176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document ID-DOI="10.11646/zootaxa.5249.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687190" ID-ZooBank="912A41A4-BB47-40FE-A6EA-A7871C9E5B08" approvalRequired="5" approvalRequired_for_textStreams="5" checkinTime="1677655564240" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2023" docId="03A439097B7BFF83F991FEFE7CFAFABA" docLanguage="en" docName="zootaxa.5249.2.2.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cuspicera Ingrisch 2023" docType="treatment" docVersion="2" lastPageNumber="203" masterDocId="FF9D41717B77FF8EF906FFB57D73FFC3" masterDocTitle="New and little-known genera and species of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) from New Guinea" masterLastPageNumber="212" masterPageNumber="190" pageNumber="202" updateTime="1677656025920" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New and little-known genera and species of Agraeciini (Orthoptera: Tettigoniidae: Conocephalinae) from New Guinea</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingrisch, Sigfrid</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5249</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>190</mods:start>
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<mods:end>212</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.2.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5249.2.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7687190</mods:identifier>
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<mods:identifier type="ZooBank">912A41A4-BB47-40FE-A6EA-A7871C9E5B08</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7687236" ID-Zenodo-Dep="7687236" LSID="urn:lsid:plazi:treatment:03A439097B7BFF83F991FEFE7CFAFABA" httpUri="http://treatment.plazi.org/id/03A439097B7BFF83F991FEFE7CFAFABA" lastPageId="13" lastPageNumber="203" pageId="12" pageNumber="202">
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<subSubSection box="[151,442,330,357]" pageId="12" pageNumber="202" type="nomenclature">
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||
<paragraph blockId="12.[151,442,330,393]" box="[151,442,330,357]" pageId="12" pageNumber="202">
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||
<heading bold="true" box="[151,442,330,357]" fontSize="11" level="1" pageId="12" pageNumber="202" reason="1">
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||
<emphasis bold="true" box="[151,442,330,357]" pageId="12" pageNumber="202">
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||
<emphasis bold="true" box="[151,341,330,357]" italics="true" pageId="12" pageNumber="202">
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<taxonomicName authorityName="Ingrisch" authorityYear="2023" box="[151,273,331,357]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="genus">Cuspicera</taxonomicName>
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alata
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</emphasis>
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sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="203" pageId="12" pageNumber="202" type="description">
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<paragraph blockId="12.[151,442,330,393]" box="[151,217,367,393]" pageId="12" pageNumber="202">
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<figureCitation box="[151,217,367,393]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="12" pageNumber="202">Fig. 6</figureCitation>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7687216" ID-Zenodo-Dep="7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="12" pageNumber="202" startId="12.[151,250,1904,1928]" targetBox="[178,1409,419,1877]" targetPageId="12">
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<paragraph blockId="12.[151,1437,1904,2037]" pageId="12" pageNumber="202">
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<emphasis bold="true" box="[151,274,1904,1928]" pageId="12" pageNumber="202">FIGURE 6.</emphasis>
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<emphasis box="[280,442,1905,1929]" italics="true" pageId="12" pageNumber="202">
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<taxonomicName authorityName="Ingrisch" authorityYear="2023" box="[280,384,1905,1928]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="genus">Cuspicera</taxonomicName>
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alata
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</emphasis>
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<emphasis bold="true" box="[447,527,1905,1929]" pageId="12" pageNumber="202">sp. nov.</emphasis>
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male holotype: A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I).
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</paragraph>
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</caption>
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<paragraph blockId="13.[151,1437,150,1401]" pageId="13" pageNumber="203">
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<materialsCitation collectingDate="1977-04-21" collectionCode="BPBM" collectorName="J. L. Gressitt & Honolulu" country="Papua New Guinea" county="Mt. Karimui" elevation="1500" latitude="-6.502687" location="New" longLatPrecision="1" longitude="144.82265" municipality="Chimbu Province" pageId="13" pageNumber="203" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[151,255,151,177]" pageId="13" pageNumber="203">Holotype</typeStatus>
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(male):
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<collectingCountry box="[345,561,151,177]" name="Papua New Guinea" pageId="13" pageNumber="203">Papua New Guinea</collectingCountry>
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:
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<location LSID="urn:lsid:plazi:treatment:03A439097B7BFF83F991FEFE7CFAFABA:8ED2DEC47B7AFF83FB3FFF227F1DFF72" box="[569,622,151,177]" country="Papua New Guinea" county="Mt. Karimui" latitude="-6.502687" longLatPrecision="1" longitude="144.82265" municipality="Chimbu Province" name="New" pageId="13" pageNumber="203">New</location>
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Guinea (NE),
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<collectingMunicipality box="[779,976,151,177]" pageId="13" pageNumber="203">Chimbu Province</collectingMunicipality>
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,
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<collectingCounty box="[985,1125,151,177]" pageId="13" pageNumber="203">Mt. Karimui</collectingCounty>
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, elev.
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<quantity box="[1191,1273,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="203" unit="m" value="1500.0">
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<elevation box="[1191,1273,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="203" unit="m" value="1500.0">1500 m</elevation>
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</quantity>
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||
(
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<geoCoordinate box="[1285,1432,150,177]" degrees="6.50268693" direction="south" orientation="latitude" pageId="13" pageNumber="203" precision="1" value="-6.502687">-6.50268693</geoCoordinate>
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,
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<geoCoordinate box="[151,274,187,213]" degrees="144.82264" direction="east" orientation="longitude" pageId="13" pageNumber="203" precision="1" value="144.82265">144.82264</geoCoordinate>
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),
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<date box="[294,414,187,213]" pageId="13" pageNumber="203" value="1977-04-21">
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<collectingDate box="[294,414,187,213]" pageId="13" pageNumber="203" value="1977-04-21">21.iv.1977</collectingDate>
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</date>
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, coll.
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<collectorName box="[481,619,187,213]" pageId="13" pageNumber="203">J.L. Gressitt</collectorName>
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—depository:
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<collectorName box="[779,885,187,213]" pageId="13" pageNumber="203">Honolulu</collectorName>
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(
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<collectionCode box="[899,983,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="203" type="Museum">BPBM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="203" type="diagnosis">
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<paragraph blockId="13.[151,1437,150,1401]" pageId="13" pageNumber="203">
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<emphasis bold="true" box="[199,322,222,248]" pageId="13" pageNumber="203">Diagnosis.</emphasis>
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<emphasis box="[329,420,223,249]" italics="true" pageId="13" pageNumber="203">C. alata</emphasis>
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<emphasis bold="true" box="[428,518,223,249]" pageId="13" pageNumber="203">sp. nov.</emphasis>
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differs from
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<taxonomicName authority="Ingrisch, 2023" authorityName="Ingrisch" authorityYear="2023" box="[668,782,223,249]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="luxoria" status="sp. nov.">
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<emphasis box="[668,782,223,249]" italics="true" pageId="13" pageNumber="203">C. luxoria</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[790,880,223,249]" pageId="13" pageNumber="203">
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<taxonomicNameLabel box="[790,880,223,249]" pageId="13" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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by the black vertical band on the face missing in
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<taxonomicName authorityName="Ingrisch" authorityYear="2023" box="[151,266,259,285]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="luxoria">
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<emphasis box="[151,266,259,285]" italics="true" pageId="13" pageNumber="203">C. luxoria</emphasis>
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</taxonomicName>
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; the male cerci are of different shape and provided with internal projections from subapical area and in basal half instead of at end and at base. The titillators have the ventral-apical margins compressed and narrow but with apical area modified instead of widened throughout and simple (in
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<taxonomicName authorityName="Ingrisch" authorityYear="2023" box="[955,1068,331,357]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="luxoria">
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<emphasis box="[955,1068,331,357]" italics="true" pageId="13" pageNumber="203">C. luxoria</emphasis>
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</taxonomicName>
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), the stridulatory file is provided with 183 teeth instead of only 106 teeth and for that has the lateral margin of the tegmen extended by a projecting flap while it is simply rounded in
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<taxonomicName authorityName="Ingrisch" authorityYear="2023" box="[530,644,403,429]" class="Insecta" family="Tettigoniidae" genus="Cuspicera" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="luxoria">
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<emphasis box="[530,644,403,429]" italics="true" pageId="13" pageNumber="203">C. luxoria</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="13" pageNumber="203" type="description">
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||
<paragraph blockId="13.[151,1437,150,1401]" pageId="13" pageNumber="203">
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<emphasis bold="true" box="[199,345,438,464]" pageId="13" pageNumber="203">Description.</emphasis>
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Medium sized species. Pronotum prolonged, with vaulted apical area covering stridulatory area of tegmen; lateral lobes narrow with a projecting ventral lobe in about mid-length (
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<figureCitation box="[1101,1191,475,501]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Fig. 6B</figureCitation>
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). Prosternal spines of medium length; mesosternal lobes subspinous with obtuse tip; metasternal lobes subangularly rounded; hind knees with a single apical spine on both sides. Femora with the following number of spines on ventral margins: fore femur 5 anterior, 5 posterior; mid femur 5 anterior, 2 posterior near base; hind femur 7 anterior over the whole length, without posterior spines. Hind knee lobes with one spine on both sides.
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</paragraph>
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<paragraph blockId="13.[151,1437,150,1401]" pageId="13" pageNumber="203">
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Male. Pronotum lateral lobes with ventral expansion in about mid-length and with dorsal anterior area flat, posterior area swollen. Tegmina almost completely reduced to stridulatory apparatus, covered by pronotum when at rest; stridulatory file on underside of left tegmen
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||
<quantity box="[739,826,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="203" unit="mm" value="1.2">1.2 mm</quantity>
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long, with small and very densely arranged teeth that count over 180 teeth or about 150 teeth per
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<quantity box="[641,708,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="203" unit="mm" value="1.0">1 mm</quantity>
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||
, and even in the area with the largest teeth count 71 teeth per
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||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="203" unit="mm" value="0.5">0.5 mm</quantity>
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or 14 teeth per
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||
<quantity box="[371,457,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="203" unit="mm" value="0.1">0.1 mm</quantity>
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||
(
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||
<figureCitation box="[472,559,799,825]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Fig. 6E</figureCitation>
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). Tenth abdominal tergite markedly wider than long with concave anterior and posterior margins; disc along midline little subsided but resulting furrow bridged in mid-length and there about as high as laterally (
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<figureCitation box="[379,470,871,897]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Fig. 6H</figureCitation>
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||
). Cerci short conical, just before end with a compressed internal process with convex apical margin and on proximal margin with a semicircular projection from basal area; in or little before mid-length of cercus with another compressed and rounded internal projection (
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<figureCitation box="[943,1025,943,969]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Fig. 6I</figureCitation>
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||
). Subgenital plate central area with approaching lateral margins, lateral margins upcurved and then widened to compressed, convex, lateral rims; central area at base little excised and along midline with indication of a faint carina; apical margin angularly excised (
|
||
<figureCitation captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Fig. 6H</figureCitation>
|
||
), swollen on both sides and downcurved, carrying small styli. Titillators with basal area curved laterad, simple band-shaped but little widened, central area long and narrow, forming a sclerotized, dark band that widens apically and gives rise to a short, about triangular process, while the central area of the band curved laterad and distad and supports a roughly triangular membranous surface that is densely covered with clinging hairs. That structure is in subapical area bent again and forms a roughly long-oval membranous sacculus with rugose surface and narrow end (
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<figureCitation box="[159,285,1231,1257]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[178,1409,419,1877]" captionTargetId="figure-60@12.[178,1409,417,1881]" captionTargetPageId="12" captionText="FIGURE 6. Cuspicera alata sp. nov. male holotype:A, frons, legs and sternites in anterio-ventral view; B, habitus lateral view; C, wings with left tegmen opened; D, left tegmen in top view; E, stridulatory file; F–G, titillators in ventro-apical view (F) and view on apical area (G); H, end of abdomen in dorso-apical view; I, right cercus in dorsal view.—Scales 10 mm (A–B), 1 mm (C–H), 0.5 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7687216" httpUri="https://zenodo.org/record/7687216/files/figure.png" pageId="13" pageNumber="203">Figs 6F–G</figureCitation>
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).
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</paragraph>
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<paragraph blockId="13.[151,1437,150,1401]" box="[199,401,1267,1293]" pageId="13" pageNumber="203">Female unknown.</paragraph>
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<paragraph blockId="13.[151,1437,150,1401]" box="[199,1279,1302,1329]" pageId="13" pageNumber="203">
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Measurements (
|
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<specimenCount box="[381,456,1303,1329]" pageId="13" pageNumber="203" type="male">1 male</specimenCount>
|
||
).—body w/o wings: 22; pronotum: 8.3; tegmen: 4; hind femur:
|
||
<quantity box="[1173,1275,1302,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="13" pageNumber="203" unit="mm" value="11.5">11.5 mm</quantity>
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||
.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="203" type="etymology">
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||
<paragraph blockId="13.[151,1437,150,1401]" pageId="13" pageNumber="203">
|
||
<emphasis bold="true" box="[199,334,1339,1365]" pageId="13" pageNumber="203">Etymology.</emphasis>
|
||
The new species is named for the movable apical projections of the titillators that in ventral-apical view look like wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |