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<document id="E6F84E4CCE37EA86C06363B8FD4D6B66" ID-CLB-Dataset="35725" ID-DOI="http://dx.doi.org/10.3897/zookeys.481.8225" ID-GBIF-Dataset="861ee59e-245c-4a17-8294-0f137f36c03d" ID-PMC="PMC4319097" ID-Pensoft-Pub="1313-2970-481-1" ID-PubMed="25685028" ID-ZooBank="791823C0114348FFB6B46DA47934A6F2" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-481-1" ModsDocOrigin="ZooKeys 481" ModsDocTitle="New Bythinella (Gastropoda, Bythinellidae) species from western Turkey" checkinTime="1451244685949" checkinUser="pensoft" docAuthor="Yildirim, Mehmet Zeki, Kebapci, Uemit, Koca, Seval Bahadir &amp; Yuece, Arzu" docDate="2015" docId="A0F0F7F4FA3447A5EFD13CF910A123A8" docLanguage="en" docName="ZooKeys 481: 1-13" docOrigin="ZooKeys 481" docSource="http://dx.doi.org/10.3897/zookeys.481.8225" docTitle="Bythinella istanbulensis Yildirim, Kebapci &amp; Yuece, sp. n." docType="treatment" docUuid="8958B919-4F97-4EAA-A6F8-EB7059A8FEB7" docUuidSource="ZooBank" docVersion="7" lastPageNumber="6" masterDocId="F833FFB4FFBEFFEAFFE68A6EFFFA3907" masterDocTitle="New Bythinella (Gastropoda, Bythinellidae) species from western Turkey" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1732710449497" updateUser="ExternalLinkService">
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<mods:title id="7C2E30610D333B0D12ABE733F7C62AD2">New Bythinella (Gastropoda, Bythinellidae) species from western Turkey</mods:title>
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<mods:namePart id="0ABEC516372272EF9194D57C71ABDA6B">Yildirim, Mehmet Zeki</mods:namePart>
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<mods:namePart id="282C4CB6C146297AD9AAD1A455E9D1F1">Kebapci, Uemit</mods:namePart>
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<mods:namePart id="5BEE9B945EB7AD27E3D7615742528050">Koca, Seval Bahadir</mods:namePart>
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<mods:namePart id="4E1BB10B1097CCA4792E286320DA9B57">Yuece, Arzu</mods:namePart>
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<mods:identifier id="6D7339790A9C8A8EB9E42FDF3FE9FC19" type="DOI">http://dx.doi.org/10.3897/zookeys.481.8225</mods:identifier>
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<treatment id="A0F0F7F4FA3447A5EFD13CF910A123A8" ID-GBIF-Taxon="152058185" LSID="urn:lsid:zoobank.org:act:8958B919-4F97-4EAA-A6F8-EB7059A8FEB7" httpUri="http://treatment.plazi.org/id/A0F0F7F4FA3447A5EFD13CF910A123A8" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" scope_class="Gastropoda" scope_family="Bythinellidae">
<subSubSection id="1D79A988FF02EB4D5B5CE3F8A87A0410" pageId="2" pageNumber="3" type="multiple">
<paragraph id="CB288CC441D226A40BAA0591AD2A107A" pageId="2" pageNumber="3">Taxon classification Animalia Littorinimorpha Bythinellidae</paragraph>
</subSubSection>
<subSubSection id="1020D54F6BDD44F9B83B11DC77663245" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="5E6228A9C58BD2E9F8F63959D72412C7" pageId="2" pageNumber="3">
<taxonomicName id="589EA23C878F1B26B3510144B4F5EA63" ID-CoL="P2QM" LSID="http://zoobank.org/8958B919-4F97-4EAA-A6F8-EB7059A8FEB7" authority="Yildirim, Kebapci &amp; Yuece" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella istanbulensis" order="Littorinimorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="istanbulensis">
Bythinella istanbulensis
<normalizedToken id="AB38E582867EA17831F79704B0EAB4E5" originalValue="Yıldırım">Yildirim</normalizedToken>
,
<normalizedToken id="19DFA055D0E426437DE2A499076556EF" originalValue="Kebapçı">Kebapci</normalizedToken>
&amp;
<normalizedToken id="ED2A417C863F33750BE5A0705845CC13" originalValue="Yüce">Yuece</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="1710E89339C30238B9BC4FB1BBF2DA37" pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
Figs 3, 6b
</paragraph>
</subSubSection>
<subSubSection id="F006E1DCF0941D95A669E66CF4171D62" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="EF6993C6A6DF7FCB3A0C5C386F7F1780" pageId="3" pageNumber="4">
<taxonomicName id="F13DDD91A4CE1588D4CF7A3CF33EF010" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="opaca">
<pageBreakToken id="B7BFEB0B1E7993956561CC68FC64F49F" pageId="3" pageNumber="4" start="start">Bythinella</pageBreakToken>
opaca
</taxonomicName>
,
<normalizedToken id="A4D575C1E47F3005070242BD32581C98" originalValue="Schütt">Schuett</normalizedToken>
, H., 1965 Zur Systematik und
<normalizedToken id="EA11419707EF76331B06DAE036F14EE7" originalValue="Ökologie">Oekologie</normalizedToken>
<normalizedToken id="E282E5AECFA8731F6CC23CC77D30128D" originalValue="Türkischer">Tuerkischer</normalizedToken>
<normalizedToken id="7AEE646E00F020754C7915AD9EC773D5" originalValue="Süsswasserprosobranchier">Suesswasserprosobranchier</normalizedToken>
. Zoologische Mededelingen, 41: 43-71. (misidentification).
</paragraph>
<paragraph id="F7A34042437B2663D323C625A39585C8" pageId="3" pageNumber="4">
<taxonomicName id="AA510CBADBBB0DA9D7E29EFABFECD483" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="opaca">Bythinella &quot; opaca &quot;</taxonomicName>
,
<normalizedToken id="8B676399C63FF78AC48EA54D8DDF3699" originalValue="Schütt">Schuett</normalizedToken>
, H., 1980 Zur Kenntnis griechischer Hydrobiiden, Arch.Moll.110 (4/6):115.
</paragraph>
<paragraph id="F75EB307E50DC5C29B9AE61029E8246A" pageId="3" pageNumber="4">
<taxonomicName id="F1C542418A484B25266290448D3826BA" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">Bythinella</taxonomicName>
sp. A,
<bibRefCitation id="DF802DF5BCB7F9AAD062D157433543E3" author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="11" pageNumber="12" pagination="197 - 204" title="Supplement to the Prosobranchia (Mollusca: Gastropoda) Fauna of Fresh and Brackish Waters of Turkey." volume="30" year="2006">
<normalizedToken id="6DF8B634131174ED76B43D2950859080" originalValue="Yıldırım">Yildirim</normalizedToken>
et al. 2006
</bibRefCitation>
Supplement to the
<taxonomicName id="EED4976775501BFE7111E5BC6F67C957" lsidName="Prosobranchia" pageId="3" pageNumber="4" rank="subclass" subclass="Prosobranchia">Prosobranchia</taxonomicName>
(
<taxonomicName id="CDF9FA3511C03AC76D3455A8A88110F6" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosobranchia" pageId="3" pageNumber="4" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
:
<taxonomicName id="02A7240FEA7BC7EE144529F591D5C2DB" class="Gastropoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosobranchia" pageId="3" pageNumber="4" phylum="Mollusca" rank="class">Gastropoda</taxonomicName>
) Fauna of Fresh and Brackish Waters of Turkey, Tr. J. Zool. 30: 197-204.
</paragraph>
</subSubSection>
<subSubSection id="4A2EC4B3EA757A7450DA3F50C6EED1A3" pageId="3" pageNumber="4" type="holotype">
<paragraph id="EB654B2E36FDC808BD599DF92F0F1DE8" pageId="3" pageNumber="4">Holotype</paragraph>
<paragraph id="A6EBD018C271BE26D51103DDC8C93762" pageId="3" pageNumber="4">
(Coll.
<normalizedToken id="8D2CE6C653107EDCFB2F9E28F2F36B91" originalValue="Yıldırım">Yildirim</normalizedToken>
): Shell height: 2.74 mm, width 1.43 mm (Coll. M.Z.
<normalizedToken id="29EC5425DD72DA7A41832EC3F0C51C3E" originalValue="Yıldırım">Yildirim</normalizedToken>
); TURKEY, Istanbul,
<normalizedToken id="2676C4CC44A4B74B46945256E5EC984F" originalValue="Bahçeköy">Bahcekoey</normalizedToken>
, a small spring at the entrance of
<normalizedToken id="487DAC8A974E09116D7C1259597D4F3A" originalValue="Bahçeköy">Bahcekoey</normalizedToken>
in Belgrad Forest,
<geoCoordinate id="57C7B65D019E7E411DB6129BBEC2CD35" direction="north" orientation="latitude" precision="9" value="41.184834">N41°11.09'</geoCoordinate>
,
<geoCoordinate id="09C734F086360158A78B8F316AF02F5F" direction="east" orientation="longitude" precision="92" value="28.991667">E28°59.5'</geoCoordinate>
, 23.02.2013, leg. A.
<normalizedToken id="E4D51960A089C9BC66BCFF3A673AC0DD" originalValue="Yüce">Yuece</normalizedToken>
. Paratypes: 27 ex., same data and locality as holotype; 5 ex. Zoologisches Museum Hamburg (ZMH 79661), 5 ex. Naturhistorisches Museum in Wien (NHMW 109174), rest in the Coll.
<normalizedToken id="2C8F98E5C4B8E209C316F31143C58E09" originalValue="Yıldırım">Yildirim</normalizedToken>
. Additional material: 5 ex. in the University of Giessen (Coll. Prof. Wilke), rest in the Coll.
<normalizedToken id="42CF7778FE51B532319D14956B41FAA0" originalValue="Yıldırım">Yildirim</normalizedToken>
(5 dissected).
</paragraph>
</subSubSection>
<subSubSection id="2F1133499A6B9C1E38972D57B27FB35C" pageId="4" pageNumber="5" type="type locality">
<paragraph id="97C11854CAEBDC64A6D610C503B8C2C9" pageId="4" pageNumber="5">
<pageBreakToken id="0870FE42C0016837E3B284238AEEECBF" pageId="4" pageNumber="5" start="start">Type</pageBreakToken>
locality.
</paragraph>
<paragraph id="4A19A6B53C818CA8FA1E124B3C1AFCC6" pageId="4" pageNumber="5">
A small spring at the entrance of
<normalizedToken id="B055764CDF983CB39DFAD774FF6D6E3D" originalValue="Bahçeköy">Bahcekoey</normalizedToken>
in Belgrad Forest,
<normalizedToken id="09B75540BE53E128A3DAF6FA1A29C082" originalValue="Bahçeköy">Bahcekoey</normalizedToken>
, Istanbul.
</paragraph>
</subSubSection>
<subSubSection id="114382C4D1DDB38C474D436FB3617A5F" pageId="4" pageNumber="5" type="etymology">
<paragraph id="FBB820505B57D7D1B5AB6B38471580CE" pageId="4" pageNumber="5">Etymology.</paragraph>
<paragraph id="B5F9E837E82203EFE2028E49F33E4F6C" pageId="4" pageNumber="5">Named after the type locality.</paragraph>
</subSubSection>
<subSubSection id="9D620A5CB98287691FDEDC05638ADF97" pageId="4" pageNumber="5" type="description">
<paragraph id="73FEB47C39D2D064921F79BAA0B00D30" pageId="4" pageNumber="5">Description.</paragraph>
<paragraph id="FB1BF0DC118D164CCCBA9C1226531EAE" pageId="4" pageNumber="5">
Shell broad cylindrical oval, yellowish horn-colored, surface smooth and glossy covered by a thick, solid and rough encrustation due to chemical conditions of the spring water; apex blunt;
<normalizedToken id="DD56CD966E6C7D370CA2A80A197E5DE6" originalValue="3½">31/2-</normalizedToken>
4 convex whorls, rapidly and regularly increasing to form a cylindrical shape, last whorl about 2/3 of the shell height, sutures relatively deep; aperture pear-shaped, height nearly the same as or slightly less than the penultimate whorl, columellar margin with an distinct inner lip reaching basal and apical corners of the aperture; operculum orange, umbilicus narrow and slit-shaped (Figs 2a, 6a).
</paragraph>
<paragraph id="8985C9CBBF3B429AB4B92BE2664CB9C6" pageId="4" pageNumber="5">Tubular gland relatively short and thick. Pallial roof unpigmented, head having little pigmentation (Fig. 2b), rs1 small and attached to oviduct (Fig. 2c), thus not easily discernible.</paragraph>
<paragraph id="F2788EF65761C49D93667343FE6F91AB" pageId="4" pageNumber="5">Measurements (n=28): SH: 2.67 mm (min 2.18/max 2.87), SW: 1.66 mm (min 1.37/max 1.94), SH/SW: 1.61 (min 1.39/max 1.84), SW/SH: 0.62 (min 0.54/max 0.72), ah/SH: 0.41 (min 0.37/max 0.50).</paragraph>
<caption id="5251598F515D3D6791AE900D25C05BD7" pageId="4" pageNumber="5">
<paragraph id="95E22C4D863330C310B9A8961625D874" pageId="4" pageNumber="5">
Figure 3. Shell and anatomy of
<taxonomicName id="D1854C3BC659284194B21FD2777BFB51" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella istanbulensis" order="Littorinimorpha" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="istanbulensis">Bythinella istanbulensis</taxonomicName>
sp. n.: a shell b male head and penis in natural position c penis d female genitalia. Scale bars = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="816BE4D2AE8DCC4C69692FC727FCD6C2" pageId="4" pageNumber="5" type="differential diagnosis">
<paragraph id="BA25316D38792AEA6025B9900E411FF4" pageId="4" pageNumber="5">Differential diagnosis.</paragraph>
<paragraph id="2A277BE35626B695A7C594981FA74188" pageId="4" pageNumber="5">
This new species is distinguished from other Turkish
<taxonomicName id="BCB594B9A931960EEA84B60E28EA47E4" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella" order="Littorinimorpha" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">Bythinella</taxonomicName>
species by its nearly cylindrical shell with broad and blunt apex. Small and attached rs1, unpigmented and light coloured pallial roof are the other key characters.
</paragraph>
</subSubSection>
<subSubSection id="F6873F3007C091D92BE9E863CB69773E" pageId="4" pageNumber="5" type="habitat">
<paragraph id="9B1CD594D0974034A10FD7886238D60D" pageId="4" pageNumber="5">Habitat.</paragraph>
<paragraph id="E95AC13E5D17F807A1CA391911E89635" pageId="4" pageNumber="5">Specimens were collected from the stones in a small spring and its pond.</paragraph>
</subSubSection>
<subSubSection id="02C528870DE9F35F8EB1FE81C6DF6DE3" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="remarks">
<paragraph id="AA056A7B1AC014CE14B48EA1AE50DBA0" pageId="4" pageNumber="5">Remarks.</paragraph>
<paragraph id="5D18E974797B547F9B88602F3B3D55CD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Based on shell characters only,
<bibRefCitation id="A53FE47D224AD55A6C4C2968B2EF90B2" author="Schuett, H" journalOrPublisher="Zoologische Mededelingen" pageId="10" pageNumber="11" pagination="43 - 71" title="Zur Systematik und Oekologie Tuerkischer Suesswasserprosobranchier." volume="41" year="1965">
<normalizedToken id="3DFADA73308F37924C467F6E2A099227" originalValue="Schütt">Schuett</normalizedToken>
(1965)
</bibRefCitation>
misidentified
<taxonomicName id="7198BC92D5B8CF3C055BA21805E3A822" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<pageBreakToken id="A33CEBC1E28D6F45AF7E3D4BEC2BBC8C" pageId="5" pageNumber="6" start="start">Bythinella</pageBreakToken>
</taxonomicName>
specimens from Belgrad Forest as
<taxonomicName id="0D6907425CD6151B16E52E03AB11A3BE" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Bythinella opaca</taxonomicName>
(Frauenfeld, 1857), a species originally described from Carniola (Slovenia) and Italy.
<taxonomicName id="BFB013345DBF3522B5F15DFF8ADE0342" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella angelitae" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="angelitae">Bythinella angelitae</taxonomicName>
was introduced as a replacement name for the preoccupied name
<taxonomicName id="A61B7EF7444A1AEF3CBD37994C04DB89" class="Gastropoda" family="Assimineidae" genus="Paludinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Paludinella opaca</taxonomicName>
Frauenfeld, 1857, a homonym of
<taxonomicName id="F731D460A431A6D09F1793EBC20DA363" class="Gastropoda" family="Assimineidae" genus="Paludinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Paludinella opaca</taxonomicName>
M. von Gallenstein, 1848, by
<bibRefCitation id="6F4DE999B798D8762462A768BBCE72E8" author="Haase, M" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="1 - 16" title="Identifying species of Bythinella (Caenogastropoda: Rissooidea): A plea for an integrative opproach." volume="1563" year="2007">Haase et al. (2007)</bibRefCitation>
. The two closely related species
<taxonomicName id="AAD1032F0B1B59F26963781A8953E67B" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella angelitae" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="angelitae">Bythinella angelitae</taxonomicName>
and
<taxonomicName id="28458D05D58C61C68EB724331FD39534" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Bythinella opaca</taxonomicName>
can be morphologically separated only by radular dentition, and the geographic range of
<taxonomicName id="BFB6B7519512C1C6DA7F9693B4010574" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella angelitae" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="angelitae">Bythinella angelitae</taxonomicName>
is restricted to the border areas of Austria and Slovenia (
<bibRefCitation id="3192F372B924B1A521B710BD9C6A3C4A" author="Haase, M" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="1 - 16" title="Identifying species of Bythinella (Caenogastropoda: Rissooidea): A plea for an integrative opproach." volume="1563" year="2007">Haase et al. 2007</bibRefCitation>
). Although they have been recorded in previous studies (
<bibRefCitation id="4189574C9FDD03D143C9DCF4D86DBC56" author="Wagner, J" journalOrPublisher="Allattani Koezlemenyek" pageId="11" pageNumber="12" pagination="197 - 210" title="A Gutin-hegyseg Mollusca faunajanak alapvetese" volume="38" year="1941">Wagner 1941</bibRefCitation>
;
<bibRefCitation id="D36BC2AFDE9D360C701ED9DF74C63972" author="Grossu, AV" journalOrPublisher="Editura Acad. RPR, Bucuresti" pageId="10" pageNumber="11" pagination="1 - 220" title="Gastropoda. Prosobranchia si Opisthobranchia. Fauna Republicii Populare Romine. Mollusca" year="1956">Grossu 1956</bibRefCitation>
;
<bibRefCitation id="4A18D687E66481104F7F92DCED5400A6" author="Angelov, A" journalOrPublisher="News of the Zoological Institute with Museum, Sofia" pageId="9" pageNumber="10" pagination="411 - 413" title="Contribution to the study of the Bulgarian freshwater mollusks." volume="9" year="1960">Angelov 1960</bibRefCitation>
,
<bibRefCitation id="DCE059CFDD23EF19699A8757EFEDB590" author="Angelov, A" journalOrPublisher="Pensoft &amp; Backhuys Publ., Sofia, Leiden" pageId="9" pageNumber="10" title="Mollusca (Gastropoda et Bivalvia) aquae dulcis, catalogus Faunae Bulgaricae" year="2000">2000</bibRefCitation>
),
<taxonomicName id="18330EA9D362FB7549EB9FABFFE28462" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella austriaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="austriaca">Bythinella austriaca</taxonomicName>
(Frauenfeld, 1857) and
<taxonomicName id="0D6364F39111E89ED16F8CC0459A0085" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Bythinella opaca</taxonomicName>
(M. von Gallenstein, 1848) have not been confirmed from the eastern Balkan countries in recent studies (
<bibRefCitation id="1DC0D50AE9094B3FCC5D46CB9E1A9476" author="Falniowski, A" journalOrPublisher="Journal of Natural History" pageId="9" pageNumber="10" pagination="2955 - 2973" title="Bythinella Moquin-Tandon, 1856 (Gastropoda: Rissooidea: Bythinellidae) in Romania: species richness in a glacial refugium." url="10.1080/00222930903359636" volume="43" year="2009 a">Falniowski et al. 2009a</bibRefCitation>
,
<bibRefCitation id="655E2C317071F199B96664BACF4DB350" author="Falniowski, A" journalOrPublisher="Folia Malacologica" pageId="10" pageNumber="11" pagination="21 - 36" title="Bythinella Moquin-Tandon, 1856 (Gastropoda: Rissooidea: Bythinellidae) in Romania: its morphology with description of four new species." url="10.2478/v10125-009-0003-2" volume="17" year="2009 b">b</bibRefCitation>
). Therefore, there is a large distribution gap between the ranges of Central European taxa and that of
<taxonomicName id="6679AC25F9246338FA8A47A8B48FC0DF" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella istanbulensis" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="istanbulensis">Bythinella istanbulensis</taxonomicName>
sp. n.
</paragraph>
<paragraph id="7D0951C82CF025AC074E59DDB7387EBE" pageId="5" pageNumber="6">
Despite the conchological similarities, the new species is anatomically distinct from the Central European species
<taxonomicName id="EF20E90044794571E7E958E36153930C" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella austriaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="austriaca">Bythinella austriaca</taxonomicName>
and
<taxonomicName id="CBD52E1FCBA607F6F7D310CEC51F7580" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Bythinella opaca</taxonomicName>
. While the lighter pallial roof colouration is also observed in
<taxonomicName id="2B2F4E7605F330514471764F87A93F90" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella opaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="opaca">Bythinella opaca</taxonomicName>
(
<normalizedToken id="7F356C39B9B3210E325FC97794E88549" originalValue="Glöer">Gloeer</normalizedToken>
&amp;
<normalizedToken id="F866E3B4789DC3CF44714FFA9610B829" originalValue="Pešic">Pesic</normalizedToken>
, 2006),
<taxonomicName id="5C57167BC80C091A856C243E05408101" class="Gastropoda" family="Bythinellidae" genus="Bythinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bythinella austriaca" order="Littorinimorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="austriaca">Bythinella austriaca</taxonomicName>
has a dark pallial roof colouration. The new species differs from both of these taxa in having a very short and broad penial appendix (cf. long trumpet-shaped penial appendix in these species).
</paragraph>
</subSubSection>
</treatment>
</document>