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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1143.94951" ID-Pensoft-Pub="1313-2970-1143-93" ID-Pensoft-UUID="5E978E6177BE5B39B380B105500D7DA5" ID-ZooBank="9156C6A84BF5472FA7010C2F089CE134" ModsDocID="1313-2970-1143-93" checkinTime="1675260478037" checkinUser="pensoft" docAuthor="van Achterberg, Cornelis, Smit, John T. &amp; Ljubomirov, Toshko" docDate="2023" docId="AACD0F7042E2572C88D3FCCEEA692360" docLanguage="en" docName="ZooKeys 1143: 93-163" docOrigin="ZooKeys 1143" docPubDate="2023-01-31" docSource="http://dx.doi.org/10.3897/zookeys.1143.94951" docTitle="Eumenes subpomiformis Bluethgen 1938" docType="treatment" docVersion="1" id="5E978E6177BE5B39B380B105500D7DA5" lastPageNumber="93" masterDocId="5E978E6177BE5B39B380B105500D7DA5" masterDocTitle="Review of the European Eumenes Latreille (Hymenoptera, Vespidae) using morphology and DNA barcodes, with an illustrated key to species" masterLastPageNumber="163" masterPageNumber="93" pageNumber="93" updateTime="1675260478037" updateUser="pensoft">
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<mods:title>Review of the European Eumenes Latreille (Hymenoptera, Vespidae) using morphology and DNA barcodes, with an illustrated key to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Achterberg, Cornelis</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, P. O. 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">kees@vanachterberg.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smit, John T.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1568-5183</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, P. O. 9517, 2300 RA Leiden, Netherlands &amp; European Invertebrate Survey - Netherlands, P. O. 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ljubomirov, Toshko</mods:namePart>
<mods:affiliation>Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Tzar Osvoboditel Boulevard 1, 1000 Sofia, Bulgaria</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1143</mods:number>
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<mods:start>93</mods:start>
<mods:end>163</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1143.94951</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1143.94951</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:AACD0F7042E2572C88D3FCCEEA692360" httpUri="http://treatment.plazi.org/id/AACD0F7042E2572C88D3FCCEEA692360" lastPageNumber="93" pageId="0" pageNumber="93">
<subSubSection pageId="0" pageNumber="93" type="nomenclature">
<paragraph pageId="0" pageNumber="93">
<taxonomicName ID-CoL="3CDPJ" ID-ENA="2910884" LSID="AACD0F70-42E2-572C-88D3-FCCEEA692360" authority="Bluethgen, 1938" authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">
Eumenes subpomiformis
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1938
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="description">
<paragraph pageId="0" pageNumber="93">
<figureCitation captionStart="Figures 227–235" captionStartId="F28" captionText="Figures 227 - 235. Eumenes subpomiformis Bluethgen, Bulgaria, female 227 metasoma lateral 228 first metasomal tergite dorsal 229 first tergite ventral 230 mesosoma dorsal 231 second metasomal tergite latero-dorsal 232 head anterior 233 head and propleuron lateral 234 hind tarsal claw 235 antenna." figureDoi="10.3897/zookeys.1143.94951.figures227-235" httpUri="https://binary.pensoft.net/fig/803451" pageId="0" pageNumber="93">Figs 227-235</figureCitation>
<figureCitation captionStart="Figures 236–243" captionStartId="F29" captionText="Figures 236 - 243. Eumenes subpomiformis Bluethgen, Bulgaria, male 236 metasoma lateral 237 first metasomal tergite dorsal 238 first tergite ventral 239 head and mesosoma dorsal 240 propodeum dorsal 241 head anterior 242 head and mesosoma lateral 243 apical hook of antenna lateral." figureDoi="10.3897/zookeys.1143.94951.figures236-243" httpUri="https://binary.pensoft.net/fig/803452" pageId="0" pageNumber="93">, 236-243</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="reference_group">
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">Eumenes subpomiformis</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1938: 480, 496;
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 101-103,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="561 - 584" refId="B52" refString="Gusenleitner, J, 1999. Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855. Linzer Biologische Beitrage 31 (2): 561 - 584" title="Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855." volume="31" year="1999">1999</bibRefCitation>
: 574,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="5 - 107" refId="B53" refString="Gusenleitner, J, 2013. Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae). Linzer Biologische Beitrage 45 (1): 5 - 107" title="Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae)." volume="45" year="2013">2013</bibRefCitation>
: 29;
<bibRefCitation author="Tobias, VI" editor="Medvedev, G" journalOrPublisher="Hymenoptera 3 (1)" pageId="0" pageNumber="93" pagination="1 - 584" refId="B82" refString="Tobias, VI, Kurzenko, NV, 1978. Sem. 2. Eumenidae, 152-173. In: Medvedev, G, Ed., Keys to the identification of insects of European USSR. Hymenoptera 3(1): 1 - 584" title="Sem. 2. Eumenidae, 152 - 173." volumeTitle="Keys to the identification of insects of European USSR." year="1978">Tobias and Kurzenko 1978</bibRefCitation>
: 161;
<bibRefCitation author="Castro, L" journalOrPublisher="Catalogus de la entomofauna aragonesa" pageId="0" pageNumber="93" pagination="3 - 8" refId="B18" refString="Castro, L, 1997. Familia Vespidae: Subfamilia Eumeninae. Catalogus de la entomofauna aragonesa 16: 3 - 8" title="Familia Vespidae: Subfamilia Eumeninae." volume="16" year="1997">Castro 1997</bibRefCitation>
: 4;
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Bembix" pageId="0" pageNumber="93" pagination="13 - 28" refId="B78" refString="Schmid-Egger, C, Schmidt, K, 2002. Kritisches Verzeichnis der deutschen Arten der Faltenwespen (Hymenoptera, Vespidae). Bembix 16: 13 - 28" title="Kritisches Verzeichnis der deutschen Arten der Faltenwespen (Hymenoptera, Vespidae)." volume="16" year="2002">Schmid-Egger and Schmidt 2002</bibRefCitation>
: 18;
<bibRefCitation author="Woydak, H-G" journalOrPublisher="Abhandlungen aus dem Westfalischen Museum fur Naturkunde" pageId="0" pageNumber="93" pagination="1 - 133" refId="B86" refString="Woydak, H-G, 2006. Hymenoptera Aculeata Westfalica. Die Faltenwespen von Nordrhein-Westfalen (Hymenoptera, Vespoidea; Vespidae und Eumenidae) (Soziale Papier und Lehmwespen). Abhandlungen aus dem Westfalischen Museum fur Naturkunde 68 (1): 1 - 133" title="Hymenoptera Aculeata Westfalica. Die Faltenwespen von Nordrhein-Westfalen (Hymenoptera, Vespoidea; Vespidae und Eumenidae) (Soziale Papier und Lehmwespen)." volume="68" year="2006">Woydak 2006</bibRefCitation>
: 46-47;
<bibRefCitation author="Castro, L" journalOrPublisher="Boletin de la SEA" pageId="0" pageNumber="93" pagination="259 - 278" refId="B19" refString="Castro, L, Sanza, F, 2009. Aportacion al conocimiento de los Vespidae (Hymenoptera) de Sierra Nevada (Espana), con algunos comentarios taxonomicos. Boletin de la SEA 45: 259 - 278" title="Aportacion al conocimiento de los Vespidae (Hymenoptera) de Sierra Nevada (Espana), con algunos comentarios taxonomicos." volume="45" year="2009">Castro and Sanza 2009</bibRefCitation>
: 267;
<bibRefCitation author="Arens, W" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="481 - 522" refId="B5" refString="Arens, W, 2012. Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil. Linzer Biologische Beitrage 44 (1): 481 - 522" title="Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil." volume="44" year="2012">Arens 2012</bibRefCitation>
: 489;
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" pagination="5 - 40" refId="B76" refString="Schmid-Egger, C, 2010. Rote Liste der Wespen Deutschlands. Hymenoptera Aculeata: Grabwespen (Ampulicidae, Crabronidae, Sphecidae), Wegwespen (Pompilidae), Goldwespen (Chrysididae), Faltenwespen (Vespidae), Spinnenameisen (Mutillidae), Dolchwespen (Scoliidae), Rollwespen (Tiphiidae) und Keulhornwespen (Sapygidae). Ampulex 1: 5 - 40" title="Rote Liste der Wespen Deutschlands. Hymenoptera Aculeata: Grabwespen (Ampulicidae, Crabronidae, Sphecidae), Wegwespen (Pompilidae), Goldwespen (Chrysididae), Faltenwespen (Vespidae), Spinnenameisen (Mutillidae), Dolchwespen (Scoliidae), Rollwespen (Tiphiidae) und Keulhornwespen (Sapygidae)." volume="1" year="2010">Schmid-Egger 2010</bibRefCitation>
: 23,
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" pagination="13 - 50" refId="B77" refString="Schmid-Egger, C, 2011. Hymenoptera Aculeata from &quot;Parc national du Mercantour&quot; (France) and &quot;Parco delle Alpi Marittime&quot; (Italy) in the south-western Alps. Ampulex 3: 13 - 50" title="Hymenoptera Aculeata from &quot; Parc national du Mercantour &quot; (France) and &quot; Parco delle Alpi Marittime &quot; (Italy) in the south-western Alps." volume="3" year="2011">2011</bibRefCitation>
: 44;
<bibRefCitation author="Neumeyer, R" journalOrPublisher="Mitteilungen der Schweizerischen entomologischen Gesellschaft" pageId="0" pageNumber="93" pagination="359 - 369" refId="B68" refString="Neumeyer, R, 2014. Kommentierte liste der in der Schweiz vorkommenden Faltenwespen (Hymenoptera: Vespidae). Mitteilungen der Schweizerischen entomologischen Gesellschaft 87: 359 - 369" title="Kommentierte liste der in der Schweiz vorkommenden Faltenwespen (Hymenoptera: Vespidae)." volume="87" year="2014">Neumeyer 2014</bibRefCitation>
: 367,
<bibRefCitation author="Neumeyer, R" journalOrPublisher="Fauna Helvetica" pageId="0" pageNumber="93" pagination="1 - 381" refId="B69" refString="Neumeyer, R, 2019. Vespidae. Fauna Helvetica 31: 1 - 381" title="Vespidae." volume="31" year="2019">2019</bibRefCitation>
: 276;
<bibRefCitation author="Baldock, D" journalOrPublisher="Monografias Sociedad Entomologiva Aragonesa" pageId="0" pageNumber="93" pagination="1 - 73" refId="B6" refString="Baldock, D, Castro, L, Cross, I, Schmid-Egger, C, Smit, J, Wood, TJ, 2020. The vespiform wasps of Portugal (Hymenoptera: Scolioidea, Tiphioidea, Pompiloidea &amp; Vespoidea). Monografias Sociedad Entomologiva Aragonesa 34: 1 - 73" title="The vespiform wasps of Portugal (Hymenoptera: Scolioidea, Tiphioidea, Pompiloidea &amp; Vespoidea)." volume="34" year="2020">Baldock et al. 2020</bibRefCitation>
: 44.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis" subGenus="Eumenes">Eumenes (Eumenes) subpomiformis</taxonomicName>
;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 133-134 (literature before 1972);
<bibRefCitation author="Verges Serra, F" journalOrPublisher="Graellsia" pageId="0" pageNumber="93" pagination="137 - 152" refId="B85" refString="Verges Serra, F, 1985. Estudio monografico de los Eumenes de Espana (Hymenoptera, Eumenidae). Graellsia 41: 137 - 152" title="Estudio monografico de los Eumenes de Espana (Hymenoptera, Eumenidae)." volume="41" year="1985">
<normalizedToken originalValue="Vergés">Verges</normalizedToken>
Serra 1985
</bibRefCitation>
: 147;
<bibRefCitation author="Castro, L" journalOrPublisher="Zapateri, Revista aragonesa de entomologia" pageId="0" pageNumber="93" pagination="21 - 34" refId="B17" refString="Castro, L, 1992. Sobre los eumenidos (Hym., Vespoidea) del valle medio del Ebro. Zapateri, Revista aragonesa de entomologia 2: 21 - 34" title="Sobre los eumenidos (Hym., Vespoidea) del valle medio del Ebro." volume="2" year="1992">Castro 1992</bibRefCitation>
: 25;
<bibRefCitation author="Sanza, F" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" refId="B74" refString="Sanza, F, 1997. Revision de los eumenidos de la Peninsula Iberica (Hymenoptera: Eumenidae) (tesis doctoral, inedita). Unpublished thesis, Dept de Biologia Animal, Ecologia, Parasitologia, Edafologia y Quimica Agricola de la Universidad de Salamanca, Salamanca, i-vii + 1-541." title="Revision de los eumenidos de la Peninsula Iberica (Hymenoptera: Eumenidae) (tesis doctoral, inedita). Unpublished thesis, Dept de Biologia Animal, Ecologia, Parasitologia, Edafologia y Quimica Agricola de la Universidad de Salamanca, Salamanca, i-vii + 1 - 541." year="1997">Sanza 1997</bibRefCitation>
: 463;
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" refId="B75" refString="Schmid-Egger, C, 2004. Bestimmungsschluessel fuer die deutschen Arten der solitaeren Faltenwespen (Hymenoptera: Eumeninae). Deutscher Jugendbund fuer Naturbeobachtung Hamburg, 54-102. 3. ueberarbeitete Auflage. Hamburg." title="Bestimmungsschluessel fuer die deutschen Arten der solitaeren Faltenwespen (Hymenoptera: Eumeninae). Deutscher Jugendbund fuer Naturbeobachtung Hamburg, 54 - 102. 3. ueberarbeitete Auflage. Hamburg." year="2004">Schmid-Egger 2004</bibRefCitation>
: 73;
<bibRefCitation DOI="https://doi.org/10.3406/linly.2006.13644" author="Gereys, B" journalOrPublisher="Bulletin de la Societe Linneenne de Lyon" pageId="0" pageNumber="93" pagination="383 - 392" refId="B37" refString="Gereys, B, 2006. Mise a jour de la nomenclature des Vespidae (Hymenoptera) de France metropolitaine et premier inventaire du departement des Alpes-de-Haute-Provence (France) (Deuxieme partie). Bulletin de la Societe Linneenne de Lyon 75 (10): 383 - 392, DOI: https://doi.org/10.3406/linly.2006.13644" title="Mise a jour de la nomenclature des Vespidae (Hymenoptera) de France metropolitaine et premier inventaire du departement des Alpes-de-Haute-Provence (France) (Deuxieme partie)." url="https://doi.org/10.3406/linly.2006.13644" volume="75" year="2006">Gereys 2006</bibRefCitation>
: 387,
<bibRefCitation author="Gereys, B" journalOrPublisher="Faune de France" pageId="0" pageNumber="93" pagination="1 - 330" refId="B40" refString="Gereys, B, 2016. Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae). Faune de France 98: 1 - 330" title="Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae)." volume="98" year="2016">2016</bibRefCitation>
: 137;
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5137.1.1" author="Dal Pos, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="93" pagination="1 - 111" refId="B21" refString="Dal Pos, D, Carpenter, JM, Uliana, M, 2022. The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae). Zootaxa 5137 (1): 1 - 111, DOI: https://doi.org/10.11646/zootaxa.5137.1.1" title="The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae)." url="https://doi.org/10.11646/zootaxa.5137.1.1" volume="5137" year="2022">Dal Pos et al. 2022</bibRefCitation>
: 16.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authority="subpomiformis" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis subsp. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="subpomiformis" subSpecies="subpomiformis">Eumenes subpomiformis subpomiformis</taxonomicName>
;
<bibRefCitation author="Giordani Soika, A" journalOrPublisher="In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna Italiana" pageId="0" pageNumber="93" pagination="1 - 9" refId="B47" refString="Giordani Soika, A, Borsato, W, 1995. Hymenoptera Vespoidea. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna Italiana 103: 1 - 9" title="Hymenoptera Vespoidea." volume="103" year="1995">Giordani Soika and Borsato 1995</bibRefCitation>
: 7;
<bibRefCitation author="Borsato, W" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Venezia" pageId="0" pageNumber="93" pagination="127 - 150" refId="B14" refString="Borsato, W, Turrisi, GF, 2004. Contributo alla conoscenza degli Eumenidae di Sicilia (Hymenoptera, Vespoidea). Bollettino del Museo Civico di Storia Naturale di Venezia 55: 127 - 150" title="Contributo alla conoscenza degli Eumenidae di Sicilia (Hymenoptera, Vespoidea)." volume="55" year="2004">Borsato and Turrisi 2004</bibRefCitation>
: 145.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluthgen" authorityYear="1956" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis subsp. crassipunctatus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="subpomiformis" subSpecies="crassipunctatus">Eumenes subpomiformis crassipunctatus</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1956: 3;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 133 (literature before 1972);
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 101-103 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">E. subpomiformis</taxonomicName>
):
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. sareptanus" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="sareptanus">E. sareptanus</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="notes">
<paragraph pageId="0" pageNumber="93">Notes.</paragraph>
<paragraph pageId="0" pageNumber="93">
As pointed out by
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner (1972)</bibRefCitation>
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. subpomiformis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. pomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="pomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. pomiformis</emphasis>
</taxonomicName>
(&quot;
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes pomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="pomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes pomiformis</emphasis>
</taxonomicName>
steht der Art
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes subpomiformis</emphasis>
</taxonomicName>
sehr nahe und nicht der Art
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes lunulatus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="lunulatus">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes lunulatus</emphasis>
</taxonomicName>
&quot;) and is easily misidentified when the medium-sized or long setae of the propleuron of
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. subpomiformis</emphasis>
</taxonomicName>
are not well exposed (head too much down), depressed or damaged. He also correctly denounced the differences in shape of the clypeus as illustrated by
<bibRefCitation DOI="https://doi.org/10.1002/mmnd.48019380208" author="Bluethgen, P" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="93" pagination="434 - 496" refId="B7" refString="Bluethgen, P, 1938. Beitraege zur Kenntnis der palaearktischen Eumeniden (Hym. Vespidae). Deutsche Entomologische Zeitschrift 1938: 434 - 496, DOI: https://doi.org/10.1002/mmnd.48019380208" title="Beitraege zur Kenntnis der palaearktischen Eumeniden (Hym. Vespidae)." url="https://doi.org/10.1002/mmnd.48019380208" volume="1938" year="1938">
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
(1938)
</bibRefCitation>
(&quot;Die Form des Clypeus, wie sie
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
<normalizedToken originalValue="für">fuer</normalizedToken>
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes subpomiformis</emphasis>
</taxonomicName>
angibt (Ausschnittecken nach den Seiten gezogen) tritt auch bei
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes pomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="pomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes pomiformis</emphasis>
</taxonomicName>
auf&quot;.) What remains in both sexes for separation according to the keys by
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner (1972</bibRefCitation>
,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="561 - 584" refId="B52" refString="Gusenleitner, J, 1999. Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855. Linzer Biologische Beitrage 31 (2): 561 - 584" title="Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855." volume="31" year="1999">1999</bibRefCitation>
) is the length of the setae on the propleuron (with equal shorter setae in
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. pomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="pomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. pomiformis</emphasis>
</taxonomicName>
and with unequal longer setae in
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. subpomiformis</emphasis>
</taxonomicName>
). However, the setosity seems rather variable (especially in males) and should be used in combination with other characters. Recent molecular research (
<bibRefCitation author="Neumeyer, R" journalOrPublisher="Mitteilungen der Schweizerischen entomologischen Gesellschaft" pageId="0" pageNumber="93" pagination="307 - 320" refId="B70" refString="Neumeyer, R, Praz, C, 2015. DNA barcoding of the potter wasp genus Eumenes Latreille, 1802 (Hymenoptera: Vespidae: Eumeninae) in Switzerland. Mitteilungen der Schweizerischen entomologischen Gesellschaft 88: 307 - 320" title="DNA barcoding of the potter wasp genus Eumenes Latreille, 1802 (Hymenoptera: Vespidae: Eumeninae) in Switzerland." volume="88" year="2015">Neumeyer and Praz 2015</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1062.70763" author="Schmid-Egger, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="93" pagination="49 - 72" refId="B79" refString="Schmid-Egger, C, Schmidt, S, 2021. Unexpected diversity in Central European Vespoidea (Hymenoptera, Mutillidae, Myrmosidae, Sapygidae, Scoliidae, Tiphiidae, Thynnidae, Vespidae), with description of two species of Smicromyrme Thomson, 1870. ZooKeys 1062: 49 - 72, DOI: https://doi.org/10.3897/zookeys.1062.70763" title="Unexpected diversity in Central European Vespoidea (Hymenoptera, Mutillidae, Myrmosidae, Sapygidae, Scoliidae, Tiphiidae, Thynnidae, Vespidae), with description of two species of Smicromyrme Thomson, 1870." url="https://doi.org/10.3897/zookeys.1062.70763" volume="1062" year="2021">Schmid-Egger and Schmidt 2021</bibRefCitation>
; this paper) revealed distinct genetic differences between
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. subpomiformis</emphasis>
</taxonomicName>
and
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. pomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="pomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. pomiformis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neighbour-Joining tree for barcoded European Eumenes species. Both E. coarctatus specimens (sampled 5 y ago for another project and not available for examination) among E. coronatus are obviously misidentified and, therefore, in quotation marks. The numbers are RMNH unique identifiers except the Finnish numbers of both dark specimens of E. c. coarctatus from Finland." figureDoi="10.3897/zookeys.1143.94951.figure3" httpUri="https://binary.pensoft.net/fig/803427" pageId="0" pageNumber="93">3</figureCitation>
) despite their overall similarity.
</paragraph>
<paragraph pageId="0" pageNumber="93">
Specimens in RMNH identified by
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
(in 1950 and 1955) as
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. pomiformis subsp. barbatulus" order="Hymenoptera" pageId="0" pageNumber="93" rank="subSpecies" species="pomiformis" subSpecies="barbatulus">
<emphasis italics="true" pageId="0" pageNumber="93">E. pomiformis barbatulus</emphasis>
</taxonomicName>
belong either to
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">E. subpomiformis</emphasis>
</taxonomicName>
(Portugal; females with mostly comparatively short setae on propleuron and deeply emarginate clypeus) or to
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. coarctatus" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="coarctatus">
<emphasis italics="true" pageId="0" pageNumber="93">E. coarctatus</emphasis>
</taxonomicName>
(most specimens (with medium-sized to long setae on propleuron) from Portugal, Spain, France, Algeria, Morocco).
</paragraph>
<caption doi="10.3897/zookeys.1143.94951.figures227-235" httpUri="https://binary.pensoft.net/fig/803451" pageId="0" pageNumber="93" start="Figures 227–235" startId="F28">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figures 227-235.</emphasis>
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes subpomiformis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, Bulgaria, female
<emphasis bold="true" pageId="0" pageNumber="93">227</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">228</emphasis>
first metasomal tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="93">229</emphasis>
first tergite ventral
<emphasis bold="true" pageId="0" pageNumber="93">230</emphasis>
mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="93">231</emphasis>
second metasomal tergite latero-dorsal
<emphasis bold="true" pageId="0" pageNumber="93">232</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="93">233</emphasis>
head and propleuron lateral
<emphasis bold="true" pageId="0" pageNumber="93">234</emphasis>
hind tarsal claw
<emphasis bold="true" pageId="0" pageNumber="93">235</emphasis>
antenna.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1143.94951.figures236-243" httpUri="https://binary.pensoft.net/fig/803452" pageId="0" pageNumber="93" start="Figures 236–243" startId="F29">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figures 236-243.</emphasis>
<taxonomicName authorityName="Bluethgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes subpomiformis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="subpomiformis">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes subpomiformis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, Bulgaria, male
<emphasis bold="true" pageId="0" pageNumber="93">236</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">237</emphasis>
first metasomal tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="93">238</emphasis>
first tergite ventral
<emphasis bold="true" pageId="0" pageNumber="93">239</emphasis>
head and mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="93">240</emphasis>
propodeum dorsal
<emphasis bold="true" pageId="0" pageNumber="93">241</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="93">242</emphasis>
head and mesosoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">243</emphasis>
apical hook of antenna lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="distribution">
<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="93">
C and S Europe, but unknown from Sardinia (
<bibRefCitation author="Giordani Soika, A" journalOrPublisher="In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna Italiana" pageId="0" pageNumber="93" pagination="1 - 9" refId="B47" refString="Giordani Soika, A, Borsato, W, 1995. Hymenoptera Vespoidea. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna Italiana 103: 1 - 9" title="Hymenoptera Vespoidea." volume="103" year="1995">Giordani Soika and Borsato 1995</bibRefCitation>
); outside Europe known from Morocco, Israel, Lebanon, and Asia Minor. In Switzerland found up to 1920 m altitude (
<bibRefCitation author="Neumeyer, R" journalOrPublisher="Fauna Helvetica" pageId="0" pageNumber="93" pagination="1 - 381" refId="B69" refString="Neumeyer, R, 2019. Vespidae. Fauna Helvetica 31: 1 - 381" title="Vespidae." volume="31" year="2019">Neumeyer 2019</bibRefCitation>
) as in Greece (
<bibRefCitation author="Arens, W" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="481 - 522" refId="B5" refString="Arens, W, 2012. Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil. Linzer Biologische Beitrage 44 (1): 481 - 522" title="Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil." volume="44" year="2012">Arens 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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