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<document id="5A10A9193091327643CA90C91CF08727" ID-CLB-Dataset="32088" ID-DOI="10.11646/zootaxa.4320.2.8" ID-GBIF-Dataset="5d3d2c6f-5260-4279-b6ae-dbebe61f8316" ID-ISSN="1175-5326" ID-Zenodo-Dep="891908" ID-ZooBank="1E026A12-Aca9-4A9D-9245-2Fe09E582Fc6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505465222306" checkinUser="plazi" docAuthor="Marusik, Yuri M." docDate="2017" docId="76614023B3280C2455B52292FDC2575D" docLanguage="en" docName="zootaxa.4320.2.8.pdf.imf" docOrigin="Zootaxa 4320 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dzhungarocosa Marusik, 2017, gen. n." docType="treatment" docVersion="8" lastPageNumber="341" masterDocId="8A58385BB3290C2655222368FFD35400" masterDocTitle="A survey of East Palaearctic Lycosidae (Araneae). 13. A new genus of spiny-legs Pardosinae from Eastern Kazakhstan" masterLastPageNumber="350" masterPageNumber="339" pageNumber="340" updateTime="1698468534854" updateUser="ExternalLinkService">
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<mods:title id="F8A6900D13D9C33E0CF81B6BEA03E106">A survey of East Palaearctic Lycosidae (Araneae). 13. A new genus of spiny-legs Pardosinae from Eastern Kazakhstan</mods:title>
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<mods:namePart id="1A3AB7523B1D53F03FA8DAE301DE6BB2">Marusik, Yuri M.</mods:namePart>
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<mods:date id="3D60E9366B99D263764180297AAF7581">2017</mods:date>
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<mods:number id="184A7EB6BF969454A44A8E76D88B2BF4">2017-09-15</mods:number>
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<treatment id="76614023B3280C2455B52292FDC2575D" ID-DOI="http://doi.org/10.5281/zenodo.6046126" ID-GBIF-Taxon="134090021" ID-Zenodo-Dep="6046126" LSID="urn:lsid:plazi:treatment:76614023B3280C2455B52292FDC2575D" httpUri="http://treatment.plazi.org/id/76614023B3280C2455B52292FDC2575D" lastPageId="2" lastPageNumber="341" pageId="1" pageNumber="340">
<subSubSection id="B6D2A2BEB3280C2755B52292FE7C5614" box="[151,431,506,532]" pageId="1" pageNumber="340" type="nomenclature">
<paragraph id="FE77F135B3280C2755B52292FE7C5614" blockId="1.[151,431,506,532]" box="[151,431,506,532]" pageId="1" pageNumber="340">
<heading id="A53F4659B3280C2755B52292FE7C5614" bold="true" box="[151,431,506,532]" fontSize="11" level="1" pageId="1" pageNumber="340" reason="1">
<emphasis id="CCBC2D27B3280C2755B52292FE7C5614" bold="true" box="[151,431,506,532]" pageId="1" pageNumber="340">
<taxonomicName id="39C88AB6B3280C2755B52292FE855614" ID-CoL="4992" box="[151,342,506,532]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCBC2D27B3280C2755B52292FE855614" bold="true" box="[151,342,506,532]" italics="true" pageId="1" pageNumber="340">Dzhungarocosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D78F905CB3280C27547E2292FE7C5614" box="[348,431,506,532]" pageId="1" pageNumber="340" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6D2A2BEB3280C2755B52129FD5C565A" box="[151,655,577,602]" pageId="1" pageNumber="340" type="materials_examined">
<paragraph id="FE77F135B3280C2755B52129FD5C565A" blockId="1.[151,1437,577,2006]" box="[151,655,577,602]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755B52129FEF8565A" bold="true" box="[151,299,577,602]" pageId="1" pageNumber="340">Type species</emphasis>
.
<taxonomicName id="39C88AB6B3280C27541B2129FD9A565A" box="[313,585,577,602]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="omelkoi" status="sp. nov.">
<emphasis id="CCBC2D27B3280C27541B2129FD9A565A" box="[313,585,577,602]" italics="true" pageId="1" pageNumber="340">Dzhungarocosa omelkoi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2757722129FD5C565A" bold="true" box="[592,655,577,602]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2757722129FD5C565A" box="[592,655,577,602]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B6D2A2BEB3280C2755E5210DFBE6570E" pageId="1" pageNumber="340" type="etymology">
<paragraph id="FE77F135B3280C2755E5210DFC2156A2" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755E5210DFE95567E" bold="true" box="[199,326,613,638]" pageId="1" pageNumber="340">Etymology</emphasis>
. The generic name is derived from the
<typeStatus id="21734F97B3280C27562B210EFCEA567E" box="[777,825,614,638]" pageId="1" pageNumber="340">type</typeStatus>
locality of the
<typeStatus id="21734F97B3280C2756CA210EFBCB567E" box="[1000,1048,614,638]" pageId="1" pageNumber="340">type</typeStatus>
species, Dzhungarian Alatau, and end with -cosa, typical ending for
<taxonomicName id="39C88AB6B3280C27573421E1FD5B56A2" authorityName="Sundevall" authorityYear="1833" box="[534,648,649,674]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
genera. The gender is feminine.
</paragraph>
<paragraph id="FE77F135B3280C2755E521C5FBE6570E" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755E521C5FF2C56C6" bold="true" box="[199,255,685,710]" pageId="1" pageNumber="340">Note</emphasis>
. The new genus belongs to Pardosinae (sensu Zyuzin, 1993), judging by the ventral origin of the embolus, the tip of the embolus resting in a small tegular depression, thick and sclerotized tegular apophysis and well sclerotized basal part of palea lacking synembolus and concealed by tegulum.
</paragraph>
</subSubSection>
<subSubSection id="B6D2A2BEB3280C2755E52071FCD351B9" pageId="1" pageNumber="340" type="diagnosis">
<paragraph id="FE77F135B3280C2755E52071FACA579E" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755E52071FE925732" bold="true" box="[199,321,793,818]" pageId="1" pageNumber="340">Diagnosis.</emphasis>
<taxonomicName id="39C88AB6B3280C27546B2071FE2A5732" box="[329,505,793,818]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCBC2D27B3280C27546B2071FE2A5732" box="[329,505,793,818]" italics="true" pageId="1" pageNumber="340">Dzhungarocosa</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2757232071FD835732" bold="true" box="[513,592,793,818]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2757232071FD835732" box="[513,592,793,818]" pageId="1" pageNumber="340" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
differs from other Pardosinae genera with numerous (4–6) pairs of ventral tibial spines on leg I and occurring in North Holarctic by having unmodified embolus (modified in all others), not reduced palea with 2 or 3 processes (1 process, or palea reduced), epigyne lacking distinct anterior pockets (present in all other genera) and with large septal pockets raised over septal plain (pockets absent or not raised).
</paragraph>
<paragraph id="FE77F135B3280C2755E520C1FF09512A" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755E520C1FEBA57C2" bold="true" box="[199,361,937,962]" pageId="1" pageNumber="340">Relationships</emphasis>
. Although the 3 species belonging to this new genus have 4–6 pairs of ventral tibial spines on leg I, arranged in slanting row (
<figureCitation id="66F3EDB0B3280C27572520A5FD2D57E6" box="[519,766,973,998]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1852,1875]" captionTargetBox="[161,1427,304,1826]" captionTargetId="figure@3.[151,1436,303,1831]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Habitus and retrolateral view of leg I in Dzhungarocosa zhishengi sp. n. (1 – 4), D. omelkoi sp. n. (5 – 8) and D. ballarini sp. n. (9 – 12). 1, 5, 9 male, dorsal; 2, 6, 10 female, dorsal; 3, 7, 11 male leg I; 4, 8, 12 female leg I. Scale = 2 mm for 1 – 2, 5 – 6, 9 – 10; 1 mm for 3 – 4, 7 – 8, 11 – 12." httpUri="https://zenodo.org/record/891910/files/figure.png" pageId="1" pageNumber="340">Figs 3–4, 7–8, 11–12</figureCitation>
) like in
<taxonomicName id="39C88AB6B3280C27564120A5FBDB57E6" authorityName="Dahl" authorityYear="1908" box="[867,1032,973,998]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C27564120A5FBDB57E6" box="[867,1032,973,998]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
,
<taxonomicName id="39C88AB6B3280C27513A20A5FB5057E6" box="[1048,1155,973,998]" class="Arachnida" family="Lycosidae" genus="Gulocosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C27513A20A5FB5057E6" box="[1048,1155,973,998]" italics="true" pageId="1" pageNumber="340">Gulocosa</emphasis>
</taxonomicName>
,
<taxonomicName id="39C88AB6B3280C2751B620A5FAF557E6" box="[1172,1318,973,998]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2751B620A5FAF557E6" box="[1172,1318,973,998]" italics="true" pageId="1" pageNumber="340">Mongolicosa</emphasis>
</taxonomicName>
and other genera belonging to the
<taxonomicName id="39C88AB6B3280C2754862099FD9A500A" authorityName="Dahl" authorityYear="1908" box="[420,585,1009,1034]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2754862099FD9A500A" box="[420,585,1009,1034]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
-complex (see
<bibRefCitation id="9A598CC4B3280C2757CE2099FC1A500A" author="Marusik" box="[748,969,1009,1034]" pageId="1" pageNumber="340" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="CCBC2D27B3280C275672209AFC52500A" box="[848,897,1009,1034]" italics="true" pageId="1" pageNumber="340">et al</emphasis>
. 2004
</bibRefCitation>
,
<bibRefCitation id="9A598CC4B3280C2756F62099FBC3500A" author="Marusik" box="[980,1040,1009,1034]" pageId="1" pageNumber="340" refString="Marusik, Y. M., Omelko, M. M. &amp; Koponen, S. (2015) A survey of East Palaearctic Lycosidae (Araneae). 11. Two new genera from the Acantholycosa complex. Zootaxa, 3985 (2), 252 - 264. https: // doi. org / 10.11646 / zootaxa. 3985.2.4" type="journal article" year="2015">2015</bibRefCitation>
), it seems that
<taxonomicName id="39C88AB6B3280C2751962099FAB7500A" box="[1204,1380,1009,1034]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCBC2D27B3280C2751962099FAB7500A" box="[1204,1380,1009,1034]" italics="true" pageId="1" pageNumber="340">Dzhungarocosa</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2750492099FF7D502E" bold="true" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2750492099FF7D502E" pageId="1" pageNumber="340" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
is not related to any of these genera. All species in the genera belonging to the
<taxonomicName id="39C88AB6B3280C27516D277DFB27502E" authorityName="Dahl" authorityYear="1908" box="[1103,1268,1045,1070]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C27516D277DFB27502E" box="[1103,1268,1045,1070]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
-complex have modified embolus; either thick (
<taxonomicName id="39C88AB6B3280C2754DD2751FDA55052" box="[511,630,1081,1106]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2754DD2751FDA55052" box="[511,630,1081,1106]" italics="true" pageId="1" pageNumber="340">Sibirocosa</emphasis>
</taxonomicName>
,
<taxonomicName id="39C88AB6B3280C2757A12751FD3D5052" box="[643,750,1081,1106]" class="Arachnida" family="Lycosidae" genus="Gulocosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2757A12751FD3D5052" box="[643,750,1081,1106]" italics="true" pageId="1" pageNumber="340">Gulocosa</emphasis>
</taxonomicName>
), or very (
<taxonomicName id="39C88AB6B3280C2756442751FC2B5052" box="[870,1016,1081,1106]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2756442751FC2B5052" box="[870,1016,1081,1106]" italics="true" pageId="1" pageNumber="340">Mongolicosa</emphasis>
</taxonomicName>
) or relatively broad (
<taxonomicName id="39C88AB6B3280C2751C52751FA5F5052" authorityName="Dahl" authorityYear="1908" box="[1255,1420,1081,1106]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2751C52751FA5F5052" box="[1255,1420,1081,1106]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
), often having a basal spine, while in species belonging to the new genus, embolus is not modified and is relatively short. The palea in
<taxonomicName id="39C88AB6B3280C27545927E9FDF8509A" box="[379,555,1153,1178]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCBC2D27B3280C27545927E9FDF8509A" box="[379,555,1153,1178]" italics="true" pageId="1" pageNumber="340">Dzhungarocosa</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C27571727E9FD5B509A" bold="true" box="[565,648,1153,1178]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C27571727E9FD5B509A" box="[565,648,1153,1178]" pageId="1" pageNumber="340" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
has 2 broad processes (or ridges), while in the genera belonging to
<taxonomicName id="39C88AB6B3280C2755B527CDFEEF50BE" authorityName="Dahl" authorityYear="1908" box="[151,316,1189,1214]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2755B527CDFEEF50BE" box="[151,316,1189,1214]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
-complex the palea is either reduced (
<taxonomicName id="39C88AB6B3280C2757C527CDFC8E50BE" box="[743,861,1189,1214]" class="Arachnida" family="Lycosidae" genus="Sibirocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2757C527CDFC8E50BE" box="[743,861,1189,1214]" italics="true" pageId="1" pageNumber="340">Sibirocosa</emphasis>
</taxonomicName>
,
<taxonomicName id="39C88AB6B3280C27564F27CDFC0B50BE" box="[877,984,1189,1214]" class="Arachnida" family="Lycosidae" genus="Melecosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C27564F27CDFC0B50BE" box="[877,984,1189,1214]" italics="true" pageId="1" pageNumber="340">Melecosa</emphasis>
</taxonomicName>
), unmodified (
<taxonomicName id="39C88AB6B3280C2751A127CDFAC650BE" box="[1155,1301,1189,1214]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2751A127CDFAC650BE" box="[1155,1301,1189,1214]" italics="true" pageId="1" pageNumber="340">Mongolicosa</emphasis>
</taxonomicName>
) or has one process (
<taxonomicName id="39C88AB6B3280C2755D927A1FE7350E2" authorityName="Dahl" authorityYear="1908" box="[251,416,1225,1250]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2755D927A1FE7350E2" box="[251,416,1225,1250]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
,
<taxonomicName id="39C88AB6B3280C27548F27A2FDFD50E1" box="[429,558,1226,1249]" class="Arachnida" family="Lycosidae" genus="Pyrenecosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C27548F27A2FDFD50E1" box="[429,558,1226,1249]" italics="true" pageId="1" pageNumber="340">Pyrenecosa</emphasis>
</taxonomicName>
). The new genus has bifurcated terminal apophysis with rounded tips, while in
<taxonomicName id="39C88AB6B3280C2755B52785FEEF5106" authorityName="Dahl" authorityYear="1908" box="[151,316,1261,1286]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2755B52785FEEF5106" box="[151,316,1261,1286]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
and related genera the terminal apophysis is unbranched and either sharply pointed or has an apical spine.
</paragraph>
<paragraph id="FE77F135B3280C2755E5265DFA4E5172" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
Females of the new genus lack distinct anterior pockets (present in all genera belonging to the
<taxonomicName id="39C88AB6B3280C2751CC265DFA40514E" authorityName="Dahl" authorityYear="1908" box="[1262,1427,1333,1358]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="CCBC2D27B3280C2751CC265DFA40514E" box="[1262,1427,1333,1358]" italics="true" pageId="1" pageNumber="340">Acantholycosa</emphasis>
</taxonomicName>
- complex) and have very large septal pockets extending septal plain. This character is unknown in other Pardosinae.
</paragraph>
<paragraph id="FE77F135B3280C2755E52615FCD351B9" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
Both palpal and epigynal morphology do not allow to trace relationships of the new genus with other Pardosinae or any species group in
<emphasis id="CCBC2D27B3280C27570126C9FD2A51BA" box="[547,761,1441,1466]" italics="true" pageId="1" pageNumber="340">
<taxonomicName id="39C88AB6B3280C27570126C9FD5251BA" box="[547,641,1441,1466]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus" sensu="lato">Pardosa</taxonomicName>
<taxonomicNameLabel id="D78F905CB3280C2757AA26CAFD2A51BA" box="[648,761,1441,1466]" pageId="1" pageNumber="340" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="B6D2A2BEB3280C2455E526ADFDC2575D" lastPageId="2" lastPageNumber="341" pageId="1" pageNumber="340" type="description">
<paragraph id="FE77F135B3280C2755E526ADFDB552B6" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
<emphasis id="CCBC2D27B3280C2755E526ADFE8A51DE" bold="true" box="[199,345,1477,1502]" pageId="1" pageNumber="340">Description.</emphasis>
Total length: male 7.5–9.5, female 8.0–10.4. Carapace length: male 3.7–4.8, female 4.0–4.7; width: male 3.0–3.9; female 3.2–3.9. Carapace black or dark-brown or black with yellow stripes around the fovea and near edges (female of
<taxonomicName id="39C88AB6B3280C2754E72566FD815226" box="[453,594,1549,1574]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B3280C2754E72566FD815226" box="[453,594,1549,1574]" italics="true" pageId="1" pageNumber="340">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2757782565FD485226" bold="true" box="[602,667,1549,1574]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2757782565FD485226" box="[602,667,1549,1574]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Clypeus yellow with black stripe on its lower edge or with a gray triangular spot in the middle, or uniformly yellow (female of
<taxonomicName id="39C88AB6B3280C27566A255AFC17524A" box="[840,964,1585,1610]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="omelkoi" status="sp. nov.">
<emphasis id="CCBC2D27B3280C27566A255AFC17524A" box="[840,964,1585,1610]" italics="true" pageId="1" pageNumber="340">D. omelkoi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2756E92559FBD9524A" bold="true" box="[971,1034,1585,1610]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2756E92559FBD9524A" box="[971,1034,1585,1610]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Chelicerae black or dark-gray (
<taxonomicName id="39C88AB6B3280C2750A3255AFF2A526E" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="ballarini" status="sp. nov.">
<emphasis id="CCBC2D27B3280C2750A3255AFF2A526E" italics="true" pageId="1" pageNumber="340">D. ballarini</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C275423253DFE91526E" bold="true" box="[257,322,1621,1646]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C275423253DFE91526E" box="[257,322,1621,1646]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) with yellow stripes in males and uniformly yellow-brown (
<taxonomicName id="39C88AB6B3280C2756D4253EFBAF526E" box="[1014,1148,1621,1646]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="ballarini" status="sp. nov.">
<emphasis id="CCBC2D27B3280C2756D4253EFBAF526E" box="[1014,1148,1621,1646]" italics="true" pageId="1" pageNumber="340">D. ballarini</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2751A7253DFB15526E" bold="true" box="[1157,1222,1621,1646]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2751A7253DFB15526E" box="[1157,1222,1621,1646]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or yellow basally with brown top in other species. Sternum and labium black or dark-gray in all species. Endites brown- or yellowgray with a yellow spot on the inner side.
</paragraph>
<paragraph id="FE77F135B3280C2755E525A9FC77536A" blockId="1.[151,1437,577,2006]" pageId="1" pageNumber="340">
Palps with the same colouration as the legs in females. Legs coloured by combination of black or gray and yellow rings in females. In males legs bright-yellow with black rings in
<taxonomicName id="39C88AB6B3280C2756EF258EFB8852FE" box="[973,1115,1765,1790]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B3280C2756EF258EFB8852FE" box="[973,1115,1765,1790]" italics="true" pageId="1" pageNumber="340">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C275141258DFB7752FE" bold="true" box="[1123,1188,1765,1790]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C275141258DFB7752FE" box="[1123,1188,1765,1790]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
or different segments and legs have different colouration (other species). Abdomen black or dark-gray with unclear pattern or without it. Leg I spination pattern: Fe: d1-1-1 p0-1(0)-2(3) r0(2)-1(0)-1; Pt: d1-1 p0(1)-0 r1(0)-0; Ti: d1-0-1 p1(2)-0-0 r1-0-0, pv6(4)-rv6(5)-
<date id="8A76D7F5B3280C2754182439FE84536A" box="[314,343,1873,1898]" pageId="1" pageNumber="340">v2</date>
ap; Mt: p1(0)-1-0-2ap, r1(0)-1-0-2ap,
<date id="8A76D7F5B3280C275622243AFCEA536A" box="[768,825,1874,1898]" pageId="1" pageNumber="340">v2-2</date>
(1)-0-1ap.
</paragraph>
<paragraph id="FE77F135B3280C2455E5241DFB5C5588" blockId="1.[151,1437,577,2006]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="341" pageId="1" pageNumber="340">
Male palp. Colouration: general colour black, patella yellow, coxae–femur black except yellow patella or black with yellow spots in femur and patella (
<taxonomicName id="39C88AB6B3280C27574724F2FD2753B2" box="[613,756,1945,1970]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B3280C27574724F2FD2753B2" box="[613,756,1945,1970]" italics="true" pageId="1" pageNumber="340">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B3280C2757DE24F1FCED53B2" bold="true" box="[764,830,1945,1970]" pageId="1" pageNumber="340">
<taxonomicNameLabel id="D78F905CB3280C2757DE24F1FCED53B2" box="[764,830,1945,1970]" pageId="1" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Tibia relatively long, about 0.56–0.62 of cymbium length. Cymbium black, modified setae absent; tip relatively long, about 1/3 of cymbial length, with one small claw. Tegulum unmodified, not extending ventrally. Tegular apophysis with two arms, both sharply pointed; anterior arm (
<emphasis id="CCBC2D27B32B0C24541523D4FE8554D3" box="[311,342,188,211]" italics="true" pageId="2" pageNumber="341">Aa</emphasis>
) larger than posterior (
<emphasis id="CCBC2D27B32B0C24574723D5FD5754D4" box="[613,644,189,212]" italics="true" pageId="2" pageNumber="341">Pa</emphasis>
). Anterior arm triangular or claw-like. Tegular conductor (
<emphasis id="CCBC2D27B32B0C24501323D5FA9B54D4" box="[1329,1352,189,212]" italics="true" pageId="2" pageNumber="341">Tc</emphasis>
) large. Palea with 2 processes, anterior (
<emphasis id="CCBC2D27B32B0C2457322388FDFC54F7" box="[528,559,224,247]" italics="true" pageId="2" pageNumber="341">Ap</emphasis>
), and posterior (
<emphasis id="CCBC2D27B32B0C2457CC2389FCDE54F8" box="[750,781,225,248]" italics="true" pageId="2" pageNumber="341">Pp</emphasis>
). Posterior process-ridge wide, subdivided into 2 parts or entire. Terminal (
<emphasis id="CCBC2D27B32B0C24547F226DFEA6551C" box="[349,373,261,284]" italics="true" pageId="2" pageNumber="341">Ta</emphasis>
) apophysis large, bifurcated near terminal part, tips rounded. Embolic division with rounded pit (
<emphasis id="CCBC2D27B32B0C2455EA2241FF345540" box="[200,231,297,320]" italics="true" pageId="2" pageNumber="341">Rp</emphasis>
) between palea and base of embolus. Embolus relatively short, especially in
<taxonomicName id="39C88AB6B32B0C24514D2241FB245540" box="[1135,1271,295,320]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="ballarini" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C24514D2241FB245540" box="[1135,1271,295,320]" italics="true" pageId="2" pageNumber="341">D. ballarini</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C245023224FFA975540" bold="true" box="[1281,1348,295,320]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C245023224FFA975540" box="[1281,1348,295,320]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="39C88AB6B32B0C2450A32241FF235565" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="omelkoi" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C2450A32241FF235565" italics="true" pageId="2" pageNumber="341">D. omelkoi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C2455D52224FEEB5565" bold="true" box="[247,312,332,357]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C2455D52224FEEB5565" box="[247,312,332,357]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; not modified, lacking tooth, not twisted around axes. Embolus either straight (
<taxonomicName id="39C88AB6B32B0C24519F2225FA995565" box="[1213,1354,332,357]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C24519F2225FA995565" box="[1213,1354,332,357]" italics="true" pageId="2" pageNumber="341">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C2450712224FA405565" bold="true" box="[1363,1427,332,357]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C2450712224FA405565" box="[1363,1427,332,357]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or slightly waved near the base. Tip of embolus slightly widened, abrupt and bent dorsally.
</paragraph>
<paragraph id="FE77F135B32B0C2455E522FCFD21563D" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="341">
Epigyne. Fovea and anterior pockets absent. Septum well developed, with either thin or wide and bifurcated stem (
<taxonomicName id="39C88AB6B32B0C2455FF22D1FEB955D0" box="[221,362,439,464]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C2455FF22D1FEB955D0" box="[221,362,439,464]" italics="true" pageId="2" pageNumber="341">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C24545022DFFE6155D0" bold="true" box="[370,434,439,464]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C24545022DFFE6155D0" box="[370,434,439,464]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Stem (
<emphasis id="CCBC2D27B32B0C24573622D0FDFB55CF" box="[532,552,440,463]" italics="true" pageId="2" pageNumber="341">St</emphasis>
) anteriorly turn into a sclerotized arches (
<emphasis id="CCBC2D27B32B0C24512422D1FBF655D0" box="[1030,1061,441,464]" italics="true" pageId="2" pageNumber="341">Ea</emphasis>
). Base of septum (
<emphasis id="CCBC2D27B32B0C2451DF22D1FACA55D0" box="[1277,1305,441,464]" italics="true" pageId="2" pageNumber="341">Bs</emphasis>
) with large either anterior or posterior pockets (
<emphasis id="CCBC2D27B32B0C24571B22B5FD8B55F4" box="[569,600,477,500]" italics="true" pageId="2" pageNumber="341">Po</emphasis>
), ventral (external) wall of pocket strongly extends plain of septum and makes pockets well visible in anterior, posterior or lateral view (
<figureCitation id="66F3EDB0B32B0C24565B2297FA855618" box="[889,1366,511,536]" captionStart-0="FIGURES 52 – 66" captionStart-1="FIGURES 67 – 74" captionStartId-0="8.[151,264,1598,1621]" captionStartId-1="9.[151,264,1495,1518]" captionTargetBox-0="[159,1427,185,1574]" captionTargetBox-1="[152,1434,628,1475]" captionTargetId-0="figure@8.[159,1427,185,1578]" captionTargetId-1="figure@9.[152,1434,628,1475]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 52 – 66. Epigyne of Dzhungarocosa zhishengi sp. n. (52 – 56), D. omelkoi sp. n. (57 – 61) and D. ballarini sp. n. (62 – 66). 52, 57, 62 ventral; 53, 58, 63 macerated, ventral; 54, 59, 64 macerated, dorsal; 55, 60, 65 anterior; 56, 61, 66 posterior. Scale = 0.2 mm. Abbreviations: Li lip of the epigyne, Rh head of the receptacle, Po septal pocket." captionText-1="FIGURES 67 – 74. Epigyne of Dzhungarocosa zhishengi sp. n. (67 – 68), D. omelkoi sp. n. (69 – 71) and D. ballarini sp. n. (72 – 74). 67, 69, 72 ventral; 68, 70, 73 ventro-lateral, showing septal pocket; 71, 74 posterior. Scale = 0.1 mm (if indicated). Abbreviations: Bs base of septum, Ea sclerotized arches, Po posterior pockets, St septal stem." httpUri-0="https://zenodo.org/record/891922/files/figure.png" httpUri-1="https://zenodo.org/record/891924/files/figure.png" pageId="2" pageNumber="341">Figs 56–57, 61–62, 66–68, 70–71, 73–74</figureCitation>
). Lips (
<emphasis id="CCBC2D27B32B0C245582214DFF64563C" box="[160,183,549,572]" italics="true" pageId="2" pageNumber="341">Li</emphasis>
) not touching each other, separated by septal base.
</paragraph>
<paragraph id="FE77F135B32B0C2455E5212FFA875660" blockId="2.[151,1437,151,861]" box="[199,1364,583,608]" pageId="2" pageNumber="341">
Receptacles tubular, with club-like heads (
<emphasis id="CCBC2D27B32B0C245781212FFD115660" box="[675,706,583,608]" italics="true" pageId="2" pageNumber="341">Rh</emphasis>
), base of receptacles diverging, heads slightly converging.
</paragraph>
<paragraph id="FE77F135B32B0C2455E52104FA575684" blockId="2.[151,1437,151,861]" box="[199,1412,620,645]" pageId="2" pageNumber="341">
<emphasis id="CCBC2D27B32B0C2455E52104FEFD5685" bold="true" box="[199,302,620,645]" pageId="2" pageNumber="341">Habitats</emphasis>
. All three species were collected among stone screes (
<figureCitation id="66F3EDB0B32B0C2456AD2104FBDC5685" box="[911,1039,620,645]" captionStart="FIGURES 75 – 80" captionStartId="10.[151,264,1944,1967]" captionTargetBox="[151,1436,481,1923]" captionTargetId="figure@10.[151,1436,481,1923]" captionTargetPageId="10" captionText="FIGURES 75 – 80. Habitats and type localities of Dzhungarocosa zhishengi sp. n. (75, 79, 80 — square), D. ballarini sp. n. (76, 79, 80 — circle) and D. omelkoi sp. n. (77 – 79, 80 — star)." httpUri="https://zenodo.org/record/891926/files/figure.png" pageId="2" pageNumber="341">Figs 75–78</figureCitation>
) on elevations over 2900 meters.
</paragraph>
<paragraph id="FE77F135B32B0C2455E521E7FDC2575D" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="341">
<emphasis id="CCBC2D27B32B0C2455E521E7FC7356A8" bold="true" box="[199,928,655,680]" pageId="2" pageNumber="341">
Composition and relationships within
<taxonomicName id="39C88AB6B32B0C2457AC21E7FC9756A8" box="[654,836,655,680]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCBC2D27B32B0C2457AC21E7FC9756A8" bold="true" box="[654,836,655,680]" italics="true" pageId="2" pageNumber="341">Dzhungarocosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D78F905CB32B0C24566921E7FC4056A8" box="[843,915,655,680]" pageId="2" pageNumber="341" rank="genus">gen. n</taxonomicNameLabel>
..
</emphasis>
Among three species currently considered in the genus
<taxonomicName id="39C88AB6B32B0C24542821DDFE5C56CD" box="[266,399,692,717]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="ballarini" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C24542821DDFE5C56CD" box="[266,399,692,717]" italics="true" pageId="2" pageNumber="341">D. ballarini</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C2454B521DCFE0B56CD" bold="true" box="[407,472,692,717]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C2454B521DCFE0B56CD" box="[407,472,692,717]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="39C88AB6B32B0C2454C421DDFDB156CD" box="[486,610,692,717]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="omelkoi" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C2454C421DDFDB156CD" box="[486,610,692,717]" italics="true" pageId="2" pageNumber="341">D. omelkoi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C24574821DCFD7956CD" bold="true" box="[618,682,692,717]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C24574821DCFD7956CD" box="[618,682,692,717]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="39C88AB6B32B0C2457C021DDFCBC56CD" box="[738,879,692,717]" class="Arachnida" family="Lycosidae" genus="Dzhungarocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="CCBC2D27B32B0C2457C021DDFCBC56CD" box="[738,879,692,717]" italics="true" pageId="2" pageNumber="341">D. zhishengi</emphasis>
</taxonomicName>
<emphasis id="CCBC2D27B32B0C24565521DCFC6456CD" bold="true" box="[887,951,692,717]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="D78F905CB32B0C24565521DCFC6456CD" box="[887,951,692,717]" pageId="2" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the two former species are closely related and have similar colour pattern, number of ventral tibial spines on leg I, tegular apophysis with claw-like anterior arm, bent embolus and similarly shaped epigyne septum. The latter species stays apart, but shares with the remaining species the shape of the terminal apophysis, unmodified embolus, raised septal pockets and sclerotized arches in anterior part of epigyne.
</paragraph>
</subSubSection>
</treatment>
</document>
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