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<document ID-CLB-Dataset="55738" ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-GBIF-Dataset="8b26ea49-e662-41f7-a213-47f4b598c91c" ID-PMC="PMC9848896" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-PubMed="36761078" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="A11627C74A08512DAFE3B8632BA87703" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus mahafaly Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="68F229F9-571D-4B71-8F98-0B89218D660E" docUuidSource="ZooBank" docVersion="5" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1732660529870" updateUser="ExternalLinkService">
<mods:mods id="390F7AB17578D27552B52D5D57845747" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1A5377F88EABEB0476E91D0B934596CD">
<mods:title id="DE1C4A780FEDCB929113D5205BCBE743">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
</mods:titleInfo>
<mods:name id="4E7522C918598A309A3AA9213343468C" type="personal">
<mods:role id="D3B97192B479F0CC3709E3F4CCD6E3B4">
<mods:roleTerm id="D5DBCA592AD634245E9BA64C035F09C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D410E857C448F0AFDD42C5828804F6EF">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="225268440D2A26AFE0503B03CC706B47">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier id="F173765663AE14FC8158E499B78204E8" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="63C1A3D726D68040F85F4E1EE6ADD133" type="personal">
<mods:role id="F56B541100F42AAAB5F3FD78996AD563">
<mods:roleTerm id="6D29B4789797FE5D273741E3E764B2D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEDD00476256B3114F36D7DF8D54F595">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="072BE40C88DF85BCB22ACB5A6FA70E42" type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation id="6C3B31BC7034EF32487FE1DDED63721C">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3D29BA620E02250550B87EDD40F1DEF4">text</mods:typeOfResource>
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<mods:titleInfo id="71FB1AA6A814832C06B2EAF5FCB79633">
<mods:title id="642011D4A83447CA28BB4AC44D6EBE59">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="4937DBD0C281EA80082A5A7333F4D66E">
<mods:date id="D2554963986A0C428513B585CBEBD7BA">2022</mods:date>
<mods:detail id="B20BA51CBD51614FC44496D4ABA3F277" type="pubDate">
<mods:number id="A4C9D3344B15720CD911D28DC5C7CF79">2022-05-03</mods:number>
</mods:detail>
<mods:detail id="A0AF178B5A28813673C943E1FB38F1A3" type="volume">
<mods:number id="659EC4FCE55A1AE25F31B2A8E01230C7">1098</mods:number>
</mods:detail>
<mods:extent id="43678D5DA0F5FE14C8A1AD9DE9F919CE" unit="page">
<mods:start id="F16CE601F3F5C8EA67A015F51913DA5E">1</mods:start>
<mods:end id="C01BF89391560DA09B10216C3065403E">180</mods:end>
</mods:extent>
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<mods:location id="4204E48426D83CF57B4E6817BD7B3F40">
<mods:url id="C785A98035B020A0B1D52B8982776271">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:url>
</mods:location>
<mods:classification id="AD43EFA270FB76F2A04E6527510C87D0">journal article</mods:classification>
<mods:identifier id="8AED0649ECFB9ECB021451B507EC47CC" type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
<mods:identifier id="C4FFDECB0163297DBC91206C2FB67454" type="Pensoft-Pub">1313-2970-1098-1</mods:identifier>
<mods:identifier id="E1817C7C0044964904152435A7752D2D" type="ZooBank">B4F4033F296E43CCBE54B9413BC19268</mods:identifier>
<mods:identifier id="45024FADCA5E718F53EA65D2EC2621DC" type="Pensoft-UUID">B22184E75B1A59C9B985C0DD9F092196</mods:identifier>
</mods:mods>
<treatment id="A11627C74A08512DAFE3B8632BA87703" ID-GBIF-Taxon="195190287" LSID="urn:lsid:zoobank.org:act:68F229F9-571D-4B71-8F98-0B89218D660E" httpUri="http://treatment.plazi.org/id/A11627C74A08512DAFE3B8632BA87703" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="B3EB8A55DF37DF70C2B1EF74833AE26A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="31FCD0E520A0F14C7D6B3B1DFD43049C" pageId="0" pageNumber="1">
<taxonomicName id="522C9B2A2C466F692261105A8FEC6DBA" LSID="http://zoobank.org/68F229F9-571D-4B71-8F98-0B89218D660E" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus mahafaly" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahafaly" status="sp. nov.">Camponotus mahafaly</taxonomicName>
<taxonomicNameLabel id="1FC3576CEE12B6E130F31BBAEBF37D04" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0BD2A858179DA75FA6BE24185A2489F2" pageId="0" pageNumber="1" type="description">
<paragraph id="FA6221E141552635962C0957C485DDFE" pageId="0" pageNumber="1">
<figureCitation id="4FBC2E17FEB91A35C0F3E10CA877B0F0" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Head in full-face view A C. mahafaly (CASENT 0115287) B C. tapia (CASENT 0493939) C C. quadrimaculatus (CASENT 0096044)." figureDoi="10.3897/zookeys.1098.73223.figure15" httpUri="https://binary.pensoft.net/fig/678074" pageId="0" pageNumber="1">Figs 15A</figureCitation>
<figureCitation id="7F927EC294A9DF01E052A027978BB0B2" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Mesosoma in lateral view A C. imitator (CASENT 0452849) B C. mahafaly (CASENT 0115287)." figureDoi="10.3897/zookeys.1098.73223.figure16" httpUri="https://binary.pensoft.net/fig/678075" pageId="0" pageNumber="1">, 16B</figureCitation>
<figureCitation id="F84D2B0E15F8547CEDCB980DC05B60EF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Head in full-face view A C. mahafaly (CASENT 0115287) B C. hovahovoides (CASENT 0487242)." figureDoi="10.3897/zookeys.1098.73223.figure20" httpUri="https://binary.pensoft.net/fig/678079" pageId="0" pageNumber="1">, 20A</figureCitation>
<figureCitation id="9099FC03AA43A04EF54812B1611F3146" captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Camponotus mahafaly A lateral view B head in full-face view C dorsal view of minor worker CASENT 0115287 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure66" httpUri="https://binary.pensoft.net/fig/678125" pageId="0" pageNumber="1">, 66</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="700836682B3E3181BB98CC5BA19EEAC5" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="2C6C82105A2D642370E196BC80D530B8" pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph id="CA65E781AA62AAD9B22727B131236F99" pageId="0" pageNumber="1">
<emphasis id="233A72A04E94B95A05C49B79635F234C" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis id="7F956D5765FEC919DFF69A76E4839FFD" bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
: Androy Region, District of Tsihombe, 74 km S of Tsihombe, Reserve Speciale Cap Ste Marie,
<geoCoordinate id="C6C5686A74DF5BAF27A1383420FD1FE9" degrees="25.58767" direction="south" orientation="latitude" precision="1" value="-25.58767">-25.58767</geoCoordinate>
,
<geoCoordinate id="E280897E69B9FFA49448AEE561A92815" degrees="45.163" direction="east" orientation="longitude" precision="55" value="45.163">45.163</geoCoordinate>
, 36 m, spiny bush, Malaise trap, 30 Apr-11 May 2003, (
<normalizedToken id="5F27E6870AF44F2D592941120509305E" originalValue="Rin’ha">Rin'ha</normalizedToken>
, Mike) CASENT0115206 (CAS).
</paragraph>
</subSubSection>
<subSubSection id="E7E184E919806C9DD603EE7A32D0C5C7" pageId="0" pageNumber="1" type="paratypes">
<paragraph id="99C930EA1F822AB69904AE986EE7FE58" pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph id="97A50696CFEE2F067971DD0B01534CE0" pageId="0" pageNumber="1">2 minor workers of same data as holotype but respectively with collection codes: MG-23-34 and MG-23-33 and specimen codes: CASENT0115287 and CASENT0115718 (CAS).</paragraph>
</subSubSection>
<subSubSection id="B272929C6AE04621315075C5B54CE9C6" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="E8DAD38CC463CE54C8010144F1324BEA" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="CE3923D2CE9EF3EBF6F9BA1DDDE98FDA" pageId="0" pageNumber="1">
<materialsCitation id="F44D0E6A68D42E6A670F3ED036FE12CF" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="RS Cap Ste Marie, PN Andohahela, Parcelle II, de Tsinjoriaky, RS Cap Sainte Marie, M. E. Irwin" country="Madagascar" county="Androy Region" elevation="176" latitude="-24.65033" location="Mike" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" specimenCount="1" stateProvince="Toliara">
<emphasis id="3F382C309131D29FEC5F3F9EB319CDD2" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="467FC9C73632AC642AD539E2C3A9A0D3" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion id="FD2612D8B00BCD038DBFC61C671DC032" country="Madagascar" name="Toliara">Toliara</collectingRegion>
</emphasis>
:
<collectingCounty id="818E3DAAAB62E0D460AD0D65E782EBD7">Androy Region</collectingCounty>
,
<collectingMunicipality id="718DDEFAFF5280ED28D7609106850C17">District of Tsihombe</collectingMunicipality>
,
<locationDeviation id="9346412B5CD6D7B5FF97103C539B2E0D" location="Tsihombe">
<quantity id="CEDA4349C5C506C14DDFDF132F7E38EF" metricMagnitude="4" metricUnit="m" metricValue="7.4" unit="km" value="74.0">74 km</quantity>
S of Tsihombe
</locationDeviation>
,
<collectorName id="625729115AF6478A7A083D6AF3CDEA0A">RS Cap Ste Marie</collectorName>
,
<geoCoordinate id="51FB6136084085E9639B8085F6680DF1" degrees="25.58767" direction="south" orientation="latitude" precision="1" value="-25.58767">-25.58767</geoCoordinate>
,
<geoCoordinate id="0BC7ECE4569F7F4C2050A1AAD1454A93" degrees="45.163" direction="east" orientation="longitude" precision="55" value="45.163">45.163</geoCoordinate>
, 36 m, spiny bush, (
<location id="31EBF8DA77244F57AC4F1B8BFAF063A5" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:31EBF8DA77244F57AC4F1B8BFAF063A5" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Mike" stateProvince="Toliara">Mike</location>
,
<location id="019883B84201399BA7D3CC0EB45769F5" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:019883B84201399BA7D3CC0EB45769F5" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Frank Parker" stateProvince="Toliara">Frank Parker</location>
,
<normalizedToken id="CF6E9901CACAAEAB61BCE5A2348FE8EB" originalValue="Rin’ha">
<location id="C51C67131BB68310FA1F640828319995" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:C51C67131BB68310FA1F640828319995" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Rin'ha" stateProvince="Toliara">Rin'ha</location>
</normalizedToken>
) (CAS);
<location id="D47C07B33D79DC02071109B6FAD21005" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:D47C07B33D79DC02071109B6FAD21005" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Androy Region" stateProvince="Toliara">Androy Region</location>
,
<location id="907B681DAEA43A6159A274BE1962F8C5" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:907B681DAEA43A6159A274BE1962F8C5" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="District of Tsihombe" stateProvince="Toliara">District of Tsihombe</location>
,
<locationDeviation id="AA1C29E222B19090362EF5D4A84695CF" location="Tsihombe">
<quantity id="F8DD121F13B4109E008DFA671E2A5963" metricMagnitude="4" metricUnit="m" metricValue="7.4" unit="km" value="74.0">74 km</quantity>
S of Tsihombe
</locationDeviation>
,
<collectorName id="98F327A6FF51AAB54AEABE0A543BEF34">RS Cap Ste Marie</collectorName>
,
<geoCoordinate id="2FBE5A89706C8BD5ACED3DEDAAA449A0" degrees="25.58767" direction="south" orientation="latitude" precision="1" value="-25.58767">-25.58767</geoCoordinate>
,
<geoCoordinate id="621250B9A98C6707989C547B9A45050A" degrees="45.163" direction="east" orientation="longitude" precision="55" value="45.163">45.163</geoCoordinate>
, 152 m,
<location id="B33E25B447EC9429F30F7DAB88A2E9D0" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:B33E25B447EC9429F30F7DAB88A2E9D0" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Bush" stateProvince="Toliara">Bush</location>
(
<location id="AD93D1717AB48A17A6E7BD961F96E9BB" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:AD93D1717AB48A17A6E7BD961F96E9BB" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Mike" stateProvince="Toliara">Mike</location>
,
<normalizedToken id="9A43A686604379AE111FAF7120E012CD" originalValue="Rin’ha">
<location id="57720F2C058F774FECD68BCA306D6241" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:57720F2C058F774FECD68BCA306D6241" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Rin'ha" stateProvince="Toliara">Rin'ha</location>
</normalizedToken>
) (CAS);
<location id="C7668ACC2E053908B2614FA5453BC3F3" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:C7668ACC2E053908B2614FA5453BC3F3" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Anosy Region" stateProvince="Toliara">Anosy Region</location>
,
<location id="49F62EECDF335771EFE7FD1D11581A46" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:49F62EECDF335771EFE7FD1D11581A46" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="District of Fort-Dauphin" stateProvince="Toliara">District of Fort-Dauphin</location>
,
<collectorName id="2B14256D9947C13472B72D502AF0377F">PN Andohahela</collectorName>
,
<collectorName id="898BDEED52513BDC87B50B3283B4CE79">Parcelle II</collectorName>
,
<location id="2F511CC22BE2074E15A0AA978B7F414F" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:2F511CC22BE2074E15A0AA978B7F414F" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Tsimela" stateProvince="Toliara">Tsimela</location>
,
<locationDeviation id="280DF7D0CA65AB318144CA44562589C7" location="Fort-Dauphin">
<quantity id="6701FF15D2F15FC64BFDFAAA6AF74CCD" metricMagnitude="4" metricUnit="m" metricValue="4.2" unit="km" value="42.0">42 km</quantity>
W of Fort-Dauphin
</locationDeviation>
,
<geoCoordinate id="A0D16703F2DEEE17F43991395B4637BD" degrees="24.93683" direction="south" orientation="latitude" precision="1" value="-24.93683">-24.93683</geoCoordinate>
,
<geoCoordinate id="25ACF70389F2909A5AED9BD04F0CDDE2" degrees="46.62667" direction="east" orientation="longitude" precision="1" value="46.62667">46.62667</geoCoordinate>
,
<quantity id="83D7FF1558D34FAE1BC7DC3B1B6439B5" metricMagnitude="2" metricUnit="m" metricValue="1.76" unit="m" value="176.0">
<elevation id="5C58CEE2FDD208B3C4671E93584E7278" metricMagnitude="2" metricUnit="m" metricValue="1.76" unit="m" value="176.0">176 m</elevation>
</quantity>
, transition forest (Michael Irwin, Frank Parker,
<normalizedToken id="1CC66255C22413D0B74C46B1F347C2E9" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS);
<location id="7AB8B1562A15C208CE747F03D001D8D0" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:7AB8B1562A15C208CE747F03D001D8D0" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Foret de Tsinjoriaky" stateProvince="Toliara">
<normalizedToken id="BC4A401A6AE2DF6DD9CA6100E44F8E47" originalValue="Forêt">Foret</normalizedToken>
<collectorName id="1F1CCF474D6C44E39BF00DA66A535C58">de Tsinjoriaky</collectorName>
</location>
,
<quantity id="96701ED0AEED1F177344EB250867FC6C" metricMagnitude="3" metricUnit="m" metricValue="6.2" unit="km" value="6.2">6.2 km</quantity>
<geoCoordinate id="8B000760D38F63CC389565CE8997AD72" degrees="84" direction="east" orientation="longitude" precision="55555" value="84.0">84° E</geoCoordinate>
<location id="6C7F6857ED4CF03717D3BF61ED9F726F" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:6C7F6857ED4CF03717D3BF61ED9F726F" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Tsifota" stateProvince="Toliara">Tsifota</location>
,
<geoCoordinate id="A2F082AB1C4A6A7232EE2E2D263D2B99" degrees="22.80222" direction="south" orientation="latitude" precision="1" value="-22.80222">-22.80222</geoCoordinate>
,
<geoCoordinate id="4FE948504E099BFF644C244A12379D48" degrees="43.42067" direction="east" orientation="longitude" precision="1" value="43.42067">43.42067</geoCoordinate>
,
<quantity id="68BFEA31E17489C79D227EA2B8A62C5B" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">
<elevation id="E6234578F93B8FED8E054ED33A3291AE" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">70 m</elevation>
</quantity>
, spiny forest/thicket (Fisher-Griswold Arthropod Team) (CAS);
<collectorName id="D04FE77465D5F3F71609072CF55AAD22">RS Cap Sainte Marie</collectorName>
,
<locationDeviation id="1601287F1F6B1DAF830EC397CE96A92B" location="Marovato">
<quantity id="F11FF0B041212477047E01BEB9079198" metricMagnitude="4" metricUnit="m" metricValue="1.23" unit="km" value="12.3">12.3 km</quantity>
262° W Marovato
</locationDeviation>
,
<geoCoordinate id="C6344196030253C5E4D25F5584AD46E2" degrees="25.58167" direction="south" orientation="latitude" precision="1" value="-25.58167">-25.58167</geoCoordinate>
,
<geoCoordinate id="A6AFB645E077631E8161C683CE5311D0" degrees="45.16833" direction="east" orientation="longitude" precision="1" value="45.16833">45.16833</geoCoordinate>
,
<quantity id="3BFDF9035F1EFFA98A12509DD8C2D99F" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="F5215CE2E48FB5FF7D1EFFDFC6AB6068" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
, spiny forest/thicket (Fisher-Griswold Arthropod Team) (CAS);
<locationDeviation id="1E157A181B43C3FBCDD7E1F947D21714" location="Itampolo">
<quantity id="B7E792D010B82C6B2154123B69609457" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
E Itampolo
</locationDeviation>
, malaise across path of plateau of
<location id="6496A91A8BDD58DA46F583261AEAD33D" LSID="urn:lsid:plazi:treatment:A11627C74A08512DAFE3B8632BA87703:6496A91A8BDD58DA46F583261AEAD33D" country="Madagascar" county="Androy Region" latitude="-24.65033" longLatPrecision="1" longitude="43.96317" municipality="District of Tsihombe" name="Andrimpano Forest" stateProvince="Toliara">Andrimpano Forest</location>
,
<geoCoordinate id="5A04C09A3D1E25AC3E62E81F60C72B52" degrees="24.65033" direction="south" orientation="latitude" precision="1" value="-24.65033">-24.65033</geoCoordinate>
,
<geoCoordinate id="06119CB01C26F997821ABDAF13147F7F" degrees="43.96317" direction="east" orientation="longitude" precision="1" value="43.96317">43.96317</geoCoordinate>
,
<quantity id="5367195FF7F15C5262AC1EB68651C4B7" metricMagnitude="2" metricUnit="m" metricValue="1.3" unit="m" value="130.0">
<elevation id="D6617C1BF14B23C48A6CA63F5F54F737" metricMagnitude="2" metricUnit="m" metricValue="1.3" unit="m" value="130.0">130 m</elevation>
</quantity>
, dry forest (
<collectorName id="B8F27E999B34B5C445268F30052F72E5">M.E. Irwin</collectorName>
,
<normalizedToken id="151287512AD1AB8E438D26D7EBF60221" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7198EB09FE0885285E20CD8C82E948E1" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="6303E49988C5D0A8CBC33AD5B2844CDB" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="9637D924417FEB3C546754D9F0ECD77A" pageId="0" pageNumber="1">With head in full-face view, lateral margins of head anterior to eye level parallel and lacking erect hairs, lateral and anteromedian clypeal margins continuously forming broad convexity; scape without erect hairs; mesosoma long and low; body color yellowish to brown.</paragraph>
</subSubSection>
<subSubSection id="9F5DF5527C7049B262A8733FAD9B466F" pageId="0" pageNumber="1" type="description">
<paragraph id="61F54FD21EF38F306FD10ABD6218A84A" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="0B87D70DF0C7C6CD45C7A18409CD9202" pageId="0" pageNumber="1">
<emphasis id="74FB527309CE6D8A14DCC266E02325E8" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides anterior to level of eye parallel, rounding evenly to posterior margin behind eye level; eye protruding and large (EL/CS: 0.32
<normalizedToken id="4DDD34FC608BD92E10A16B8B94936283" originalValue="±">+/-</normalizedToken>
0.01; 0.30-0.35), breaking lateral cephalic margin, level of its posterior margin situated approximately at posterior 1/4 of head (PoOc/CL: 0.23
<normalizedToken id="987FA8483B5570E0C7772DB7C6DF36A6" originalValue="±">+/-</normalizedToken>
0.01; 0.20-0.24); frontal carinae close to each other (FR/CS: 0.27
<normalizedToken id="F508DB229A8DC01332EBFE2208B82C5E" originalValue="±">+/-</normalizedToken>
0.01; 0.25-0.28), posteriorly parallel, distance between them smaller than their smallest distance to eye; clypeus without well-defined anterolateral angle, its anteromedian margin broadly convex; mandible with two apical teeth normally spaced; antennal scape relatively long (SL/CS: 1.40
<normalizedToken id="C22853E3B466FB793968DB7373A2E64D" originalValue="±">+/-</normalizedToken>
0.04; 1.33-1.45). Promesonotum weakly convex; mesonotum with posterior portion flat immediately anterior to weakly visible metanotal groove; propodeal dorsum almost straight (MPH/ML: 0.37
<normalizedToken id="82D2B18EED8BD2595EBCC9469B125AAA" originalValue="±">+/-</normalizedToken>
0.03; 0.34-0.50); rounding to declivity, 2
<normalizedToken id="8657B04E325FD4BAF76EE96AB60D82D0" originalValue="×">x</normalizedToken>
as long as declivity. Petiolar node short and high, with dorsal margin inclined posteriorly, rounding to anterior margin; height of anterior face 2/3 of that of posterior face; femur of hind leg rounded axially, without twist near base.
</paragraph>
<paragraph id="25722FC250B467670E2F7C53D80E10FE" pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; lateral margin of head anterior to eye level, with no erect hairs posterior to eye level; two erect hairs present close to posterior margin; antennal scape covered only with appressed hairs; erect hairs lacking on promesonotum; posterodorsal corner of propodeum with a pair of erect hairs. Integument shining; body color yellowish.</paragraph>
<paragraph id="D6FEB113465CD094EC6B5C06BB753DB7" pageId="0" pageNumber="1">
<emphasis id="6A9A34894913FA09760EF5EB8429FD9D" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="AE26BADC11D3C6F48D7706894B3C7569" pageId="0" pageNumber="1" type="distribution">
<paragraph id="345F86336D2D0EDBD475E81B1EFF297F" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="C40DA701188C9E5942199C0C2AFFA2E6" pageId="0" pageNumber="1">
<taxonomicName id="85CD5D382F72B35764EC521F3AA1A794" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus mahafaly" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahafaly">
<emphasis id="90E490D13BF3EEC84AECEF7DE0AC4723" italics="true" pageId="0" pageNumber="1">Camponotus mahafaly</emphasis>
</taxonomicName>
is geographically restricted to dry forest and spiny bush and forest habitats of the southwest and south of Madagascar (Fig.
<figureCitation id="6828D4321B9BA2CD10C4F47079C6D2F5" captionStart="Figure 66" captionStartId="F66" captionText="Figure 66. Camponotus mahafaly A lateral view B head in full-face view C dorsal view of minor worker CASENT 0115287 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure66" httpUri="https://binary.pensoft.net/fig/678125" pageId="0" pageNumber="1">66D</figureCitation>
). Nest sites are generally located in rotten logs, and foraging occurs on the surface of the ground.
</paragraph>
<caption id="BB57D7E4A14069DAEF59F7FDCC600C15" doi="10.3897/zookeys.1098.73223.figure66" httpUri="https://binary.pensoft.net/fig/678125" pageId="0" pageNumber="1" start="Figure 66" startId="F66">
<paragraph id="F8D6109BC3514DEC349E333E781BB8FD" pageId="0" pageNumber="1">
<emphasis id="B59F8916CAB3E1B00A238D8E81FADB00" bold="true" pageId="0" pageNumber="1">Figure 66.</emphasis>
<taxonomicName id="E2F4FFC8105F5311D43321CD94F81D12" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus mahafaly" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahafaly">
<emphasis id="3C018EB13A6CA0B48C5D2E09982A3C00" italics="true" pageId="0" pageNumber="1">Camponotus mahafaly</emphasis>
</taxonomicName>
<emphasis id="279BCC413045FC764A442DF582B8565E" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis id="A9A011A020626B077A32BD114D37B300" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis id="1248BF2F739D3D197FAA350FA312E579" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of minor worker CASENT0115287
<emphasis id="4D335D7BF289B4639EF7AF2792F9E2B4" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E869E9F08CC7B8E7A31EA492CB672AF9" pageId="0" pageNumber="1" type="discussion">
<paragraph id="EC16729AB57FFDB6319C3A44C7B1D394" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="E24DC3989CB1860FA82009AC51731EFF" pageId="0" pageNumber="1">
<taxonomicName id="95E6EC1F76A2163CF506A84B59336C2B" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus mahafaly" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahafaly">
<emphasis id="4D09C1020F3CF1C62C11BA4DB2D338BC" italics="true" pageId="0" pageNumber="1">Camponotus mahafaly</emphasis>
</taxonomicName>
might be confounded with
<taxonomicName id="BE49D5AFB7DE498A91EA0DE0AF01E365" lsidName="C. cemeryi" pageId="0" pageNumber="1" rank="species" species="cemeryi">
<emphasis id="5B8CDF2C14CE83F0F06C6154E4408D0E" italics="true" pageId="0" pageNumber="1">C. cemeryi</emphasis>
</taxonomicName>
because the petiole of both species is not nodelike, but in the latter species the antennal scape is covered with suberect hairs and the mesosoma is generally short and high, with a dorsal outline that is strongly convex.
</paragraph>
<paragraph id="620E6F3CD75BA668D18CA0A2CB406C54" pageId="0" pageNumber="1">
The qualitative morphology-based study of this species agrees with the multivariate morphometrics to support the taxonomic determination of
<taxonomicName id="304D22C832BD3AB7933648E0FB2BED1F" lsidName="C. mahafaly" pageId="0" pageNumber="1" rank="species" species="mahafaly">
<emphasis id="68438035B7271D4DE2DC51C3EF8B57CA" italics="true" pageId="0" pageNumber="1">C. mahafaly</emphasis>
</taxonomicName>
. Separation of this species is confirmed by LDA with 100% success.
</paragraph>
</subSubSection>
<subSubSection id="79B6752ADE14FAE14077925E1E4D6DD8" pageId="0" pageNumber="1" type="etymology">
<paragraph id="4475EEDAB6F4ABC543F2EEF079CD8EB5" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="9DFCBB9C2C87A811E0D4DE3E2C5B6DAF" pageId="0" pageNumber="1">
The species name
<emphasis id="780E5CA1349118FE1F9032D10BB1B03B" italics="true" pageId="0" pageNumber="1">mahafaly</emphasis>
is a singular non-Latin noun used in apposition and refers to the Mahafaly region in the southwest of Madagascar.
</paragraph>
</subSubSection>
</treatment>
</document>
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