785 - 788" refId="ref1788" refString="Fanti F., Kupryjanowicz J., 2017. A new soldier beetle from Eocene Baltic amber. Acta Palaeontologica Polonica, 62 (4): 785 - 788." type="journal article" year="2017">Fanti &amp; Kupryjanowicz (2017)</bibRefCitation>
,
<taxonomicName id="4C5526FEFFACBA259B09FBBBFE631346" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2017" box="[303,412,1074,1098]" class="Insecta" family="Cantharidae" genus="Kuskaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B921816FFFACBA259B09FBBBFE631346" box="[303,412,1074,1098]" italics="true" pageId="4" pageNumber="5">Kuskaella</emphasis>
</taxonomicName>
could be a specialized genus; however, the elongated elytra and the unspecialized urites make considering this taxon as the basis of the lineage of
<taxonomicName id="4C5526FEFFACBA259BF0FB10FDA613BD" authorityName="Kiesenwetter" authorityYear="1852" box="[470,601,1177,1201]" pageId="4" pageNumber="1" rank="subFamily" subFamily="Malthininae">Malthininae</taxonomicName>
, close to Malchinini, from which it differs in the peculiar aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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