162 lines
13 KiB
XML
162 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7200" ID-GBIF-Dataset="dced855b-268d-4353-8a42-247a7f67b621" ID-PMC="PMC4023237" ID-Pensoft-Pub="1313-2970-403-1" ID-PubMed="24843264" ID-ZBK="87A96A762AA14B02A4439D986DDCAD64" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-1" ModsDocOrigin="ZooKeys 403" ModsDocTitle="Three new species of the genus Scaphicoma Motschulsky, 1863 (Coleoptera, Staphylinidae, Scaphidiinae) from Northern Sulawesi, Indonesia" checkinTime="1451245968335" checkinUser="pensoft" docAuthor="Ogawa, Ryo, Loebl, Ivan & Maeto, Kaoru" docDate="2014" docId="E5E6385EECA5FAC9F657A4B5C77449CB" docLanguage="en" docName="ZooKeys 403: 1-13" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7200" docTitle="Scaphicoma subflava Ogawa & Loebl, sp. n." docType="treatment" docUuid="5EF0186B-9ECA-4998-805B-BC38A0450214" docUuidSource="ZooBank" docVersion="4" lastPageNumber="6" masterDocId="FFFBFF87855AFFC0D601FF89FF85FFAF" masterDocTitle="Three new species of the genus Scaphicoma Motschulsky, 1863 (Coleoptera, Staphylinidae, Scaphidiinae) from Northern Sulawesi, Indonesia" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1668158242690" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Three new species of the genus Scaphicoma Motschulsky, 1863 (Coleoptera, Staphylinidae, Scaphidiinae) from Northern Sulawesi, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ogawa, Ryo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Loebl, Ivan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maeto, Kaoru</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>403</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>13</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.403.7200</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7200</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-403-1</mods:identifier>
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<mods:identifier type="ZBK">87A96A762AA14B02A4439D986DDCAD64</mods:identifier>
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<mods:identifier type="ZooBank">87A96A762AA14B02A4439D986DDCAD64</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052602" LSID="urn:lsid:zoobank.org:act:5EF0186B-9ECA-4998-805B-BC38A0450214" httpUri="http://treatment.plazi.org/id/E5E6385EECA5FAC9F657A4B5C77449CB" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName LSID="http://zoobank.org/5EF0186B-9ECA-4998-805B-BC38A0450214" authority="Ogawa & Loebl" class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma subflava" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="subflava">
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Scaphicoma subflava Ogawa &
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<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
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Figs 1a, 2, 5
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="diagnosis">
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<paragraph pageId="2" pageNumber="3">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Most of body yellowish-brown. Body size relatively moderate. Antennomere XI about 1.6 times as long as VIII; IV and V each shorter than VI. Protarsomeres
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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and V each about two times as long as IV. Mesotarsomeres I about 1.2 times as long as II and III; V about 1.5 times as long as IV. Metatarsomeres I about 1.5 times as long as II and III; II and III each about 1.5 times as long as IV and V. Male sternite VII with middle of apical margin strongly concave. Parameres asymmetrical. Internal sac on basal portion covered with scale-like sclerites, and with pair of sclerites on apical portion. Bursa copulatrix sclerotized.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
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<paragraph pageId="2" pageNumber="3">Description.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Body, shining. Most of body including head, pronotum and elytra yellowish-brown, except for darkened mesoventrite (Fig. 1a). Antennae yellowish-brown, except of antennomeres
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<normalizedToken originalValue="VII–XI">VII-XI</normalizedToken>
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dark yellowish-brown. Head, pronotum and elytra sparsely and finely pubescent.
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Head with interocular distance almost as eye width. Punctuation sparse and fine. Antennomeres
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<normalizedToken originalValue="I–VI">I-VI</normalizedToken>
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with a few macrosetae,
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<normalizedToken originalValue="VII–XI">VII-XI</normalizedToken>
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covered with some microsetae; VI about two times as long as III; IV and V each shorter than VI; VII almost as VIII; XI about 1.6 times as long as VIII (Fig. 2e).
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</paragraph>
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<caption pageId="2" pageNumber="3">
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<paragraph pageId="2" pageNumber="3">
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Figure 1. Habitus of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Scaphicoma</taxonomicName>
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spp. in dorsal and lateral views. a
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma subflava" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="subflava">Scaphicoma subflava</taxonomicName>
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sp. n. b
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma bidentia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bidentia">Scaphicoma bidentia</taxonomicName>
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sp. n. c
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma quadrifasciata" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="quadrifasciata">Scaphicoma quadrifasciata</taxonomicName>
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sp. n. Scale: 1.0 mm.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="3">
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<paragraph pageId="2" pageNumber="3">
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Figure 2. Genitalia and antenna of
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma subflava" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="subflava">Scaphicoma subflava</taxonomicName>
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sp. n. a, b Male genitalia; c, d female genitalia e male antenna. a, c, Dorsal view; b, d, lateral view. Scale: 0.25 mm.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="4">
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<pageBreakToken pageId="3" pageNumber="4" start="start">Pronotum</pageBreakToken>
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almost as wide as long, lateral keel invisible in dorsal view. Punctuation sparse and fine, as on head. Scutellum concealed.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">Elytra longer than wide, widest at basal 1/6, gradually narrowed to apex, with minute serration at inner part of posterior margin. Punctuation coarser than on pronotum (Fig. 5e, f). Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and not joined with lateral striae.</paragraph>
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<paragraph pageId="3" pageNumber="4">Propygidium densely and finely punctuate. Pygidium sparsely and finely punctuate, slightly emarginated at apex.</paragraph>
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<paragraph pageId="3" pageNumber="4">Hypomeron finely punctuate. Lateral portion of mesoventrite coarsely and sparsely punctuate; medial portion finely and sparsely punctuate, with fine pubescence. Lateral portion of metaventrite from base to basal 1/3 sparsely and coarsely punctuate, with apical portion moderately concave. Mesocoxa almost six times as wide as space between them; mesocoxal area moderately broadened. Metepimeron almost as long as wide, with microsculpture. Metacoxa about eight times as wide as metacoxal process. Metanepisternum about six times as long as wide. Lateral portion of ventrite I from base to basal 1/3 densely and coarsely punctuate. Ventrite VI strongly pointed at apicomedian portion.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Meso- and metafemora with microsculpture, sparsely and coarsely punctuate. Protarsomeres
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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and V each about two times as long as IV. Mesotarsomeres I about 1.2 times as long as II and III; V about 1.5 times as long as IV. Metatarsomeres I about 1.5 times as long as II and III; II and III each about 1. 5 times as long as IV and V.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Male. Ventrite V strongly emarginated at apex. Protarsomeres
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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with tenent setae (Fig. 5b), enlarged. Aedeagus about 0.91 mm long; parameres asymmetrical; internal sac on basal portion covered with scale-like sclerites, and with a pair of sclerites (Fig. 2a).
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Female</pageBreakToken>
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. Ventrite V slightly emarginate or truncate. Protarsomeres
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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without tenent setae, not enlarged. Gonostylus elongate. Distal gonocoxite normal and elongate; vagina membranous, without robust sclerites; bursa copulatrix strongly sclerotized (Fig. 2c, d). Spermatheca as Fig. 2c.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="measurements">
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<paragraph pageId="4" pageNumber="5">Measurements</paragraph>
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<paragraph pageId="4" pageNumber="5">(n = 6). Length (PL+EL): 2.55-2.75 mm; width (PW, EW): 1.04-1.13 mm, 1.09-1.21 mm. HW: 0.55-0.58 mm. ID: 0.16-0.19 mm. PL/PW: 0.95-1.07. EL/EW:1.30-1.43. Approximate ratio of each antennal length (width) from base to apex as follows (n = 1): 1.7 (0.7): 0.9 (0.6): 1.0 (0.2): 1.6 (0.2): 1.5 (0.2): 1.9 (0.2): 1.5 (0.3): 1.5 (0.2): 1.7 (0.3): 1.9 (0.3): 2.5 (0.2).</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="specimens examined">
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<paragraph pageId="4" pageNumber="5">Specimens examined.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Holotype, 1 ♂, Mt. Tilongkabila (Gunung Tilongkabila), N. Sulawesi, alt. ca. 800m, 0°34'
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<geoCoordinate direction="north" orientation="latitude" precision="555" value="28.52">28.52N</geoCoordinate>
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, 123°11'
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<geoCoordinate direction="east" orientation="longitude" precision="555" value="30.61">30.61E</geoCoordinate>
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, 8. VI. 2012, R. Ogawa leg. (MZBI); Paratypes, 2♂2♀, Same data above (EUMJ); 1♂, Mt. Tilongkabila (Gunung Tilongkabila), N. Sulawesi, alt. ca. 800-1300 m, 0°34'28.52N, 0°35'18.14N, 123°11'30.61E, 123°13'22.71E, 9. VI. 2012, R. Ogawa leg. (HUOI); 1♂, Palu, Palopo, C. Sulawesi, 25-27. VIII. 1990, A. Riedel leg. (MHNG).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Indonesia: northern and central Sulawesi.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="etymology">
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<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="6">This specific name is the Latin subflava adjective meaning somewhat yellowish.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="remarks">
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<paragraph pageId="5" pageNumber="6">Remarks.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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This species was illustrated in
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<bibRefCitation author="Leschen, RAB" journalOrPublisher="Coleopterists Society Monographs" pageId="11" pageNumber="12" pagination="1 - 63" title="Phylogeny and classification of Scaphisomatini (Staphylinidae: Scaphidiinae) with notes on mycophagy, termitophily and functional morphology." url="10.1649/0010-065X(2005)059[0001:PACOSS]2.0.CO;2" volume="3" year="2005">Leschen and Lobl (2005)</bibRefCitation>
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, thought unidentified. It is very similar to the Javanese
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphicoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphicoma pallens" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pallens">Scaphicoma pallens</taxonomicName>
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(Achard, 1921) by the body color and the shapes of male genitalia, but it is easily distinguished by the distinctive male genitalia with internal sac bearing sclerites.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |