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<mods:title id="6FD51084538967AB1AA8FB13F5D93C0B">Taxonomic studies of Diospyros L. (Ebenaceae) from the Malagasy region. IV. Synoptic revision of the Squamosa group in Madagascar and the Comoro Islands</mods:title>
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<mods:namePart id="B3F9649DFD7DD4D5B998956084EDC1E7">Schatz, George E.</mods:namePart>
<mods:affiliation id="FFE4A623713B93EF8AC67DCADCF82B33">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, MO, 63110 (United States) george. schatz @ mobot. org</mods:affiliation>
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<mods:namePart id="55C49BA09A903428DB2D693B6349F01C">Lowry II, Porter P.</mods:namePart>
<mods:affiliation id="40C45445AC0CD8B13400679D32158A55">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, MO, 63110 (United States) and Institut de Systématique, Évolution et Biodiversité, ISYEB Muséum national d'Histoire naturelle, Centre national de la Recherche scientifique, Sorbonne Université, École pratique des Hautes Études, Université des Antilles</mods:affiliation>
<mods:nameIdentifier id="B2C594A316ED9AB0BA519A3D2FB57426" type="email">pete.lowry@mobot.org</mods:nameIdentifier>
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<mods:title id="5B4B131EDF43E9A65491FC381FAB6EA7">Adansonia</mods:title>
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<mods:date id="098246CB89C1A1BE81845ED4CA91B045">2020</mods:date>
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<taxonomicName id="33FD7DE1FFB3070FFF4A12392F96453B" ID-CoL="9V73P" authority="Capuron ex G.E. Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB3070FFF4A12392FB4455B" bold="true" box="[179,429,1811,1838]" italics="true" pageId="4" pageNumber="203">Diospyros antakaranae</emphasis>
Capuron ex G.E. Schatz &amp; Lowry
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,
<taxonomicNameLabel id="DDBA670BFFB3070FFE63121E2FF5453B" box="[410,492,1844,1870]" pageId="4" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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(
<figureCitation id="6CC61AE7FFB3070FFE0212192C20453B" box="[507,569,1843,1870]" captionStart="FIG" captionStartId="5.[132,143,1687,1704]" captionTargetId="figure@5.[698,1267,787,1364]" captionTargetPageId="5" captionText="FIG. 1. — Diospyros antakaranae Capuron ex G.E. Schatz &amp; Lowry, sp. nov.: A, fruiting branch; B, detail of leaf venation; C, fruit. A-C, Service Forestier 8331 (P03975401). Illustration by A. Jouy. Scale bars: A, 4 cm; B, C, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3954010" httpUri="https://zenodo.org/record/3954010/files/figure.png" pageId="4" pageNumber="203">Fig. 1</figureCitation>
)
</paragraph>
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<subSubSection id="BCE755E9FFB3070FFF7D12412F154598" pageId="4" pageNumber="203" type="diagnosis">
<paragraph id="F4420662FFB3070FFF7D12412F154598" blockId="4.[132,776,1899,2029]" pageId="4" pageNumber="203">
<taxonomicName id="33FD7DE1FFB3070FFF7D12412CA845F6" authority="Capuron ex G.E. Schatz &amp; Lowry" authorityName="Capuron ex G. E. Schatz &amp; Lowry" authorityYear="2020" box="[132,689,1899,1923]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB3070FFF7D12412F4F45F6" box="[132,342,1899,1923]" italics="true" pageId="4" pageNumber="203">Diospyros antakaranae</emphasis>
Capuron ex G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB3070FFD4512412D1145F7" box="[700,776,1899,1922]" pageId="4" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from
<taxonomicName id="33FD7DE1FFB3070FFEA012AC2C6345E8" authority="Bojer ex A. DC." authorityName="A. DC." box="[345,634,1925,1949]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="species" species="squamosa">
<emphasis id="C689DA70FFB3070FFEA012AC2FCB45E8" box="[345,466,1926,1949]" italics="true" pageId="4" pageNumber="203">D. squamosa</emphasis>
Bojer ex A. DC.
</taxonomicName>
by its smaller, elliptic to narrowly elliptic leaves with a cuneate to attenuate base and mature fruit densely covered with appressed golden trichomes
<emphasis id="C689DA70FFB3070FFF7D12FF2E8D4599" box="[132,148,2005,2028]" italics="true" pageId="4" pageNumber="203">c.</emphasis>
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long.
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TYPUS. —
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.
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Region [Prov.
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, fr.,
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<collectorName id="590863B4FFB3070FFBFC12062A664531" box="[1029,1151,1836,1860]" pageId="4" pageNumber="203">Razafitsalama</collectorName>
1213
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(
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,
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[
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[
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[
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.
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. —
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<collectingCountry id="8CEA46F2FFB3070FFC4A12AF2A3845E8" box="[947,1057,1925,1949]" name="Madagascar" pageId="4" pageNumber="203">Madagascar</collectingCountry>
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.
<collectingCounty id="1D237EEEFFB3070FFBD312AF2ABC45E8" box="[1066,1189,1925,1949]" pageId="4" pageNumber="203">Diana Region</collectingCounty>
[Prov.
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]:
<collectingMunicipality id="14269C18FFB3070FFAA712AC2BB545E8" box="[1374,1452,1926,1949]" pageId="4" pageNumber="203">Ramena</collectingMunicipality>
,
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,
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345
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(
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,
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,
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,
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,
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<collectingDate id="9007D94AFFB3070FFC0C12902A7545A7" box="[1013,1132,1978,2002]" pageId="4" pageNumber="203" value="2011-12-29">29.XII.2011</collectingDate>
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, late
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fl.,
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<collectorName id="590863B4FFB3070FFB2012902B5B45A7" box="[1241,1346,1978,2002]" pageId="4" pageNumber="203">Nusbaumer</collectorName>
et al. 3284
</emphasis>
(
<collectionCode id="92EC9EA7FFB3070FFCCD12FF2D514598" box="[820,840,2005,2029]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="4" pageNumber="203" type="Herbarium">G</collectionCode>
,
<collectionCode id="92EC9EA7FFB3070FFCA812FF2D794599" box="[849,864,2005,2028]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="203" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB3070FFC9C12FE2DCD4599" box="[869,980,2004,2028]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="203" type="Herbarium">P01060531</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="44950C3FFFB3070EFC1412FF2F95456C" ID-GBIF-Occurrence="2816746333" collectingDate="2005-08-11" collectionCode="TAN, W" collectorName="Rakotonasolo &amp; Rapanarivo" country="Madagascar" lastPageId="5" lastPageNumber="204" location="Antsampano" pageId="4" pageNumber="203" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB3070FFC1412FF2A7F4598" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB3070FFC1412FF2A7F4598" box="[1005,1126,2005,2029]" country="Madagascar" name="Antsampano" pageId="4" pageNumber="203">Antsampano</location>
,
<date id="804320A2FFB3070FFB9712FE2AF44599" box="[1134,1261,2004,2028]" pageId="4" pageNumber="203" value="2005-08-11">
<collectingDate id="9007D94AFFB3070FFB9712FE2AF44599" box="[1134,1261,2004,2028]" pageId="4" pageNumber="203" value="2005-08-11">11.VIII.2005</collectingDate>
</date>
, bud,
<emphasis id="C689DA70FFB3070EFADA12FF2F3B456C" italics="true" lastPageId="5" lastPageNumber="204" pageId="4" pageNumber="203">
<collectorName id="590863B4FFB3070FFADA12FF2B8E4598" box="[1315,1431,2005,2029]" pageId="4" pageNumber="203">Rakotonasolo</collectorName>
&amp;
<collectorName id="590863B4FFB2070EFF7D12282EF4456C" box="[132,237,1794,1817]" pageId="5" pageNumber="204">Rapanarivo</collectorName>
1036
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFED7122B2F7A456C" box="[302,355,1793,1817]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="5" pageNumber="204" type="Herbarium">TAN</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFE93122B2F9D456C" box="[362,388,1793,1817]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="204" type="Herbarium">W</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFE6C122B2CC94541" ID-GBIF-Occurrence="2816746352" collectingDate="2011-12-18" collectionCode="G, P" collectorName="Ramandimbimanana &amp; Randimbiarison" country="Madagascar" location="Montagne d'Ambre" pageId="5" pageNumber="204" specimenCode="P01060546" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFE6C122B2C56456C" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFE6C122B2C56456C" box="[405,591,1793,1817]" country="Madagascar" name="Montagne d'Ambre" pageId="5" pageNumber="204">Montagne dAmbre</location>
,
<date id="804320A2FFB2070EFDAE122B2CD7456C" box="[599,718,1793,1817]" pageId="5" pageNumber="204" value="2011-12-18">
<collectingDate id="9007D94AFFB2070EFDAE122B2CD7456C" box="[599,718,1793,1817]" pageId="5" pageNumber="204" value="2011-12-18">18.XII.2011</collectingDate>
</date>
,
<specimenCount id="E2FBCDEBFFB2070EFD2E122B2CF3456C" box="[727,746,1793,1817]" pageId="5" pageNumber="204" type="male"></specimenCount>
fl.,
<emphasis id="C689DA70FFB2070EFF7D12312C0F4541" box="[132,534,1819,1844]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFF7D12312F5D4546" box="[132,324,1819,1843]" pageId="5" pageNumber="204">Ramandimbimanana</collectorName>
&amp;
<collectorName id="590863B4FFB2070EFEA612312FF74546" box="[351,494,1819,1843]" pageId="5" pageNumber="204">Randimbiarison</collectorName>
181
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFDDB12362C2F4541" box="[546,566,1820,1844]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="204" type="Herbarium">G</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFDC712362C544546" box="[574,589,1820,1843]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFDAB12312CA64546" box="[594,703,1819,1843]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P01060546</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFD2112362F22451D" ID-GBIF-Occurrence="2816746351" collectingDate="2011-12-20" collectionCode="G, P" collectorName="Randimbiarison &amp; Ramandimbimanana" country="Madagascar" location="Montagne d'Ambre" pageId="5" pageNumber="204" specimenCode="P01060533" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same locality,
<date id="804320A2FFB2070EFF37121C2F5A453B" box="[206,323,1846,1870]" pageId="5" pageNumber="204" value="2011-12-20">
<collectingDate id="9007D94AFFB2070EFF37121C2F5A453B" box="[206,323,1846,1870]" pageId="5" pageNumber="204" value="2011-12-20">20.XII.2011</collectingDate>
</date>
, ♂ fl.,
<emphasis id="C689DA70FFB2070EFE86121C2D1F453B" box="[383,774,1846,1870]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFE86121C2C17453B" box="[383,526,1846,1870]" pageId="5" pageNumber="204">Randimbiarison</collectorName>
&amp;
<collectorName id="590863B4FFB2070EFDD0121C2CF3453B" box="[553,746,1846,1870]" pageId="5" pageNumber="204">Ramandimbimanana</collectorName>
40
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFF75127A2EB9451D" box="[140,160,1872,1896]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="204" type="Herbarium">G</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFF51127B2EAE451D" box="[168,183,1873,1896]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFF44127A2F32451D" box="[189,299,1872,1896]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P01060533</specimenCode>
])
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFEBD127A2FB745F6" ID-GBIF-Occurrence="2816746330" collectingDate="2005-01-25" collectionCode="I, MO, P, TAN" collectorName="Randrianarivelo" country="Madagascar" location="Mahavanona" pageId="5" pageNumber="204" specimenCode="P01031022" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFEBD127A2FA6451D" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFEBD127A2FA6451D" box="[324,447,1872,1896]" country="Madagascar" name="Mahavanona" pageId="5" pageNumber="204">Mahavanona</location>
,
<date id="804320A2FFB2070EFE3F127A2C3D451D" box="[454,548,1872,1896]" pageId="5" pageNumber="204" value="2005-01-25">
<collectingDate id="9007D94AFFB2070EFE3F127A2C3D451D" box="[454,548,1872,1896]" pageId="5" pageNumber="204" value="2005-01-25">
25.
<collectionCode id="92EC9EA7FFB2070EFE1A127A2FF2451D" box="[483,491,1872,1896]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="5" pageNumber="204" type="Herbarium">I</collectionCode>
.2005
</collectingDate>
</date>
, fr.,
<emphasis id="C689DA70FFB2070EFDB0127A2EBE45F6" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFDB0127A2CC0451D" box="[585,729,1872,1896]" pageId="5" pageNumber="204">Randrianarivelo</collectorName>
et al. 163
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFF4A12412EC745F6" box="[179,222,1899,1923]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="204" type="Herbarium">MO</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFF1F12412EEC45F7" box="[230,245,1899,1922]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFF0312412F7F45F6" box="[250,358,1899,1923]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P01031022</specimenCode>
],
<collectionCode id="92EC9EA7FFB2070EFE8C12412FB145F6" box="[373,424,1899,1923]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="5" pageNumber="204" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFE4F12412F2E45CD" ID-GBIF-Occurrence="2816746317" collectingDate="1952-08-31" collectionCode="G, K, MO, P, TEF" collectorName="Service Forestier &amp; Randriamanatina" country="Madagascar" location="Montagne d'Ambre" pageId="5" pageNumber="204" specimenCode="P03974935, P03974936" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFE4F12412C7045F6" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFE4F12412C7045F6" box="[438,617,1899,1923]" country="Madagascar" name="Montagne d'Ambre" pageId="5" pageNumber="204">Montagne dAmbre</location>
,
<date id="804320A2FFB2070EFD8812412CF545F6" box="[625,748,1899,1923]" pageId="5" pageNumber="204" value="1952-08-31">
<collectingDate id="9007D94AFFB2070EFD8812412CF545F6" box="[625,748,1899,1923]" pageId="5" pageNumber="204" value="1952-08-31">31.VIII.1952</collectingDate>
</date>
,
<specimenCount id="E2FBCDEBFFB2070EFD0D12412D1E45F6" box="[756,775,1899,1923]" pageId="5" pageNumber="204" type="female"></specimenCount>
fl.,
<emphasis id="C689DA70FFB2070EFF5912AC2C1545E8" box="[160,524,1925,1949]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFF5912AC2F3E45E8" box="[160,295,1926,1949]" pageId="5" pageNumber="204">Service Forestier</collectorName>
(
<collectorName id="590863B4FFB2070EFEC812AF2FC845E8" box="[305,465,1925,1949]" pageId="5" pageNumber="204">Randriamanatina</collectorName>
) 5795
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFDEE12AF2C3245E8" box="[535,555,1925,1949]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="204" type="Herbarium">G</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFDCA12AC2C5C45E8" box="[563,581,1926,1949]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="204" type="Herbarium">K</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFDB412AF2C6145E8" box="[589,632,1925,1949]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="204" type="Herbarium">MO</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFD7912AC2C9645E8" box="[640,655,1926,1949]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFD6D12AF2D1945E8" box="[660,768,1925,1949]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P03974935</specimenCode>
,
<specimenCode id="A45BAE19FFB2070EFF7D128A2EEB45CD" box="[132,242,1952,1976]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P03974936</specimenCode>
],
<collectionCode id="92EC9EA7FFB2070EFEFB128A2F2945CD" box="[258,304,1952,1976]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="5" pageNumber="204" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFEC6128A2C0A45A7" ID-GBIF-Occurrence="2816746337" collectingDate="1954-02-19" collectionCode="MO, P, TEF" collectorName="Service Forestier" country="Madagascar" location="Massif d'Ampitiliantsambo" pageId="5" pageNumber="204" specimenCode="P03975401" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFEC6128A2C2145CD" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFEC6128A2C2145CD" box="[319,568,1952,1976]" country="Madagascar" name="Massif d'Ampitiliantsambo" pageId="5" pageNumber="204">Massif dAmpitiliantsambo</location>
,
<date id="804320A2FFB2070EFDB9128A2CBC45CD" box="[576,677,1952,1976]" pageId="5" pageNumber="204" value="1954-02-19">
<collectingDate id="9007D94AFFB2070EFDB9128A2CBC45CD" box="[576,677,1952,1976]" pageId="5" pageNumber="204" value="1954-02-19">19.II.1954</collectingDate>
</date>
, fr.,
<emphasis id="C689DA70FFB2070EFD33128A2F1D45A7" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFD33128A2ED745A7" pageId="5" pageNumber="204">Service Forestier</collectorName>
8331
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFEF612902F2545A7" box="[271,316,1978,2002]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="204" type="Herbarium">MO</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFEBD12912F4845A7" box="[324,337,1979,2002]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFEA412902FD545A7" box="[349,460,1978,2002]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P03975401</specimenCode>
],
<collectionCode id="92EC9EA7FFB2070EFE2412902C1545A7" box="[477,524,1978,2002]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="5" pageNumber="204" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFDE312902AD14562" ID-GBIF-Occurrence="2816746328" collectingDate="1966-01-26" collectionCode="G, MO, P, TEF, W" collectorName="Service Forestier" country="Madagascar" location="Massif de l'Ambongoabo" municipality="Diego-Suarez" pageId="5" pageNumber="204" specimenCode="P03975365, P03975366" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFDE312902D1A45A7" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFDE312902D1A45A7" box="[538,771,1978,2002]" country="Madagascar" municipality="Diego-Suarez" name="Massif de l'Ambongoabo" pageId="5" pageNumber="204">Massif de lAmbongoabo</location>
, à louest de
<collectingMunicipality id="14269C18FFB2070EFF0F12FF2F6F4598" box="[246,374,2005,2029]" pageId="5" pageNumber="204">Diégo-Suarez</collectingMunicipality>
,
<date id="804320A2FFB2070EFE7812FF2FC64598" box="[385,479,2005,2029]" pageId="5" pageNumber="204" value="1966-01-26">
<collectingDate id="9007D94AFFB2070EFE7812FF2FC64598" box="[385,479,2005,2029]" pageId="5" pageNumber="204" value="1966-01-26">26.I.1966</collectingDate>
</date>
, fr.,
<emphasis id="C689DA70FFB2070EFDF512FF2CC64598" box="[524,735,2005,2029]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFDF512FF2C834599" box="[524,666,2005,2028]" pageId="5" pageNumber="204">Service Forestier</collectorName>
24457
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFD1712FF2D1A4598" box="[750,771,2005,2029]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="5" pageNumber="204" type="Herbarium">G</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFCD413D52D404562" box="[813,857,1791,1815]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="204" type="Herbarium">MO</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFC9A122A2D6B4562" box="[867,882,1792,1815]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P</collectionCode>
[
<specimenCode id="A45BAE19FFB2070EFC8113D52DFE4562" box="[888,999,1791,1815]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P03975365</specimenCode>
,
<specimenCode id="A45BAE19FFB2070EFC0B13D52A784562" box="[1010,1121,1791,1815]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="5" pageNumber="204" type="Herbarium">P03975366</specimenCode>
],
<collectionCode id="92EC9EA7FFB2070EFB8D122A2A864562" box="[1140,1183,1792,1815]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="5" pageNumber="204" type="Herbarium">TEF</collectionCode>
,
<collectionCode id="92EC9EA7FFB2070EFB5E13D52AD84562" box="[1191,1217,1791,1815]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="204" type="Herbarium">W</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFB2B122A2AC24547" ID-GBIF-Occurrence="2816746304" collectingDate="1979-11-12" collectionCode="TEF" collectorName="Service Forestier" country="Madagascar" location="Montagne des Francais" pageId="5" pageNumber="204" specimenCount="1" typeStatus="paratype">
<location id="F12250B9FFB2070EFB2B122A2BB54562" LSID="urn:lsid:plazi:treatment:7C54B774FFB3070CFF4A12392EE74306:F12250B9FFB2070EFB2B122A2BB54562" box="[1234,1452,1791,1816]" country="Madagascar" name="Montagne des Francais" pageId="5" pageNumber="204">Montagne des Français</location>
,
<date id="804320A2FFB2070EFCD412302D834547" box="[813,922,1818,1842]" pageId="5" pageNumber="204" value="1979-11-12">
<collectingDate id="9007D94AFFB2070EFCD412302D834547" box="[813,922,1818,1842]" pageId="5" pageNumber="204" value="1979-11-12">12.XI.1979</collectingDate>
</date>
, ster.,
<emphasis id="C689DA70FFB2070EFC2912302A834547" box="[976,1178,1818,1842]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFC2912302A434544" box="[976,1114,1818,1841]" pageId="5" pageNumber="204">Service Forestier</collectorName>
29601
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFB5C12302ACD4547" box="[1189,1236,1818,1842]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="5" pageNumber="204" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="44950C3FFFB2070EFB1A12302A6B4539" ID-GBIF-Occurrence="2816746332" collectingDate="1979-11-12" collectionCode="TEF" collectorName="Service Forestier" country="Madagascar" location="Montagne des Francais" pageId="5" pageNumber="204" specimenCount="1" typeStatus="paratype">
same locality and date, ster.,
<emphasis id="C689DA70FFB2070EFCA4121E2A374539" box="[861,1070,1844,1868]" italics="true" pageId="5" pageNumber="204">
<collectorName id="590863B4FFB2070EFCA4121E2DF24539" box="[861,1003,1844,1868]" pageId="5" pageNumber="204">Service Forestier</collectorName>
29602
</emphasis>
(
<collectionCode id="92EC9EA7FFB2070EFBC2121E2A724539" box="[1083,1131,1844,1868]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="5" pageNumber="204" type="Herbarium">TEF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="A08256EAFFB2070EFF7D13BD2CD844CB" ID-DOI="http://doi.org/10.5281/zenodo.3954010" ID-Zenodo-Dep="3954010" httpUri="https://zenodo.org/record/3954010/files/figure.png" pageId="5" pageNumber="204" targetBox="[131,1456,211,1651]" targetPageId="5">
<paragraph id="F4420662FFB2070EFF7D13BD2CD844CB" blockId="5.[132,1456,1687,1726]" pageId="5" pageNumber="204">
FIG. 1. —
<taxonomicName id="33FD7DE1FFB2070EFF2F13BD2CAD44DD" authority="Capuron ex G.E. Schatz &amp; Lowry" authorityName="Capuron ex G. E. Schatz &amp; Lowry" authorityYear="2020" box="[214,692,1687,1704]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="204" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB2070EFF2F13BD2F8D44DD" box="[214,404,1687,1704]" italics="true" pageId="5" pageNumber="204">Diospyros antakaranae</emphasis>
Capuron ex G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB2070EFD4413BD2D1D44DD" box="[701,772,1687,1704]" pageId="5" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="C689DA70FFB2070EFCF213BD2D0144DD" bold="true" box="[779,792,1687,1704]" pageId="5" pageNumber="204">A</emphasis>
, fruiting branch;
<emphasis id="C689DA70FFB2070EFC5013BD2DAF44DD" bold="true" box="[937,950,1687,1704]" pageId="5" pageNumber="204">B</emphasis>
, detail of leaf venation;
<emphasis id="C689DA70FFB2070EFB7B13BD2A8944DD" bold="true" box="[1154,1168,1687,1704]" pageId="5" pageNumber="204">C</emphasis>
, fruit. A-C,
<emphasis id="C689DA70FFB2070EFB0A13BD2BA944DD" box="[1267,1456,1687,1704]" italics="true" pageId="5" pageNumber="204">Service Forestier 8331</emphasis>
(P03975401). Illustration by A. Jouy. Scale bars: A, 4 cm; B, C, 1 cm.
</paragraph>
</caption>
<subSubSection id="BCE755E9FFB2070EFCD412402DF54598" pageId="5" pageNumber="204" type="etymology">
<paragraph id="F4420662FFB2070EFCD412402DF54598" blockId="5.[813,1456,1898,2029]" pageId="5" pageNumber="204">
ETYMOLOGY. — The name
<taxonomicName id="33FD7DE1FFB2070EFBC812412B1845F6" authority="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc, 2020" authorityName="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc" authorityYear="2020" box="[1073,1281,1899,1923]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="204" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB2070EFBC812412B1845F6" box="[1073,1281,1899,1923]" italics="true" pageId="5" pageNumber="204">Diospyros antakaranae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB2070EFAF512412B4C45F7" box="[1292,1365,1899,1922]" pageId="5" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
was written by René Capuron on specimens at P and TEF, but was never published. He appears to have based this name on the Antakarana ethnic group of northern
<collectingCountry id="8CEA46F2FFB2070EFBE712902A8845A7" box="[1054,1169,1978,2002]" name="Madagascar" pageId="5" pageNumber="204">Madagascar</collectingCountry>
, who live in the area inhabited by this new species.
</paragraph>
</subSubSection>
<subSubSection id="BCE755E9FFB1070DFF7D15F32F33432E" pageId="6" pageNumber="205" type="distribution">
<paragraph id="F4420662FFB1070DFF7D15F32F33432E" blockId="6.[132,776,217,347]" pageId="6" pageNumber="205">
DISTRIBUTION AND ECOLOGY. —
<taxonomicName id="33FD7DE1FFB1070DFE2F15F02CB54284" authority="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc, 2020" authorityName="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc" authorityYear="2020" box="[470,684,217,241]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFE2F15F02CB54284" box="[470,684,217,241]" italics="true" pageId="6" pageNumber="205">Diospyros antakaranae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFD4215F02D114284" box="[699,776,218,241]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
occurs in dry forest in the far north of
<collectingCountry id="8CEA46F2FFB1070DFE0C15DE2C704379" box="[501,617,244,268]" name="Madagascar" pageId="6" pageNumber="205">Madagascar</collectingCountry>
, extending from the northern parts of Montagne dAmbre and nearby areas situated slightly farther north to Montagne des Français, from
<quantity id="3305AB87FFB1070DFD3D14032EDC432E" metricMagnitude="2" metricUnit="m" metricValue="5.08" metricValueMax="9.0" metricValueMin="1.16" pageId="6" pageNumber="205" unit="m" value="508.0" valueMax="900.0" valueMin="116.0">116 to 900 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="BCE755E9FFB1070DFF7D14582EF843B5" pageId="6" pageNumber="205" type="biology_ecology">
<paragraph id="F4420662FFB1070DFF7D14582EF843B5" blockId="6.[132,775,370,448]" pageId="6" pageNumber="205">
PHENOLOGY. —
<taxonomicName id="33FD7DE1FFB1070DFED014592FE443FE" authority="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc, 2020" authorityName="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc" authorityYear="2020" box="[297,509,371,395]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFED014592FE443FE" box="[297,509,371,395]" italics="true" pageId="6" pageNumber="205">Diospyros antakaranae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFDF014592C4D43FF" box="[521,596,371,394]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
has been recorded in flower from August to December, and in fruit in January and February.
</paragraph>
</subSubSection>
<subSubSection id="BCE755E9FFB1070DFF7D14FD2CDE439A" box="[132,711,471,495]" pageId="6" pageNumber="205" type="vernacular_names">
<paragraph id="F4420662FFB1070DFF7D14FD2CDE439A" blockId="6.[132,711,471,495]" box="[132,711,471,495]" pageId="6" pageNumber="205">
VERNACULAR NAME. — Mampingo (
<emphasis id="C689DA70FFB1070DFE1214F22CA0439A" box="[491,697,471,495]" italics="true" pageId="6" pageNumber="205">Service Forestier 5795</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="BCE755E9FFB1070DFF7D172C2D1F4178" pageId="6" pageNumber="205" type="conservation">
<paragraph id="F4420662FFB1070DFF7D172C2D1F4178" blockId="6.[132,776,518,781]" pageId="6" pageNumber="205">
CONSERVATION STATUS. —
<taxonomicName id="33FD7DE1FFB1070DFE5D17222C98406A" authority="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc, 2020" authorityName="Schatz &amp; Lowry Ii &amp; Dc &amp; Dc" authorityYear="2020" box="[420,641,519,543]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFE5D17222C98406A" box="[420,641,519,543]" italics="true" pageId="6" pageNumber="205">Diospyros antakaranae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFD69172D2CC6406B" box="[656,735,519,542]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
has a geographic range in the form of an EOO of
<quantity id="3305AB87FFB1070DFD9517082CA4404C" box="[620,701,545,570]" metricMagnitude="5" metricUnit="m" metricValue="4.21" pageId="6" pageNumber="205" unit="km" value="421.0">421 km</quantity>
<superScript id="0388AB2AFFB1070DFD44170B2CDF4047" attach="left" box="[701,710,545,562]" fontSize="7" pageId="6" pageNumber="205">2</superScript>
and a minimum AOO of
<quantity id="3305AB87FFB1070DFEB117162F904021" box="[328,393,572,596]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="205" unit="km" value="32.0">32 km</quantity>
2. It is present in the protected areas of Montagne dAmbre and Montagne des Français. It is threatened by fire, grazing, and development outside of these protected areas, and likely also within them. With respect to the principal threat of fire, which will result in projected continuing decline in habitat quality,
<taxonomicName id="33FD7DE1FFB1070DFF2817EB2CAD40AC" authority="Capuron ex G.E. Schatz &amp; Lowry" authorityName="Capuron ex G. E. Schatz &amp; Lowry" authorityYear="2020" box="[209,692,704,729]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="antakaranae" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFF2817EB2F7C40AD" box="[209,357,704,728]" italics="true" pageId="6" pageNumber="205">D. antakaranae</emphasis>
Capuron ex G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFD4717EB2D1140AD" box="[702,776,705,728]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
exists at seven locations, and can therefore be assessed for its risk of extinction as Vulnerable [VU B1ab(iii)+2ab(iii)] (
<bibRefCitation id="906C7B93FFB1070DFD8617DF2CE04178" author="IUCN" box="[639,761,757,781]" firstAuthor="IUCN" pageId="6" pageNumber="205" refId="ref13513" refString="IUCN 2012. - IUCN Red List Categories and Criteria: Version 3.1. Second edition. IUCN, Gland, Switzerland and Cambridge, United Kingdom." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BCE755E9FFB1070DFF7D161C2B204538" pageId="6" pageNumber="205" type="description">
<paragraph id="F4420662FFB1070DFF7D161C2F0C413B" blockId="6.[132,775,822,2030]" box="[132,277,822,848]" pageId="6" pageNumber="205">DESCRIPTION</paragraph>
<paragraph id="F4420662FFB1070DFF7D167D2B204538" blockId="6.[132,775,822,2030]" lastBlockId="6.[813,1455,1397,1869]" pageId="6" pageNumber="205">
Shrub to tree
<quantity id="3305AB87FFB1070DFEE0167D2F424107" box="[281,347,855,882]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="6" pageNumber="205" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">3-7 m</quantity>
tall,
<quantity id="3305AB87FFB1070DFE6B167D2FF44107" box="[402,493,855,882]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="4.0" metricValueMin="0.6" pageId="6" pageNumber="205" unit="cm" value="23.0" valueMax="40.0" valueMin="6.0">6-40 cm</quantity>
DBH. Young stems whitish, slender,
<quantity id="3305AB87FFB1070DFEFE165D2F5E41E4" box="[263,327,887,913]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="205" unit="mm" value="1.0">1 mm</quantity>
in diam., sparsely covered with semi-erect gray trichomes to
<quantity id="3305AB87FFB1070DFEBD16BD2F9D41C4" box="[324,388,919,945]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="205" unit="mm" value="2.0">2 mm</quantity>
long, glabrescent. Lamina 1.9-5.0 ×
<quantity id="3305AB87FFB1070DFF7D169C2EE441A4" box="[132,253,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.5" metricValueMin="0.8" pageId="6" pageNumber="205" unit="cm" value="1.65" valueMax="2.5" valueMin="0.8">0.8-2.5 cm</quantity>
, narrowly elliptic to elliptic, coriaceous, glabrous above and below, base acute to cuneate to attenuate, rarely obtuse, margin micro-revolute and initially sparsely ciliate with grayish trichomes
<quantity id="3305AB87FFB1070DFE79113C2FCC4644" box="[384,469,1046,1073]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="205" unit="mm" value="0.7">0.7 mm</quantity>
long, apex acuminate, occasionally slightly falcate, the acumen to
<quantity id="3305AB87FFB1070DFDD7111C2C644624" box="[558,637,1078,1105]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="205" unit="cm" value="1.1">1.1 cm</quantity>
, the very tip rounded, midvein slightly impressed above, raised below, initially covered with sparse grayish trichomes 0.7. mm long below, venation weakly brochidodromous, 7-9 secondary veins per side, completely obscure above, slightly raised below; petiole
<quantity id="3305AB87FFB1070DFEF111FC2F7B4685" box="[264,354,1238,1264]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="205" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
, &lt;
<quantity id="3305AB87FFB1070DFE7B11FC2FDB4685" box="[386,450,1238,1264]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="205" unit="mm" value="1.0">1 mm</quantity>
in diam., terete to slightly flat adaxially, sparsely covered with semi-erect gray trichomes to
<quantity id="3305AB87FFB1070DFF7D103C2EDA4745" box="[132,195,1302,1328]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="205" unit="mm" value="2.0">2 mm</quantity>
long, glabrescent. Male flowers solitary in axils of fallen leaves, enclosed within 5 distichous, overlapping, cucullate bracts, becoming progressively larger from base to apex of the pedicel, the largest broadly ovate to round, 3.5 ×
<quantity id="3305AB87FFB1070DFD46105F2D1A47E5" box="[703,771,1397,1424]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="205" unit="mm" value="4.0">4 mm</quantity>
, sparsely covered with minute appressed trichomes to glabrous, the margins minutely ciliate; pedicel
<quantity id="3305AB87FFB1070DFDFD109F2C4047BA" box="[516,601,1461,1487]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="205" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
long,
<quantity id="3305AB87FFB1070DFD60109F2CF547BA" box="[665,748,1461,1488]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="205" unit="mm" value="0.5">0.5 mm</quantity>
in diam.; calyx urceolate,
<quantity id="3305AB87FFB1070DFE8510FF2FA4479A" box="[380,445,1493,1519]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="205" unit="mm" value="3.0">3 mm</quantity>
tall,
<quantity id="3305AB87FFB1070DFE0D10FF2C55479A" box="[500,588,1493,1520]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="205" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
in diam. at apex, with 5 triangular lobes, 0.5 ×
<quantity id="3305AB87FFB1070DFE2810DF2C0F447A" box="[465,534,1525,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="205" unit="mm" value="1.0">1 mm</quantity>
, densely covered with semi-appressed golden trichomes
<quantity id="3305AB87FFB1070DFE06133E2C68445A" box="[511,625,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="205" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
long; corolla white in vivo, tubular, fused portion
<quantity id="3305AB87FFB1070DFDF5131F2C52443A" box="[524,587,1589,1615]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="205" unit="mm" value="3.0">3 mm</quantity>
long, with 5 free, overlapping and spreading, depressed ovate lobes, 2 ×
<quantity id="3305AB87FFB1070DFD39137E2D1A441A" box="[704,771,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="205" unit="mm" value="4.0">4 mm</quantity>
, the tube glabrous to densely covered with minute appressed trichomes at apex outside, the lobes densely covered with short (
<quantity id="3305AB87FFB1070DFF31139E2F0744BA" box="[200,286,1716,1743]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="205" unit="mm" value="0.2">0.2 mm</quantity>
) appressed trichomes outside, sparsely covered with minute appressed trichomes inside; stamens
<emphasis id="C689DA70FFB1070DFD5F13FF2CAF449B" box="[678,694,1749,1774]" italics="true" pageId="6" pageNumber="205">c.</emphasis>
15-20, anthers sessile, 2.5-3 ×
<quantity id="3305AB87FFB1070DFE8D13DE2FD2457B" box="[372,459,1780,1807]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="205" unit="mm" value="0.5">0.5 mm</quantity>
. Female flowers solitary in the axils of leaves, enclosed within 5-7 distichous, overlapping cucullate bracts, collectively broadly ovate, 2-3 ×
<quantity id="3305AB87FFB1070DFD5F121E2D1A453B" box="[678,771,1844,1870]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="205" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2-4 mm</quantity>
, becoming progressively larger from base to apex, rounded to narrowly ovate, densely covered with short, matted appressed golden trichomes outside except glabrous towards the ciliate margins, glabrous inside; pedicel
<quantity id="3305AB87FFB1070DFE1012992C4F45BB" box="[489,598,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="5.0" metricValueMin="1.4" pageId="6" pageNumber="205" unit="mm" value="3.2" valueMax="5.0" valueMin="1.4">1.4-5 mm</quantity>
long,
<quantity id="3305AB87FFB1070DFD6312992D1E45BB" box="[666,775,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="6" pageNumber="205" unit="mm" value="1.6" valueMax="2.0" valueMin="1.2">1.2-2 mm</quantity>
in diam., with a dense ring of trichomes at apex subtending the receptacle; calyx urceolate, 4-merous,
<quantity id="3305AB87FFB1070DFB1F105F2B2547E5" box="[1254,1340,1397,1424]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="205" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
tall,
<quantity id="3305AB87FFB1070DFA89105C2BB647E5" box="[1392,1455,1398,1424]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="205" unit="mm" value="7.0">7 mm</quantity>
in diam. at apex, the 4 lobes triangular, 5 ×
<quantity id="3305AB87FFB1070DFB0F10BF2B5647C5" box="[1270,1359,1429,1456]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="205" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
, margins slightly revolute, densely covered with semi-appressed golden trichomes
<quantity id="3305AB87FFB1070DFC4610FF2A34479A" box="[959,1069,1493,1520]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="205" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
long; corolla not seen; ovary ovoid,
<quantity id="3305AB87FFB1070DFCD410DF2D77447A" box="[813,878,1525,1551]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="205" unit="mm" value="4.0">4 mm</quantity>
tall, densely covered with appressed white trichomes, stigma with 4 bifid branches,
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long. Fruit borne on an accrescent receptacle, to
<quantity id="3305AB87FFB1070DFBC1131E2A60443A" box="[1080,1145,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="205" unit="mm" value="5.0">5 mm</quantity>
long, 3-5-
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in diam., with a ring of downward appressed golden grayish trichomes around the base; fruit
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tall,
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in diam., slightly flattened globose, the stylar remnant
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long, moderately to densely covered towards the apex with appressed golden trichomes
<emphasis id="C689DA70FFB1070DFC1513FF2DE4449B" box="[1004,1021,1749,1774]" italics="true" pageId="6" pageNumber="205">c.</emphasis>
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long, calyx in fruit covering
<emphasis id="C689DA70FFB1070DFA8813FF2B9B449B" box="[1393,1410,1749,1774]" italics="true" pageId="6" pageNumber="205">c.</emphasis>
one half of fruit,
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tall, the lobes 10 ×
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, triangular, the apex reflexed, margins of the lobes slightly revolute, covered with dense, matted, appressed golden trichomes.
</paragraph>
</subSubSection>
<caption id="A08256EAFFB1070DFCD4118C2DEF475D" ID-DOI="http://doi.org/10.5281/zenodo.3953002" ID-Zenodo-Dep="3953002" httpUri="https://zenodo.org/record/3953002/files/figure.png" pageId="6" pageNumber="205" targetBox="[811,1457,213,1149]" targetPageId="6">
<paragraph id="F4420662FFB1070DFCD4118C2DEF475D" blockId="6.[813,1457,1190,1320]" pageId="6" pageNumber="205">
FIG. 2. — Geographic distribution of
<taxonomicName id="33FD7DE1FFB1070DFBA4118C2D7A46BB" authority="G.E. Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="betamponensis" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFBA4118C2B2C46C2" box="[1117,1333,1190,1207]" italics="true" pageId="6" pageNumber="205">Diospyros betamponensis</emphasis>
G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFC9311972DB746BB" box="[874,942,1213,1230]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
(.),
<taxonomicName id="33FD7DE1FFB1070DFC2D11972B5946BB" authority="G.E. Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" box="[980,1344,1213,1230]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="callmanderi" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFC2D11972A9246BB" box="[980,1163,1213,1230]" italics="true" pageId="6" pageNumber="205">Diospyros callmanderi</emphasis>
G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFAB111972B9546BB" box="[1352,1420,1213,1230]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="C689DA70FFB1070DFA6D11962BBC46B8" bold="true" box="[1428,1445,1212,1229]" pageId="6" pageNumber="205">🞧</emphasis>
),
<taxonomicName id="33FD7DE1FFB1070DFCD411F92A884691" authority="G.E. Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" box="[813,1169,1235,1252]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="darainensis" status="sp.nov.">
<emphasis id="C689DA70FFB1070DFCD411F92DFB4691" box="[813,994,1235,1252]" italics="true" pageId="6" pageNumber="205">Diospyros darainensis</emphasis>
G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFB6E11F92AC34691" box="[1175,1242,1235,1252]" pageId="6" pageNumber="205" rank="species">sp.nov.</taxonomicNameLabel>
(
<emphasis id="C689DA70FFB1070DFB1B11F92AEA4691" bold="true" box="[1250,1267,1235,1252]" pageId="6" pageNumber="205"></emphasis>
),
<taxonomicName id="33FD7DE1FFB1070DFAF811F92DCE468E" authority="G.E.Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="phillipsonii" status="sp.nov.">
<emphasis id="C689DA70FFB1070DFAF811F92BB64691" box="[1281,1455,1235,1252]" italics="true" pageId="6" pageNumber="205">Diospyros phillipsonii</emphasis>
G.E.Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFC2511C02A06468E" box="[988,1055,1258,1275]" pageId="6" pageNumber="205" rank="species">sp.nov.</taxonomicNameLabel>
(·), and
<taxonomicName id="33FD7DE1FFB1070DFB9C11C02BB4468E" authority="G.E. Schatz &amp; Lowry" authorityName="G. E. Schatz &amp; Lowry" authorityYear="2020" box="[1125,1453,1258,1275]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="205" phylum="Tracheophyta" rank="species" species="sennenii" status="sp. nov.">
<emphasis id="C689DA70FFB1070DFB9C11C02AE6468E" box="[1125,1279,1258,1275]" italics="true" pageId="6" pageNumber="205">Diospyros sennenii</emphasis>
G.E. Schatz &amp; Lowry
</taxonomicName>
,
<taxonomicNameLabel id="DDBA670BFFB1070DFCD4102B2D684767" box="[813,881,1281,1298]" pageId="6" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="C689DA70FFB1070DFC83102A2D954764" bold="true" box="[890,908,1280,1297]" pageId="6" pageNumber="205"></emphasis>
), mapped on the bioclimatic regions of Madagascar (after
<bibRefCitation id="906C7B93FFB1070DFA8F102B2D4E475D" author="Cornet" firstAuthor="Cornet" pageId="6" pageNumber="205" refId="ref13238" refString="CORNET A. 1974. - Essai de cartographie bioclimatique a Madagascar. Notice Explicative de l'ORSTOM No. 55." type="book" year="1974">Cornet 1974</bibRefCitation>
; see
<bibRefCitation id="906C7B93FFB1070DFC7A103D2DF5475D" author="Schatz" box="[899,1004,1303,1320]" firstAuthor="Schatz" pageId="6" pageNumber="205" pagination="1 - 9" refId="ref13689" refString="SCHATZ G. E. 2000. - Endemism in the Malagasy tree flora, in LOURENCO W. R. &amp; GOODMAN S. M. (eds), Diversity and Endemism in Madagascar. Memoires de la Societe biogeographique de Paris. Societe de Biogeographie, Paris: 1 - 9." type="book chapter" year="2000">Schatz 2000</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="BCE755E9FFB1070CFCD412592EE74306" lastPageId="7" lastPageNumber="206" pageId="6" pageNumber="205" type="discussion">
<paragraph id="F4420662FFB1070DFCD412592D6145FF" blockId="6.[813,1455,1907,2030]" box="[813,888,1907,1933]" pageId="6" pageNumber="205">NOTES</paragraph>
<paragraph id="F4420662FFB1070CFCD412B92EE74306" blockId="6.[813,1455,1907,2030]" lastBlockId="7.[132,775,217,371]" lastPageId="7" lastPageNumber="206" pageId="6" pageNumber="205">
This species can be recognized from other members of the Squamosa group by its relatively small, narrowly elliptic to elliptic leaves with a cuneate to attenuate base, and its mature fruit covered with appressed golden trichomes
<emphasis id="C689DA70FFB0070CFD8D15F02C9C4286" box="[628,645,218,243]" italics="true" pageId="7" pageNumber="206">c.</emphasis>
<quantity id="3305AB87FFB0070CFD7315F32CD34281" box="[650,714,217,244]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="206" unit="mm" value="1.0">1 mm</quantity>
long. Because individuals of this species can become large trees and may therefore be targeted as a source of ebony wood, only general locality information has been provided for the cited collections.
</paragraph>
</subSubSection>
</treatment>
</document>