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<document id="9FC1CBCEED702B7122F4F6EDD73F794F" ID-CLB-Dataset="153839" ID-DOI="10.11646/zootaxa.5190.2.1" ID-GBIF-Dataset="60faa853-726d-4b71-a3cb-c21a0087515f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7120060" ID-ZooBank="92BFDA7A-28C0-4816-8A91-CFB842C95E96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664372331544" checkinUser="plazi" docAuthor="Zarei, Fatah, Esmaeili, Hamid Reza, Abbasi, Keyvan, Kovačić, Marcelo, Schliewen, Ulrich K. &amp; Stepien, Carol A." docDate="2022" docId="4154F53A8A0DFF88FF72AAECFE854DC7" docLanguage="en" docName="zootaxa.5190.2.1.pdf" docOrigin="Zootaxa 5190 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benthophilus persicus Kovacic, Esmaeili, Zarei, Abbasi &amp; Schliewen 2021" docType="treatment" docVersion="7" lastPageNumber="169" masterDocId="BD6D8D428A1CFF9AFFE5AC4EFF994C6A" masterDocTitle="Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification" masterLastPageNumber="193" masterPageNumber="151" pageNumber="168" updateTime="1699361546833" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4623328C1F94F39A23CC1BD0D302A825">Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification</mods:title>
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<mods:namePart id="F621CA2EB1417DF0036C699E9677D5FC">Zarei, Fatah</mods:namePart>
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<mods:affiliation id="630BBF7CCDDAFF0D9DA642268859B536">Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. &amp; fataahzarei @ gmail. com; https: // orcid. org / 0000 - 0001 - 5552 - 4301</mods:affiliation>
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<mods:namePart id="DBF9AAD21ABE780EE840EAB21CD5629E">Esmaeili, Hamid Reza</mods:namePart>
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<mods:affiliation id="B6D785F9550915C62ABB865A38BD4BAB">Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. &amp; hresmaeili @ shirazu. ac. ir; https: // orcid. org / 0000 - 0002 - 9736 - 397 X</mods:affiliation>
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<mods:namePart id="B047A9D36861E8EBC41E264B4968160F">Abbasi, Keyvan</mods:namePart>
<mods:affiliation id="32C556479C8C60DE5358D9BDB04A2B60">Inland Waters Aquaculture Research Center, Iranian Fisheries Sciences Research Institute, Agricultural Research, Education and Extension Organization, Bandar Anzali, Iran.</mods:affiliation>
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<mods:namePart id="7D2C67CFDF0FB0C603D3CDDD46E620BE">Kovačić, Marcelo</mods:namePart>
<mods:affiliation id="7A3D4C998BBE985856FD8A4661DCD096">Natural History Museum Rijeka, Lorenzov prolaz 1, HR- 51000 Rijeka, Croatia.</mods:affiliation>
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<mods:namePart id="C8019FFEE780C3EF01BDE85DAD0FD750">Schliewen, Ulrich K.</mods:namePart>
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<mods:affiliation id="BA3F06A973764D68E1E721C357A3B908">SNSB-Bavarian State Collection of Zoology, Department of Ichthyology, Münchhausenstr. 21, 81247 München, Germany. schliewen @ snsb. de; https: // orcid. org / 0000 - 0003 - 0187 - 4319</mods:affiliation>
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<mods:affiliation id="F43425683891B2D508447EE35042C5C5">National Museum of Natural History, Department of Vertebrate Zoology, Smithsonian Institution, Washington D. C, USA. StepienCA @ si. edu; https: // orcid. org / 0000 - 0002 - 5544 - 4333</mods:affiliation>
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<emphasis id="FB89983E8A0DFF8BFF72AAECFBC74AD6" bold="true" box="[151,1118,1698,1725]" pageId="17" pageNumber="168">
16.
<taxonomicName id="0EFD3FAF8A0DFF8BFF23AAECFBC74AD6" authority="Kovacic, Esmaeili, Zarei, Abbasi &amp; Schliewen, 2021" authorityName="Kovacic, Esmaeili, Zarei, Abbasi &amp; Schliewen" authorityYear="2021" box="[198,1118,1698,1725]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="persicus">
<emphasis id="FB89983E8A0DFF8BFF23AAECFE544AD7" bold="true" box="[198,461,1698,1725]" italics="true" pageId="17" pageNumber="168">Benthophilus persicus</emphasis>
<bibRefCitation id="AD6C39DD8A0DFF8BFE3CAAECFBC74AD6" author="Kovacic, M. &amp; Esmaeili, H. R. &amp; Zarei, F. &amp; Abbasi, K. &amp; Schliewen, U. K." box="[473,1118,1698,1725]" pageId="17" pageNumber="168" pagination="45 - 63" refId="ref27887" refString="Kovacic, M., Esmaeili, H. R., Zarei, F., Abbasi, K. &amp; Schliewen, U. K. (2021) A new species of Tadpole goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea. Zootaxa, 4980 (1), 45 - 63. https: // doi. org / 10.11646 / zootaxa. 4980.1.3" type="journal article" year="2021">Kovačić, Esmaeili, Zarei, Abbasi &amp; Schliewen, 2021</bibRefCitation>
</taxonomicName>
</emphasis>
(
<figureCitation id="51C658A98A0DFF8BFB96AAEDFB534AD7" box="[1139,1226,1699,1725]" captionStart="FIGURE 13" captionStartId="18.[151,250,927,951]" captionTargetBox="[174,1409,472,904]" captionTargetId="figure-124@18.[174,1413,453,904]" captionTargetPageId="18" captionText="FIGURE 13. Benthophilus persicus. ZM-CBSU S026-1, 35.3 mm SL, Shafaroud mouth, Gilan Province, Iran." figureDoi="http://doi.org/10.5281/zenodo.7120102" httpUri="https://zenodo.org/record/7120102/files/figure.png" pageId="17" pageNumber="168">Fig. 13</figureCitation>
), Persian Tadpole Goby
</paragraph>
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<treatmentCitation id="485C623D8A0DFF8BFF72AAA5FB894B6F" ID-DOI="http://doi.org/10.5281/zenodo.4945539" ID-Zenodo-Dep="4945539" author="Kovacic, M. &amp; Esmaeili, H. R. &amp; Zarei, F. &amp; Abbasi, K. &amp; Schliewen, U. K." box="[151,1040,1771,1797]" httpUri="http://treatment.plazi.org/id/038D87D3A805FF9DC1ECF994718BFE2A" page="47" pageId="17" pageNumber="168" year="2021">
<taxonomicName id="0EFD3FAF8A0DFF8BFF72AAA5FB894B6F" authority="Kovacic, Esmaeili, Zarei, Abbasi &amp; Schliewen, 2021: 47" authorityName="Kovacic, Esmaeili, Zarei, Abbasi &amp; Schliewen" authorityPageNumber="47" authorityYear="2021" box="[151,1040,1771,1797]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="persicus">
<emphasis id="FB89983E8A0DFF8BFF72AAA5FE164B6E" box="[151,399,1771,1797]" italics="true" pageId="17" pageNumber="168">Benthophilus persicus</emphasis>
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<bibRefCitation id="AD6C39DD8A0DFF8BFE73AAA5FB894B6F" author="Kovacic, M. &amp; Esmaeili, H. R. &amp; Zarei, F. &amp; Abbasi, K. &amp; Schliewen, U. K." box="[406,1040,1771,1797]" pageId="17" pageNumber="168" pagination="45 - 63" refId="ref27887" refString="Kovacic, M., Esmaeili, H. R., Zarei, F., Abbasi, K. &amp; Schliewen, U. K. (2021) A new species of Tadpole goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea. Zootaxa, 4980 (1), 45 - 63. https: // doi. org / 10.11646 / zootaxa. 4980.1.3" type="journal article" year="2021">Kovačić, Esmaeili, Zarei, Abbasi &amp; Schliewen, 2021: 47</bibRefCitation>
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, figs. 2–7; type locality: off
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, 37°29’N, 49°29’E;
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: ZSM 47595,
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male,
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TL,
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;
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,
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<collectionCode id="AFECDCE98A0DFF8BFBD8AB19FB204B1B" box="[1085,1209,1879,1905]" pageId="17" pageNumber="168">ZM-CBSU</collectionCode>
<specimenCode id="995BEC578A0DFF8BFB26AB19FABB4B1B" box="[1219,1314,1879,1905]" collectionCode="ZM-CBSU" pageId="17" pageNumber="168">S003-17</specimenCode>
(21)
</materialsCitation>
,
<materialsCitation id="79954E718A0DFF8BFA8CAB19FC9E4BFF" ID-GBIF-Occurrence="3914046453" collectionCode="ZM-CBSU" country="Iran" pageId="17" pageNumber="168" specimenCode="S003-115" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
ZMCBSU S003-112–113 (2),
<collectionCode id="AFECDCE98A0DFF8BFE0AAB35FDF24BFF" box="[495,619,1915,1941]" pageId="17" pageNumber="168">ZM-CBSU</collectionCode>
<specimenCode id="995BEC578A0DFF8BFD97AB34FD474BFF" box="[626,734,1914,1941]" collectionCode="ZM-CBSU" pageId="17" pageNumber="168">S003-115</specimenCode>
(1)
</materialsCitation>
,
<materialsCitation id="79954E718A0DFF8BFCF6AB35FBA24BFF" ID-GBIF-Occurrence="3914046340" box="[787,1083,1914,1941]" collectionCode="ZM-CBSU" country="Iran" isTailEnumeration="true" pageId="17" pageNumber="168" specimenCode="S003-134, 135" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
<collectionCode id="AFECDCE98A0DFF8BFCF6AB35FC164BFF" box="[787,911,1915,1941]" pageId="17" pageNumber="168">ZM-CBSU</collectionCode>
<specimenCode id="995BEC578A0DFF8BFC72AB34FB9B4BFF" box="[919,1026,1914,1941]" collectionCode="ZM-CBSU" pageId="17" pageNumber="168">S003-134</specimenCode>
–
<specimenCode id="995BEC578A0DFF8BFBF7AB34FBA24BFF" box="[1042,1083,1914,1941]" collectionCode="ZM-CBSU" pageId="17" pageNumber="168">135</specimenCode>
</materialsCitation>
(2).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0DFF8BFF72ABD1FF624BB7" pageId="17" pageNumber="168" type="etymology">
<paragraph id="C942442C8A0DFF8BFF72ABD1FF624BB7" blockId="17.[151,1436,1698,2013]" pageId="17" pageNumber="168">
<emphasis id="FB89983E8A0DFF8BFF72ABD1FEBB4BD3" bold="true" box="[151,290,1951,1977]" pageId="17" pageNumber="168">Etymology:</emphasis>
Named for
<collectingCountry id="B1EA04BC8A0DFF8BFE4CABD1FE694BD3" box="[425,496,1951,1977]" name="Iran" pageId="17" pageNumber="168">Persia</collectingCountry>
, the historic region of southwestern Asia that is associated with the area that is now
<collectingCountry id="B1EA04BC8A0DFF8BFF23AB8DFF6E4BB7" box="[198,247,1987,2013]" name="Iran" pageId="17" pageNumber="168">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0EFF88FF72ACD8FCFC4C93" pageId="18" pageNumber="169" type="distribution">
<paragraph id="C942442C8A0EFF88FF72ACD8FCFC4C93" blockId="18.[151,1436,150,429]" pageId="18" pageNumber="169">
<emphasis id="FB89983E8A0EFF88FF72ACD8FE584CDA" bold="true" box="[151,449,150,177]" pageId="18" pageNumber="169">Distribution and habitat:</emphasis>
This species is abundant on sandy bottoms in coastal areas of western South Caspian Sea (
<figureCitation id="51C658A98A0EFF88FF2BACF5FEBC4CBF" box="[206,293,187,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1648,1672]" captionTargetBox="[157,1433,1091,1624]" captionTargetId="figure-151@12.[154,1434,1090,1624]" captionTargetPageId="12" captionText="FIGURE 8. Distribution of (A) Anatirostrum profundorum, Babka gymnotrachelus, Benthophiloides brauneri, Benthophiloides turcomanus, and Benthophilus pinchuki, (B) Benthophilus baeri, and (C) Benthophilus casachicus, Benthophilus ctenolepidus, Benthophilus leptocephalus, and Benthophilus persicus." figureDoi="http://doi.org/10.5281/zenodo.7120086" httpUri="https://zenodo.org/record/7120086/files/figure.png" pageId="18" pageNumber="169">Fig. 8C</figureCitation>
). Eurybathic, depth ranges usually from
<quantity id="0E05E9C98A0EFF88FD03ACF5FCA44CBF" box="[742,829,187,213]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="7.0" metricValueMin="0.6" pageId="18" pageNumber="169" unit="m" value="38.0" valueMax="70.0" valueMin="6.0">
6–
<specimenCount id="DFFB8FA58A0EFF88FCE7ACF5FCA44CBF" box="[770,829,187,213]" pageId="18" pageNumber="169" type="generic">70 m</specimenCount>
</quantity>
. However, no specimens have yet been collected in the eastern part of the South Caspian Sea (
<bibRefCitation id="AD6C39DD8A0EFF88FD92AC91FCC14C93" author="Kovacic, M. &amp; Esmaeili, H. R. &amp; Zarei, F. &amp; Abbasi, K. &amp; Schliewen, U. K." box="[631,856,223,249]" pageId="18" pageNumber="169" pagination="45 - 63" refId="ref27887" refString="Kovacic, M., Esmaeili, H. R., Zarei, F., Abbasi, K. &amp; Schliewen, U. K. (2021) A new species of Tadpole goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea. Zootaxa, 4980 (1), 45 - 63. https: // doi. org / 10.11646 / zootaxa. 4980.1.3" type="journal article" year="2021">
Kovačić
<emphasis id="FB89983E8A0EFF88FD38AC91FC8C4C93" box="[733,789,223,249]" italics="true" pageId="18" pageNumber="169">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0EFF88FF72AD4CFE854DC7" pageId="18" pageNumber="169" type="materials_examined">
<paragraph id="C942442C8A0EFF88FF72AD4CFD7B4DE3" blockId="18.[151,1436,150,429]" pageId="18" pageNumber="169">
<emphasis id="FB89983E8A0EFF88FF72AD4CFE1A4D76" bold="true" box="[151,387,258,284]" pageId="18" pageNumber="169">Material examined:</emphasis>
<collectionCode id="AFECDCE98A0EFF88FE6CAD4DFE5E4D77" box="[393,455,259,285]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="18" pageNumber="169">PMR</collectionCode>
,
<collectionCode id="AFECDCE98A0EFF88FE37AD4DFDC84D77" box="[466,593,259,285]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
, and
<collectionCode id="AFECDCE98A0EFF88FD6EAD4DFD5C4D77" box="[651,709,259,285]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="18" pageNumber="169" type="Museum">ZSM</collectionCode>
(
<typeStatus id="1646FA8E8A0EFF88FD36AD4DFC9C4D77" box="[723,773,259,285]" pageId="18" pageNumber="169">type</typeStatus>
material), plus additional non-type material (
<bibRefCitation id="AD6C39DD8A0EFF88FB1AAD4DFE9B4D2B" author="Kovacic, M. &amp; Esmaeili, H. R. &amp; Zarei, F. &amp; Abbasi, K. &amp; Schliewen, U. K." pageId="18" pageNumber="169" pagination="45 - 63" refId="ref27887" refString="Kovacic, M., Esmaeili, H. R., Zarei, F., Abbasi, K. &amp; Schliewen, U. K. (2021) A new species of Tadpole goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea. Zootaxa, 4980 (1), 45 - 63. https: // doi. org / 10.11646 / zootaxa. 4980.1.3" type="journal article" year="2021">
Kovačić
<emphasis id="FB89983E8A0EFF88FA80AD4DFA0C4D77" box="[1381,1429,259,285]" italics="true" pageId="18" pageNumber="169">et al</emphasis>
. 2021
</bibRefCitation>
);
<materialsCitation id="79954E718A0EFF88FEF0AD69FD104D2B" ID-GBIF-Occurrence="3914046302" box="[277,649,295,321]" collectionCode="ZM-CBSU" latitude="-3.116" location="Anzali" pageId="18" pageNumber="169" specimenCount="1">
<collectionCode id="AFECDCE98A0EFF88FEF0AD69FE084D2B" box="[277,401,295,321]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S003, 116, off
<location id="CC2212F78A0EFF88FDDEAD69FD104D2B" LSID="urn:lsid:plazi:treatment:4154F53A8A0DFF88FF72AAECFE854DC7:CC2212F78A0EFF88FDDEAD69FD104D2B" box="[571,649,295,321]" latitude="-3.116" name="Anzali" pageId="18" pageNumber="169">Anzali</location>
</materialsCitation>
;
<materialsCitation id="79954E718A0EFF88FD76AD69FC604D2B" ID-GBIF-Occurrence="3914046414" box="[659,1017,294,321]" collectionCode="ZM-CBSU" latitude="-16.25" location="Astara" pageId="18" pageNumber="169" specimenCount="1">
<collectionCode id="AFECDCE98A0EFF88FD76AD69FC964D2B" box="[659,783,295,321]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S016, 25, off
<location id="CC2212F78A0EFF88FC49AD69FC604D2B" LSID="urn:lsid:plazi:treatment:4154F53A8A0DFF88FF72AAECFE854DC7:CC2212F78A0EFF88FC49AD69FC604D2B" box="[940,1017,295,321]" latitude="-16.25" name="Astara" pageId="18" pageNumber="169">Astara</location>
</materialsCitation>
;
<materialsCitation id="79954E718A0EFF88FBE6AD69FA014D2B" ID-GBIF-Occurrence="3914046387" box="[1027,1432,295,321]" collectionCode="ZM-CBSU" latitude="-22.3" location="Chaboksar" pageId="18" pageNumber="169" specimenCount="1">
<collectionCode id="AFECDCE98A0EFF88FBE6AD69FBE64D2B" box="[1027,1151,295,321]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S022, 30, off
<location id="CC2212F78A0EFF88FAF8AD69FA014D2B" LSID="urn:lsid:plazi:treatment:4154F53A8A0DFF88FF72AAECFE854DC7:CC2212F78A0EFF88FAF8AD69FA014D2B" box="[1309,1432,295,321]" latitude="-22.3" name="Chaboksar" pageId="18" pageNumber="169">Chaboksar</location>
</materialsCitation>
;
<materialsCitation id="79954E718A0EFF88FF23AD05FE3F4D0F" ID-GBIF-Occurrence="3914046352" box="[198,422,331,357]" collectionCode="ZM-CBSU" latitude="-24.1" pageId="18" pageNumber="169" specimenCount="1">
<collectionCode id="AFECDCE98A0EFF88FF23AD05FEDB4D0F" box="[198,322,331,357]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S024, 1
</materialsCitation>
, &amp;
<materialsCitation id="79954E718A0EFF88FE32AD05FC0B4D0F" ID-GBIF-Occurrence="3914046426" box="[471,914,330,357]" collectionCode="ZM-CBSU" latitude="-25.81" location="Chamkhaleh" pageId="18" pageNumber="169" specimenCount="1">
<collectionCode id="AFECDCE98A0EFF88FE32AD05FDCA4D0F" box="[471,595,331,357]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S025, 81, off
<location id="CC2212F78A0EFF88FCE4AD05FC0B4D0F" LSID="urn:lsid:plazi:treatment:4154F53A8A0DFF88FF72AAECFE854DC7:CC2212F78A0EFF88FCE4AD05FC0B4D0F" box="[769,914,331,357]" latitude="-25.81" name="Chamkhaleh" pageId="18" pageNumber="169">Chamkhaleh</location>
</materialsCitation>
;
<materialsCitation id="79954E718A0EFF88FC44AD05FD464DE3" ID-GBIF-Occurrence="3914046400" collectionCode="ZM-CBSU" collectorName="Shafaroud" country="Iran" latitude="-29.3" location="Talesh" pageId="18" pageNumber="169" specimenCount="1" stateProvince="Gilan">
<collectionCode id="AFECDCE98A0EFF88FC44AD05FB844D0F" box="[929,1053,331,357]" pageId="18" pageNumber="169">ZM-CBSU</collectionCode>
S026, 4,
<collectorName id="640821FA8A0EFF88FB77AD05FA9C4D0F" box="[1170,1285,331,357]" pageId="18" pageNumber="169">Shafaroud</collectorName>
mouth; ZMCBSU S029, 3, off
<location id="CC2212F78A0EFF88FE45AD21FE754DE3" LSID="urn:lsid:plazi:treatment:4154F53A8A0DFF88FF72AAECFE854DC7:CC2212F78A0EFF88FE45AD21FE754DE3" box="[416,492,367,393]" country="Iran" latitude="-29.3" name="Talesh" pageId="18" pageNumber="169" stateProvince="Gilan">Talesh</location>
,
<collectingRegion id="0B398ACE8A0EFF88FE12AD21FD3B4DE3" box="[503,674,367,393]" country="Iran" name="Gilan" pageId="18" pageNumber="169">Gilan Province</collectingRegion>
,
<collectingCountry id="B1EA04BC8A0EFF88FD4BAD21FD464DE3" box="[686,735,367,393]" name="Iran" pageId="18" pageNumber="169">Iran</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="C942442C8A0EFF88FF72ADDCFE854DC7" blockId="18.[151,1436,150,429]" box="[151,284,402,429]" pageId="18" pageNumber="169">
<emphasis id="FB89983E8A0EFF88FF72ADDCFF714DC6" bold="true" box="[151,232,402,428]" pageId="18" pageNumber="169">IUCN:</emphasis>
NE.
</paragraph>
</subSubSection>
</treatment>
</document>
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