165 lines
20 KiB
XML
165 lines
20 KiB
XML
<document ID-CLB-Dataset="22854" ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki & Bjoern Stelbrink," docDate="2020" docId="819F33E69E015A96978B52B692112C7D" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia pelkiae Gittenberger & Gyeltshen 2020, sp. nov." docType="treatment" docUuid="8548A11B-5AAC-4D3E-9064-6321DF57677C" docUuidSource="ZooBank" docVersion="6" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="6" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="4" updateTime="1732748779243" updateUser="ExternalLinkService">
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<mods:mods id="27F0B9745AF277286621FFE57B277922" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="60E0A0FBE08CAE42488204DCE7F92A2A">
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<mods:title id="84E42B350E2D566B3CA838E412A11DAC">The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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<mods:name id="5038D4B8283291600D00D73EEC8A1E5F" type="personal">
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<mods:roleTerm id="9B477C91565A1F0C01B8A2C31ABA08FB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C60D27E7A7D438D30EFA2E5EB3115CE7">Gittenberger, Edmund</mods:namePart>
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<mods:affiliation id="98E73BF9A392EF2CC527F0DC8962FF3D">Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier id="8D571756516FF04C448245A2D99B5819" type="email">egittenberger@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="30925E3890C76A5BC43E8B3BB16F545E" type="personal">
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<mods:roleTerm id="F60C1B48B5D2B83A40D54E4B62E06BFD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="09978C7727985A451F24D7CE470BF4CC">Leda, Pema</mods:namePart>
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<mods:affiliation id="B98C10E7821C1F1610D9BF152A1B3F1E">National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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</mods:name>
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<mods:name id="A8C00B62BFAD53E48472C04AFFEFF547" type="personal">
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<mods:roleTerm id="2E1A58008E6D409822EBFD1A47DA266F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C2404573EF0D695977DD4CB094998EFA">Wangchuk, Jigme</mods:namePart>
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<mods:affiliation id="AF8550BC035AC2A0DC50473140D0AA9E">Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:nameIdentifier id="3ACABD2DBC7785F8A6AAD82708821DCC" type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
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</mods:name>
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<mods:name id="56E6E15079B2B246E3204FAABF074890" type="personal">
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<mods:roleTerm id="F5FEFA9F711A6258C6A1B71C646908DB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E9C0D8B9AF401C627C7BBAB6CA7B5DE">Gyeltshen, Choki</mods:namePart>
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<mods:affiliation id="88507A458DF33D7AC923BBDFD9F9657F">National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:nameIdentifier id="37FD2A267D412E9437B73F6086F4BF13" type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
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<mods:roleTerm id="5AB4A600C087DC61C94129C06EA8D6C6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7FE596BF5F6161B867B11DEE462C0C12">Bjoern Stelbrink,</mods:namePart>
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<mods:affiliation id="6B9250DF5882540E9703EA197E2F038E">Department of Animal Ecology & Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany & Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="3786DDED20713FEBEB5218BB51D2364F" type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
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<mods:typeOfResource id="A71E8CB320905709E882236C12B7AAA2">text</mods:typeOfResource>
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<mods:title id="A4246579677BE632D22424612F866635">ZooKeys</mods:title>
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<mods:part id="75041C6949009A39137FF87B6B9C4E7E">
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<mods:date id="8455424638DB0CF0EB5CF237284EEBA7">2020</mods:date>
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<mods:detail id="7302AA0ADD022600FA578C6102C98EB9" type="volume">
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<mods:number id="1222F435D198EC3D0996F6872DB61F9B">929</mods:number>
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<mods:url id="F4444C0164ECA0FC04C18E0F909779A4">http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
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<mods:classification id="6FDB5376F46B0814963216A95A1569C5">journal article</mods:classification>
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<mods:identifier id="D7D3FBF070ADBD8012DC9A2CC3DAE061" type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
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<mods:identifier id="0DB3E625CA8E08F42788240AA4F8AC5F" type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
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<mods:identifier id="85CC4253896669612565DDF0E5DD7BF3" type="ZooBank">4FCA3A27-973C49F7-8264-6A34D279EC5C</mods:identifier>
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<treatment id="819F33E69E015A96978B52B692112C7D" ID-GBIF-Taxon="163735478" LSID="urn:lsid:zoobank.org:act:8548A11B-5AAC-4D3E-9064-6321DF57677C" httpUri="http://treatment.plazi.org/id/819F33E69E015A96978B52B692112C7D" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" scope_class="Gastropoda" scope_family="Amnicolidae" scope_superFamily="Rissooidea">
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<subSubSection id="05FF204B38BB4916EF94616A63C7B6BA" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="4C46255C99F4BFEB97CA8E9C7C0328D4" pageId="3" pageNumber="4">
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<taxonomicName id="5EE218F0A4301561A9EC677AC09D737A" ID-CoL="7R8DF" LSID="819F33E6-9E01-5A96-978B-52B692112C7D" authority="Gittenberger & Gyeltshen" authorityName="Gittenberger & Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae" status="sp. nov.">Erhaia pelkiae Gittenberger & Gyeltshen</taxonomicName>
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<taxonomicNameLabel id="D2BFC57E5037BAC5544E275BC44087FA" pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
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<figureCitation id="64FA876F55DA33BB330B390E54F14CAF" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">Figs 4</figureCitation>
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<figureCitation id="1CD67F4981780BFFC88C216016BA0EE3" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 5</figureCitation>
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<figureCitation id="2599DEF56E5416D31EEECFEEB04C5C9F" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="20113199AE31ADA9FCF1E63C7F365D04" pageId="4" pageNumber="5" type="materials_examined">
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||
<paragraph id="737D5A43827B628D96ED7A980DD14AA6" pageId="4" pageNumber="5">
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<pageBreakToken id="8D2E8EE9B234B8313FAEC8E8C694B1F2" pageId="4" pageNumber="5" start="start">Material</pageBreakToken>
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examined.
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</paragraph>
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<paragraph id="EF9D1AF7ECD82AF60B9DDDB9CFD7B53D" pageId="4" pageNumber="5">
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||
<emphasis id="8EE628DC29D604D989C8C010AE106A67" bold="true" pageId="4" pageNumber="5">
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<emphasis id="FDF48E481B5D2C33CFA5DEEF26124027" italics="true" pageId="4" pageNumber="5">Holotype</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation id="5CC39B051A8C5B8A15A68F7ADBDF3B6B" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">4</figureCitation>
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) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.;
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<geoCoordinate id="59623E169EEB1709F2F12494FFD528C5" degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43"N</geoCoordinate>
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,
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<geoCoordinate id="7B1D26EDAD1E69349B8EAEBC24B45928" degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10"E</geoCoordinate>
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; E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 25.X.2018; NBCB 1059.
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<emphasis id="738A2DA94D4A024FE05F65BB60BBC560" bold="true" pageId="4" pageNumber="5">
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<emphasis id="A4616A0FB32F8F91A5CC4A0D46F282C3" italics="true" pageId="4" pageNumber="5">Paratype</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation id="0042F9902F8B5F7472F08729FB623FEC" captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">5</figureCitation>
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) 1 shell; same data as for holotype; NBCB 1060.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CF02D6385514EF711F0D08D161EFA36" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="4ABED5CE50169F479037DF157F0F3FD4" pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph id="411096EC3C04197D54B49700B8B9ED65" pageId="4" pageNumber="5">Shell with a partly reflected peristome, teleoconch with spiral lirae.</paragraph>
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</subSubSection>
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<subSubSection id="2CCD98A0CC6D8E7EA4CB904601E2D7AA" pageId="4" pageNumber="5" type="shell">
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<paragraph id="B0775FEF686C398DA93B4074F0188DFE" pageId="4" pageNumber="5">Shell.</paragraph>
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<paragraph id="E2F0743843F81A4315A3DDBCB83B9470" pageId="4" pageNumber="5">
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Shell elongated ovoid, with
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<normalizedToken id="56E8B9ECB70B79B16AA2681DC1801047" originalValue="3½">31/2</normalizedToken>
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convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; light yellowish brown with fine growth lines and some brown periostracal ridges. Peristome reflected at the columellar and the basal side. Parietal and columellar side of the aperture thickened by a whitish callus. An irregular umbilical chink only in the paratype might be represent a malformation resulting from repair of the shell wall. Protoconch encrustated; teleoconch with fine spiral lines.
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</paragraph>
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</subSubSection>
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<subSubSection id="3F148EE6435970F88E7BB2F61569AEAF" pageId="4" pageNumber="5" type="description">
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<paragraph id="C3B078A425E226DFCA9404B2A20E1713" pageId="4" pageNumber="5">Measurements.</paragraph>
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<paragraph id="B88B63401C5F11513687D79E81E3C779" pageId="4" pageNumber="5">
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(
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<emphasis id="AA7B83A46EA6AE0E91A0E0E86F284BCD" italics="true" pageId="4" pageNumber="5">N</emphasis>
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= 2): holotype and paratype H 1.9 mm, B 1.1 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5C1EBA6FB212C769528A9559863F060" pageId="5" pageNumber="6" type="ecology">
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<paragraph id="232127952A5A55F6381E74303F688071" pageId="5" pageNumber="6">
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<pageBreakToken id="B17DEFBE67CE2E378ACBCB63B5813389" pageId="5" pageNumber="6" start="start">Ecology</pageBreakToken>
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</paragraph>
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<paragraph id="A2DB5253509EF1BF76B1AA3D23ECDA90" pageId="5" pageNumber="6">
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(Fig.
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<figureCitation id="C92116728FD2701EE8C720C88B15D7B3" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="5" pageNumber="6">11</figureCitation>
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). The snails are rare on the rocks in the shaded streamlet uphill the prayer wheel. See also the data for
|
||
<taxonomicName id="567371C23BBF98664BEE1D4AAC34841C" lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
|
||
<emphasis id="5BF2BD28CF509CD00D11F1B88632EE8C" italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CC52D7FFCB959DA5813D71C74054784B" pageId="5" pageNumber="6" type="notes">
|
||
<paragraph id="9965BB08AD2A43FFB260B19C86AA8610" pageId="5" pageNumber="6">Notes.</paragraph>
|
||
<paragraph id="4219553545F291A49DC5D5EFFC9230E1" pageId="5" pageNumber="6">
|
||
The differences between the sympatric
|
||
<taxonomicName id="9F831BDC3E97D25C65F9B9B3E747CD57" lsidName="E. pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
|
||
<emphasis id="BBAE4FD40575198ED4B13B71828886E8" italics="true" pageId="5" pageNumber="6">E. pelkiae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B297B0D2C4ACAC115C7C5FF973028BBD" lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
|
||
<emphasis id="4CDEBC6760291A73E80BC807C8566424" italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
|
||
</taxonomicName>
|
||
are too large to regard as sexual dimorphism.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82BFB9E0525B9541ADAAB998A4488179" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="5E27437A785E9B8F41355D97E5DF0E90" pageId="5" pageNumber="6">Etymology.</paragraph>
|
||
<paragraph id="23928ED483340EC169AA675442033082" pageId="5" pageNumber="6">
|
||
The epithet
|
||
<taxonomicName id="D72451A57E9533796E2E8643FAF91E53" lsidName="pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
|
||
<emphasis id="D972ECD25C3659DD176CC92C737157FD" italics="true" pageId="5" pageNumber="6">pelkiae</emphasis>
|
||
</taxonomicName>
|
||
refers to Ms. Pelki Yangdon, the daughter of the fourth author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |