1082 lines
157 KiB
XML
1082 lines
157 KiB
XML
<document id="63C68B78CBA783A9967CA4412B005611" ID-DOI="10.11646/zootaxa.4048.1.2" ID-GBIF-Dataset="2f5da4b3-c0ee-4b70-b661-0e2ef9bacfa5" ID-ISSN="1175-5326" ID-Zenodo-Dep="290075" ID-ZooBank="97BB73FF-9D62-4A15-B09C-0F570E26E738" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461341944160" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2015" docId="1B6F879DE15BBD7F54A1FF9DDB84891C" docLanguage="en" docName="zt04048p056.pdf" docOrigin="Zootaxa 4048 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma pontianak Ng, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="55" masterDocId="E756FFE5E150BD6D5436F955DF298F30" masterDocTitle="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species" masterLastPageNumber="56" masterPageNumber="37" pageNumber="48" updateTime="1698620270231" updateUser="plazi">
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<mods:mods id="B4510F90C498B6A4497765C0B8A7660D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8AC269CB542D05316ECD3A7129E7D007">
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<mods:title id="DD15D718307B65ABFB5C3B19277276EE">Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species</mods:title>
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<mods:name id="CD25F6CB30B85ED00F3D3450BECBE0EA" type="personal">
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<mods:role id="B5F638EE7D3E756866BCD0FE136E908F">
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<mods:roleTerm id="103C6E24C55A6184BF1CBC764D97496D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="831DDCCC72CE3CD2A8B477F70439D66F">Ng, Peter K. L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="694A51421854D7AB273A9594B90268F9">text</mods:typeOfResource>
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<mods:titleInfo id="2777FD89FFEC48364F0D430909B6D4AE">
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<mods:title id="0440F43C9E5470B714CB61C682096995">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6AAA7E6FA84E32658218C7A15AC0EB21">
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<mods:date id="30AB167729ACED5E87C01B65E3E16A83">2015</mods:date>
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<mods:detail id="6C5C5C7F9E88EB05927ECB7208521DE5" type="volume">
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<mods:number id="AD1B764BB9CD96C51566E3AC61019050">4048</mods:number>
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</mods:detail>
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<mods:detail id="DF5B6A33CD97842E2DCF39DF5DE7E9D6" type="issue">
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<mods:number id="B881F761C9C5F6526ABC32DB0706110B">1</mods:number>
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<mods:start id="124382686DFEAD4794EF4979CFA8785C">37</mods:start>
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<mods:end id="8A7CD6D88CBD932F68BA19C793401A35">56</mods:end>
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</mods:part>
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<mods:classification id="8146D511FF59A6246F86A9168C049A16">journal article</mods:classification>
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<mods:identifier id="CA3436FF2F118CC28A85E6B34FD52E1D" type="DOI">10.11646/zootaxa.4048.1.2</mods:identifier>
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<mods:identifier id="6B6A8B700DE8B820364078AA7CC19311" type="GBIF-Dataset">2f5da4b3-c0ee-4b70-b661-0e2ef9bacfa5</mods:identifier>
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<mods:identifier id="BA9DE439175E9777349CEA26958E0216" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="827C83C1A82F45948FAC059B7401F149" type="Zenodo-Dep">290075</mods:identifier>
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<mods:identifier id="4B24A04640DDB24ED0A4353826495291" type="ZooBank">97BB73FF-9D62-4A15-B09C-0F570E26E738</mods:identifier>
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</mods:mods>
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<treatment id="1B6F879DE15BBD7F54A1FF9DDB84891C" ID-DOI="http://doi.org/10.5281/zenodo.6121306" ID-GBIF-Taxon="127694316" ID-Zenodo-Dep="6121306" LSID="urn:lsid:plazi:treatment:1B6F879DE15BBD7F54A1FF9DDB84891C" httpUri="http://treatment.plazi.org/id/1B6F879DE15BBD7F54A1FF9DDB84891C" lastPageId="18" lastPageNumber="55" pageId="11" pageNumber="48">
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||
<subSubSection id="DBDC6500E15BBD6654A1FF9DDE058834" pageId="11" pageNumber="48" type="nomenclature">
|
||
<paragraph id="9379368BE15BBD6654A1FF9DDD3889D3" blockId="11.[151,529,1736,1796]" box="[151,529,1736,1763]" pageId="11" pageNumber="48">
|
||
<heading id="C83181E7E15BBD6654A1FF9DDD3889D3" bold="true" box="[151,529,1736,1763]" fontSize="11" level="1" pageId="11" pageNumber="48" reason="1">
|
||
<emphasis id="A1B2EA99E15BBD6654A1FF9DDD3889D3" bold="true" box="[151,529,1736,1763]" pageId="11" pageNumber="48">
|
||
<taxonomicName id="54C64D08E15BBD6654A1FF9DDE8589D2" ID-CoL="6KBJ5" box="[151,428,1736,1762]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15BBD6654A1FF9DDE8589D2" bold="true" box="[151,428,1736,1762]" italics="true" pageId="11" pageNumber="48">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E15BBD665585FF9CDD3889D3" box="[435,529,1737,1763]" pageId="11" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9379368BE15BBD6654A1FFBEDE058834" blockId="11.[151,529,1736,1796]" box="[151,300,1771,1796]" pageId="11" pageNumber="48">
|
||
(
|
||
<figureCitation id="0BFD2A0EE15BBD6654A9FFBEDE0D8834" box="[159,292,1771,1796]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="13.[151,250,1936,1958]" captionStartId-1="14.[151,250,636,658]" captionStartId-2="15.[151,250,1858,1880]" captionTargetBox-0="[168,1422,181,1924]" captionTargetBox-1="[151,1435,193,614]" captionTargetBox-2="[161,1422,463,1843]" captionTargetId-0="figure@13.[168,1423,181,1925]" captionTargetId-1="figure@14.[151,1436,193,615]" captionTargetId-2="figure@15.[151,1436,459,1849]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." captionText-1="FIGURE 12. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. Male abdomens. A, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 a); B, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 b); C paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c)." captionText-2="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri-0="https://zenodo.org/record/290086/files/figure.png" httpUri-1="https://zenodo.org/record/290087/files/figure.png" httpUri-2="https://zenodo.org/record/290088/files/figure.png" pageId="11" pageNumber="48">Figs. 11–13</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E15BBD6154A1FE66DA508E54" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48" type="materials_examined">
|
||
<paragraph id="9379368BE15BBD6154A1FE66DA508E54" blockId="11.[151,1436,1843,2012]" lastBlockId="12.[151,1436,151,357]" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48">
|
||
<emphasis id="A1B2EA99E15BBD6654A1FE66DE53887C" bold="true" box="[151,378,1843,1868]" pageId="11" pageNumber="48">Material examined</emphasis>
|
||
.
|
||
<typeStatus id="4C7D8829E15BBD6655BFFE66DEDF887C" box="[393,502,1843,1868]" pageId="11" pageNumber="48" type="holotype">Holotype</typeStatus>
|
||
: male (13.4 ×
|
||
<quantity id="543E9B6EE15BBD6656AFFE61DDD6887C" box="[665,767,1844,1868]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="11" pageNumber="48" unit="mm" value="13.2">13.2 mm</quantity>
|
||
) (
|
||
<collectionCode id="F5D7AE4EE15BBD665720FE61DC7D887C" box="[790,852,1844,1868]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="11" pageNumber="48">MZB</collectionCode>
|
||
Cru 4392), Gunung Kloncet, near km 67 milestone to
|
||
<taxonomicName id="54C64D08E15BBD66548AFE02DE068840" box="[188,303,1879,1904]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, on
|
||
<taxonomicName id="54C64D08E15BBD665546FE02DEC98840" box="[368,480,1879,1904]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
to Anjungan Road, Kampung Anjungan,
|
||
<geoCoordinate id="F6F2504CE15BBD6657DDFE0DDB478840" box="[1003,1134,1880,1904]" direction="north" orientation="latitude" pageId="11" pageNumber="48" precision="9" value="0.35033333">00°21.02’N</geoCoordinate>
|
||
<geoCoordinate id="F6F2504CE15BBD665048FE0DDA278840" box="[1150,1294,1880,1904]" direction="east" orientation="longitude" pageId="11" pageNumber="48" precision="9" value="109.18467">109°11.08’E</geoCoordinate>
|
||
, Kabupaten
|
||
<taxonomicName id="54C64D08E15BBD6654A1FE2EDE2388A4" box="[151,266,1915,1940]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, West Kalimantan, coll. native collectors, through dealer,
|
||
<date id="E778104BE15BBD665780FE2EDB5C88A4" box="[950,1141,1915,1940]" pageId="11" pageNumber="48" value="1999-09">September 1999</date>
|
||
.
|
||
<typeStatus id="4C7D8829E15BBD6650B2FE29DBDC88A4" box="[1156,1269,1916,1940]" pageId="11" pageNumber="48" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="85C0FD02E15BBD665133FE29DA5988A4" box="[1285,1392,1915,1940]" pageId="11" pageNumber="48" type="male">12 males</specimenCount>
|
||
,
|
||
<specimenCount id="85C0FD02E15BBD6651B6FE29DFC78888" pageId="11" pageNumber="48" type="female">12 females</specimenCount>
|
||
(
|
||
<collectionCode id="F5D7AE4EE15BBD6654C8FEF5DE158888" box="[254,316,1952,1976]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="11" pageNumber="48">MZB</collectionCode>
|
||
Cru 4393),
|
||
<specimenCount id="85C0FD02E15BBD6655F0FEF5DD108888" box="[454,569,1951,1976]" pageId="11" pageNumber="48" type="male">123 males</specimenCount>
|
||
(largest 13.9 ×
|
||
<quantity id="543E9B6EE15BBD6656DBFEF5DC7D8887" box="[749,852,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="11" pageNumber="48" unit="mm" value="13.7">13.7 mm</quantity>
|
||
, smallest 6.3 ×
|
||
<quantity id="543E9B6EE15BBD665031FECADB488888" box="[1031,1121,1951,1976]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="11" pageNumber="48" unit="mm" value="6.2">6.2 mm</quantity>
|
||
),
|
||
<specimenCount id="85C0FD02E15BBD665043FECADBD98888" box="[1141,1264,1951,1976]" pageId="11" pageNumber="48" type="female">69 females</specimenCount>
|
||
(largest 12.6 ×
|
||
<quantity id="543E9B6EE15BBD6654A1FE96DFD488EB" box="[151,253,1987,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="48" unit="mm" value="12.5">12.5 mm</quantity>
|
||
, smallest mature 8.2 ×
|
||
<quantity id="543E9B6EE15BBD665635FE91DD7788EC" box="[515,606,1988,2012]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="48" unit="mm" value="8.1">8.1 mm</quantity>
|
||
), 3 ovigerous females (
|
||
<collectionCode id="F5D7AE4EE15BBD665754FE91DCB388EC" box="[866,922,1988,2012]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="48">ZRC</collectionCode>
|
||
2015.0454), same data as
|
||
<typeStatus id="4C7D8829E15BBD6650F0FE96DA0288EC" box="[1222,1323,1987,2012]" pageId="11" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="85C0FD02E15BBD665101FE91DAB588EC" box="[1335,1436,1987,2012]" pageId="11" pageNumber="48" type="male">23 males</specimenCount>
|
||
(largest 13.7 ×
|
||
<quantity id="543E9B6EE15CBD615574F9CDDE838F80" box="[322,426,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="12" pageNumber="49" unit="mm" value="13.4">13.4 mm</quantity>
|
||
),
|
||
<specimenCount id="85C0FD02E15CBD61558BF9CDDD118F80" box="[445,568,151,176]" pageId="12" pageNumber="49" type="female">19 females</specimenCount>
|
||
(largest 14.5 ×
|
||
<quantity id="543E9B6EE15CBD6156DCF9CDDC7A8F80" box="[746,851,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="12" pageNumber="49" unit="mm" value="14.2">14.2 mm</quantity>
|
||
), 3 ovigerous females (
|
||
<collectionCode id="F5D7AE4EE15CBD61506EF9CDDBB98F80" box="[1112,1168,152,176]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="49">ZRC</collectionCode>
|
||
2015.0455), Anjungan,
|
||
<taxonomicName id="54C64D08E15CBD6154A1F9E9DE238FE5" box="[151,266,188,213]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, Kabupaten
|
||
<taxonomicName id="54C64D08E15CBD61559CF9E9DD348FE5" box="[426,541,188,213]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, West Kalimantan, coll. native collectors, from
|
||
<collectingCountry id="EBD1761BE15CBD61504DF9E9DBC48FE5" box="[1147,1261,188,213]" name="Singapore" pageId="12" pageNumber="49">Singapore</collectingCountry>
|
||
aquarium,
|
||
<date id="E778104BE15CBD6151B6F9E9DE7E8FC8" pageId="12" pageNumber="49" value="1999-09-25">25 September 1999</date>
|
||
. Non-types:
|
||
<specimenCount id="85C0FD02E15CBD6155C4F9B5DD678FC8" box="[498,590,223,248]" pageId="12" pageNumber="49" type="male">4 males</specimenCount>
|
||
(largest 13.4 × 13.0 mm),
|
||
<specimenCount id="85C0FD02E15CBD615796F98ADB3B8FC8" box="[928,1042,223,248]" pageId="12" pageNumber="49" type="female">6 females</specimenCount>
|
||
(largest 13.4 ×
|
||
<quantity id="543E9B6EE15CBD6150EAF98ADA6E8FC8" box="[1244,1351,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="12" pageNumber="49" unit="mm" value="12.6">12.6 mm</quantity>
|
||
) (
|
||
<collectionCode id="F5D7AE4EE15CBD615152F9B5DAB58FC8" box="[1380,1436,224,248]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="49">ZRC</collectionCode>
|
||
2015.0456),
|
||
<taxonomicName id="54C64D08E15CBD615510F851DEB08E2D" box="[294,409,260,285]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, Mandor area, near Anjungan, coll. local collectors,
|
||
<date id="E778104BE15CBD6157D9F851DB818E2C" box="[1007,1192,260,284]" pageId="12" pageNumber="49" value="1999-08-31">31 August 1999</date>
|
||
;
|
||
<specimenCount id="85C0FD02E15CBD615082F851DA3F8E2D" box="[1204,1302,260,285]" pageId="12" pageNumber="49" type="female">1 female</specimenCount>
|
||
(10.1 ×
|
||
<quantity id="543E9B6EE15CBD61514FF851DFED8E70" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="12" pageNumber="49" unit="mm" value="9.6">9.6 mm</quantity>
|
||
) (
|
||
<collectionCode id="F5D7AE4EE15CBD6154ECF87DDE3B8E70" box="[218,274,296,320]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="49">ZRC</collectionCode>
|
||
2015.0457), near Anjungan, Kalimantan Barat, coll. H.H. Tan & Y.Y.Goh,
|
||
<date id="E778104BE15CBD61506CF87DDBD08E70" box="[1114,1273,295,320]" pageId="12" pageNumber="49" value="1998-04-29">29 April 1998</date>
|
||
; 1 young male (7.6 ×
|
||
<quantity id="543E9B6EE15CBD6154D7F819DE118E54" box="[225,312,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="49" unit="mm" value="7.3">7.3 mm</quantity>
|
||
) (
|
||
<collectionCode id="F5D7AE4EE15CBD615578F819DEAF8E54" box="[334,390,332,356]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="49">ZRC</collectionCode>
|
||
2015.0458), near Anjungan, Kalimantan Barat, coll. H.H. Tan & Y.Y.Goh,
|
||
<date id="E778104BE15CBD6150E2F819DA5D8E54" box="[1236,1396,332,357]" pageId="12" pageNumber="49" value="1998-04-28">28 April 1998</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C7B96603E15CBD6154A1FCBBDCAF8913" httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="12" pageNumber="49" targetBox="[161,1424,395,1502]" targetPageId="12">
|
||
<paragraph id="9379368BE15CBD6154A1FCBBDCAF8913" blockId="12.[151,1435,1518,1571]" pageId="12" pageNumber="49">
|
||
<emphasis id="A1B2EA99E15CBD6154A1FCBBDE358934" bold="true" box="[151,284,1518,1540]" pageId="12" pageNumber="49">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E15CBD61551BFCBADD0C8934" box="[301,549,1518,1540]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15CBD61551BFCBADD0C8934" box="[301,549,1518,1540]" italics="true" pageId="12" pageNumber="49">Geosesarma ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15CBD615606FCBBDDAD8934" bold="true" box="[560,644,1518,1540]" pageId="12" pageNumber="49">
|
||
<taxonomicNameLabel id="BA8157E2E15CBD615606FCBBDDAD8934" box="[560,644,1518,1540]" pageId="12" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs
|
||
<emphasis id="A1B2EA99E15CBD615509FF5BDE578913" box="[319,382,1550,1571]" italics="true" pageId="12" pageNumber="49">in situ</emphasis>
|
||
. Specimens not preserved, photographs by Y. Kho.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DBDC6500E15CBD6354F1FF18DE298C06" lastPageId="14" lastPageNumber="51" pageId="12" pageNumber="49" type="diagnosis">
|
||
<paragraph id="9379368BE15CBD6154F1FF18DA5A88C2" blockId="12.[151,1437,1613,2034]" pageId="12" pageNumber="49">
|
||
<emphasis id="A1B2EA99E15CBD6154F1FF18DE138956" bold="true" box="[199,314,1613,1638]" pageId="12" pageNumber="49">Diagnosis</emphasis>
|
||
. Carapace almost square, width to length ratio 1.01–1.02, lateral margins subparallel (
|
||
<figureCitation id="0BFD2A0EE15CBD615130FF18DA7C8956" box="[1286,1365,1613,1638]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Fig. 11</figureCitation>
|
||
A, B); dorsal surface with well-defined regions, anterior regions covered with small rounded, flattened granules (
|
||
<figureCitation id="0BFD2A0EE15CBD615159FF24DF9A899E" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Fig. 11</figureCitation>
|
||
B); front deflexed, frontal lobes broad with gently convex margins; postfrontal, postorbital cristae prominent, very sharp, strong (
|
||
<figureCitation id="0BFD2A0EE15CBD615542FFECDEEC89E2" box="[372,453,1721,1746]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Fig. 11</figureCitation>
|
||
B, C); external orbital tooth triangular, outer margins curving anteriorly, tooth directed anteriorly, not extending past lateral margin (
|
||
<figureCitation id="0BFD2A0EE15CBD6156A0FF88DDCE89C6" box="[662,743,1757,1782]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Fig. 11</figureCitation>
|
||
B). Merus of third maxilliped subovate, subequal in length to ischium; exopod without trace of flagellum (
|
||
<figureCitation id="0BFD2A0EE15CBD6156AFFE54DDC5882A" box="[665,748,1793,1818]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="12" pageNumber="49">Fig. 13</figureCitation>
|
||
A). Outer surface of palm of adult male covered with small rounded granules; dorsal margin of dactylus with 11 or 12 tubercles (distal one smallest), each with pectinated tip (
|
||
<figureCitation id="0BFD2A0EE15CBD6154A9FE1CDFC68852" box="[159,239,1865,1890]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Fig. 11</figureCitation>
|
||
E, F). Ambulatory legs with relatively slender merus, with sharp subdistal spine on dorsal margin, surfaces slightly rugose (
|
||
<figureCitation id="0BFD2A0EE15CBD61557BFE38DE8E88B6" box="[333,423,1901,1926]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Figs. 11</figureCitation>
|
||
A, 13C). Male abdomen broadly triangular; somite 6 relatively broad, with gently convex lateral margins; telson semicircular (
|
||
<figureCitation id="0BFD2A0EE15CBD615601FEC4DDBA889A" box="[567,659,1937,1962]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="12" pageNumber="49">Figs. 11</figureCitation>
|
||
D, 12, 13B). G1 relatively stout, subdistal part of outer margin with distinct right-angled angled, prominent hump-like arch just before distal pectinated part; pectinated distal part elongate from lateral view, subspatuliform from marginal view, with small median cleft (
|
||
<figureCitation id="0BFD2A0EE15CBD61504CFE8CDBE188C2" box="[1146,1224,2009,2034]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="12" pageNumber="49">Fig. 13</figureCitation>
|
||
D–H, J, K, M).
|
||
</paragraph>
|
||
<caption id="C7B96603E15DBD6054A1FEC5DC7688D3" httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="13" pageNumber="50" targetBox="[168,1422,181,1924]" targetPageId="13">
|
||
<paragraph id="9379368BE15DBD6054A1FEC5DC7688D3" blockId="13.[151,1436,1936,2019]" pageId="13" pageNumber="50">
|
||
<emphasis id="A1B2EA99E15DBD6054A1FEC5DE3E8896" bold="true" box="[151,279,1936,1958]" pageId="13" pageNumber="50">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E15DBD605515FEC4DD258896" box="[291,524,1936,1958]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15DBD605515FEC4DD258896" box="[291,524,1936,1958]" italics="true" pageId="13" pageNumber="50">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E15DBD605625FEC5DD4B8896" box="[531,610,1936,1958]" pageId="13" pageNumber="50" rank="species">
|
||
<emphasis id="A1B2EA99E15DBD605625FEC5DD4B8896" bold="true" box="[531,610,1936,1958]" pageId="13" pageNumber="50">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C7B96603E15EBD6354A1FB29DE348DE0" httpUri="https://zenodo.org/record/290087/files/figure.png" pageId="14" pageNumber="51" targetBox="[151,1435,193,614]" targetPageId="14">
|
||
<paragraph id="9379368BE15EBD6354A1FB29DE348DE0" blockId="14.[151,1436,636,720]" pageId="14" pageNumber="51">
|
||
<emphasis id="A1B2EA99E15EBD6354A1FB29DE318DA2" bold="true" box="[151,280,636,658]" pageId="14" pageNumber="51">FIGURE 12</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E15EBD635512FB28DD278DA2" box="[292,526,636,658]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635512FB28DD278DA2" box="[292,526,636,658]" italics="true" pageId="14" pageNumber="51">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635622FB29DD4A8DA2" box="[532,611,636,658]" pageId="14" pageNumber="51" rank="species">
|
||
<emphasis id="A1B2EA99E15EBD635622FB29DD4A8DA2" bold="true" box="[532,611,636,658]" pageId="14" pageNumber="51">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Anjungan, Kalimantan Barat, Borneo. Male abdomens. A, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454a); B, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454b); C paratype male (11.1 × 10.8 mm) (ZRC 2015.0454c).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9379368BE15EBD6354F1FBACDE298C06" blockId="14.[151,1437,761,2025]" pageId="14" pageNumber="51">
|
||
<emphasis id="A1B2EA99E15EBD6354F1FBACDE338C22" bold="true" box="[199,282,761,786]" pageId="14" pageNumber="51">Colour</emphasis>
|
||
. The adult colours and patterns of
|
||
<taxonomicName id="54C64D08E15EBD635690FBAFDC1C8C22" box="[678,821,762,786]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635690FBAFDC1C8C22" box="[678,821,762,786]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63570BFBACDCB18C22" bold="true" box="[829,920,761,786]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63570BFBACDCB18C22" box="[829,920,761,786]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are similar to those of
|
||
<taxonomicName id="54C64D08E15EBD6350A9FBAFDA138C22" box="[1183,1338,762,786]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6350A9FBAFDA138C22" box="[1183,1338,762,786]" italics="true" pageId="14" pageNumber="51">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635174FBACDAB58C22" bold="true" box="[1346,1436,761,786]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635174FBACDAB58C22" box="[1346,1436,761,786]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE15EBD6354A9FA48DFDA8C06" box="[159,243,797,822]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="14" pageNumber="51">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E15EBD6354F1FA14DFDD8CD9" pageId="14" pageNumber="51" type="etymology">
|
||
<paragraph id="9379368BE15EBD6354F1FA14DEF28C92" blockId="14.[151,1437,761,2025]" pageId="14" pageNumber="51">
|
||
<emphasis id="A1B2EA99E15EBD6354F1FA14DE6F8C6A" bold="true" box="[199,326,833,858]" pageId="14" pageNumber="51">Etymology</emphasis>
|
||
. In Indonesian mythology, a “
|
||
<taxonomicName id="54C64D08E15EBD6356AEFA14DC248C6A" box="[664,781,833,858]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
” is a vampiric ghost. The name of the new species alludes to its eerie-looking yellow eyes and is also in reference to the
|
||
<typeStatus id="4C7D8829E15EBD63576FFA32DCA08C4F" box="[857,905,871,895]" pageId="14" pageNumber="51">type</typeStatus>
|
||
locality, the district of
|
||
<taxonomicName id="54C64D08E15EBD6350A4FA30DA2F8C4E" box="[1170,1286,869,894]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="9379368BE15EBD6354F1FAF8DFDD8CD9" blockId="14.[151,1437,761,2025]" pageId="14" pageNumber="51">
|
||
<emphasis id="A1B2EA99E15EBD6354F1FAF8DE0D8CF6" bold="true" box="[199,292,941,966]" pageId="14" pageNumber="51">Ecology</emphasis>
|
||
. According to local collectors (P. Yap, pers. comm.), the species was collected in the forest near small streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E15EBD7C54F1FAA0DC408EB8" lastPageId="17" lastPageNumber="54" pageId="14" pageNumber="51" type="discussion">
|
||
<paragraph id="9379368BE15EBD6354F1FAA0DC178BAE" blockId="14.[151,1437,761,2025]" pageId="14" pageNumber="51">
|
||
<emphasis id="A1B2EA99E15EBD6354F1FAA0DE1D8B3E" bold="true" box="[199,308,1013,1038]" pageId="14" pageNumber="51">Remarks</emphasis>
|
||
. Most of the species now known from Borneo do not have a flagellum on the exopod of the third maxilliped. The only three species that have this flagellum,
|
||
<taxonomicName id="54C64D08E15EBD635760FD4FDCD28B02" box="[854,1019,1050,1074]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E15EBD635760FD4FDCD28B02" box="[854,1019,1050,1074]" italics="true" pageId="14" pageNumber="51">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD63500FFD4FDBF68B02" box="[1081,1247,1050,1074]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD63500FFD4FDBF68B02" box="[1081,1247,1050,1074]" italics="true" pageId="14" pageNumber="51">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6350DFFD4CDA6F8B02" bold="true" box="[1257,1350,1049,1074]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6350DFFD4CDA6F8B02" box="[1257,1350,1049,1074]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD6351B2FD4FDE018B66" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E15EBD6351B2FD4FDE018B66" italics="true" pageId="14" pageNumber="51">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
, are also characterised by possessing a more trapezoidal carapace with the lateral margins diverging (
|
||
<figureCitation id="0BFD2A0EE15EBD6354A9FD34DFC58B4A" box="[159,236,1121,1146]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="14" pageNumber="51">Figs. 1</figureCitation>
|
||
A, B, 2A, 3A, B, 5A, B, 6A, B; Serène 1968). The remaining species from Borneo all have distinctly quadrate carapaces which are only slightly longer than wide.
|
||
</paragraph>
|
||
<paragraph id="9379368BE15EBD6354F1FDFFDC6A898A" blockId="14.[151,1437,761,2025]" pageId="14" pageNumber="51">
|
||
<taxonomicName id="54C64D08E15EBD6354F1FDFFDEF18BF2" box="[199,472,1194,1218]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6354F1FDFFDEF18BF2" box="[199,472,1194,1218]" italics="true" pageId="14" pageNumber="51">Geosesarma ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6355D4FDFCDD178BF2" bold="true" box="[482,574,1193,1218]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6355D4FDFCDD178BF2" box="[482,574,1193,1218]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD63564CFDFFDC258BF2" box="[634,780,1194,1218]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD63564CFDFFDC258BF2" box="[634,780,1194,1218]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635720FDFCDC5B8BF2" bold="true" box="[790,882,1193,1218]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635720FDFCDC5B8BF2" box="[790,882,1193,1218]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are each other’s closest relative, possessing the same colour pattern in life, as well as similar third maxilliped and G1 structures.
|
||
<taxonomicName id="54C64D08E15EBD635010FD9BDA1D8BD6" box="[1062,1332,1230,1254]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635010FD9BDA1D8BD6" box="[1062,1332,1230,1254]" italics="true" pageId="14" pageNumber="51">Geosesarma ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63510DFD98DABC8BD6" bold="true" box="[1339,1429,1229,1254]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63510DFD98DABC8BD6" box="[1339,1429,1229,1254]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, however, differs in the following aspects: carapace proportionately wider (width to length ratio 1.06–1.08,
|
||
<figureCitation id="0BFD2A0EE15EBD635109FDA4DA568A3A" box="[1343,1407,1265,1290]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Fig. 8</figureCitation>
|
||
B) (
|
||
<quantity id="543E9B6EE15EBD6354A9FC43DE048A1E" box="[159,301,1301,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.5781" metricValueMax="2.5908" metricValueMin="2.5654" pageId="14" pageNumber="51" unit="in" value="1.015" valueMax="1.02" valueMin="1.01">1.01–1.02 in</quantity>
|
||
<taxonomicName id="54C64D08E15EBD635502FC43DEEA8A1E" box="[308,451,1302,1326]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635502FC43DEEA8A1E" box="[308,451,1302,1326]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6355FCFC40DD0A8A1E" bold="true" box="[458,547,1301,1326]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6355FCFC40DD0A8A1E" box="[458,547,1301,1326]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD635607FC40DDA98A1E" box="[561,640,1301,1326]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Fig. 11</figureCitation>
|
||
B), the external orbital tooth is directed obliquely laterally (
|
||
<figureCitation id="0BFD2A0EE15EBD63512DFC40DA758A1E" box="[1307,1372,1301,1326]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Fig. 8</figureCitation>
|
||
A, B) (curving anteriorly in
|
||
<taxonomicName id="54C64D08E15EBD6355BBFC6FDD358A62" box="[397,540,1338,1362]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6355BBFC6FDD358A62" box="[397,540,1338,1362]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635614FC6CDD558A62" bold="true" box="[546,636,1337,1362]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635614FC6CDD558A62" box="[546,636,1337,1362]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD6356BFFC6CDDFF8A62" box="[649,726,1337,1362]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Fig. 11</figureCitation>
|
||
A, B), the postfrontal and postorbital cristae are distinct but not very sharp (
|
||
<figureCitation id="0BFD2A0EE15EBD635514FC08DE4C8A46" box="[290,357,1373,1398]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Fig. 8</figureCitation>
|
||
A–C) (very sharp and strong in
|
||
<taxonomicName id="54C64D08E15EBD6356EBFC0BDC478A46" box="[733,878,1374,1398]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6356EBFC0BDC478A46" box="[733,878,1374,1398]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63574FFC08DCFC8A46" bold="true" box="[889,981,1373,1398]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63574FFC08DCFC8A46" box="[889,981,1373,1398]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD6357D0FC08DB1C8A46" box="[998,1077,1373,1398]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Fig. 11</figureCitation>
|
||
A–C), the ambulatory meri are proportionately shorter and stouter (
|
||
<figureCitation id="0BFD2A0EE15EBD635600FCD4DDAC8AAA" box="[566,645,1409,1434]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Figs. 8</figureCitation>
|
||
A, 9C) (relatively longer and more slender in
|
||
<taxonomicName id="54C64D08E15EBD6350ABFCD7DA068AAA" box="[1181,1327,1410,1434]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6350ABFCD7DA068AAA" box="[1181,1327,1410,1434]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63510EFCD4DABC8AAA" bold="true" box="[1336,1429,1409,1434]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63510EFCD4DABC8AAA" box="[1336,1429,1409,1434]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD6354A1FCF0DFD88A8E" box="[151,241,1445,1470]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD6354CBFCF3DE308A8E" box="[253,281,1446,1470]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="14" pageNumber="51">13</figureCitation>
|
||
C), the male abdomen is proportionately narrower, especially somite 6, with the telson relatively shorter (
|
||
<figureCitation id="0BFD2A0EE15EBD6354A9FC9CDFC98AD2" box="[159,224,1481,1506]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Fig. 8</figureCitation>
|
||
D, 9B) (proportionately wider with a wide somite 6 and longer telson in
|
||
<taxonomicName id="54C64D08E15EBD635032FC9FDBBB8AD2" box="[1028,1170,1482,1506]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635032FC9FDBBB8AD2" box="[1028,1170,1482,1506]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6350AFFC9CDBD88AD2" bold="true" box="[1177,1265,1481,1506]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6350AFFC9CDBD88AD2" box="[1177,1265,1481,1506]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD6350C9FC9CDA7E8AD2" box="[1279,1367,1481,1506]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Figs. 11</figureCitation>
|
||
D, 12, 13B), the subterminal hump on the G1 is distinct but more sloping (
|
||
<figureCitation id="0BFD2A0EE15EBD6357A4FCB8DCFD8936" box="[914,980,1517,1542]" captionStart="FIGURE 9" captionStartId="11.[151,250,1514,1536]" captionTargetBox="[167,1418,200,1477]" captionTargetId="figure@11.[150,1436,193,1493]" captionTargetPageId="11" captionText="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." httpUri="https://zenodo.org/record/290084/files/figure.png" pageId="14" pageNumber="51">Fig. 9</figureCitation>
|
||
D–H) (usually sharply right-angled in
|
||
<taxonomicName id="54C64D08E15EBD6351B2FCBBDE2E891A" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6351B2FCBBDE2E891A" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635538FF44DE41891A" bold="true" box="[270,360,1553,1578]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635538FF44DE41891A" box="[270,360,1553,1578]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE15EBD635540FF44DEED891A" box="[374,452,1553,1578]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="14" pageNumber="51">Fig. 13</figureCitation>
|
||
D–H, J, K, M). The angle of the hump in
|
||
<taxonomicName id="54C64D08E15EBD6357ACFF47DB00891A" box="[922,1065,1554,1578]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6357ACFF47DB00891A" box="[922,1065,1554,1578]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635006FF44DBA3891A" bold="true" box="[1072,1162,1553,1578]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635006FF44DBA3891A" box="[1072,1162,1553,1578]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
varies slightly, and may be less distinctly right-angled, especially in smaller specimens (e.g.,
|
||
<figureCitation id="0BFD2A0EE15EBD6357FAFF60DB08897E" box="[972,1057,1589,1614]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="14" pageNumber="51">Fig. 13</figureCitation>
|
||
L). In such cases, the G1 of
|
||
<taxonomicName id="54C64D08E15EBD6351B2FF63DE2E8942" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6351B2FF63DE2E8942" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635538FF0CDE408942" bold="true" box="[270,361,1625,1650]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635538FF0CDE408942" box="[270,361,1625,1650]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD635596FF0FDD128942" box="[416,571,1626,1650]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635596FF0FDD128942" box="[416,571,1626,1650]" italics="true" pageId="14" pageNumber="51">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635674FF0CDDB58942" bold="true" box="[578,668,1625,1650]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635674FF0CDDB58942" box="[578,668,1625,1650]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are almost indistinguishable. The two species, however, can still be easily distinguished by the form of the carapaces (more quadrate in
|
||
<taxonomicName id="54C64D08E15EBD63579CFF2BDB1589A6" box="[938,1084,1662,1686]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD63579CFF2BDB1589A6" box="[938,1084,1662,1686]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635073FF28DB8889A6" bold="true" box="[1093,1185,1661,1686]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635073FF28DB8889A6" box="[1093,1185,1661,1686]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) and ambulatory legs (more slender and relatively longer in
|
||
<taxonomicName id="54C64D08E15EBD635672FFF7DDFA898A" box="[580,723,1698,1722]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635672FFF7DDFA898A" box="[580,723,1698,1722]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6356ECFFF4DC1A898A" bold="true" box="[730,819,1697,1722]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6356ECFFF4DC1A898A" box="[730,819,1697,1722]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9379368BE15EBD6254F1FF90DABF8EAA" blockId="14.[151,1437,761,2025]" lastBlockId="15.[151,1437,151,410]" lastPageId="15" lastPageNumber="52" pageId="14" pageNumber="51">
|
||
The three species currently known from Sabah,
|
||
<taxonomicName id="54C64D08E15EBD6356D7FF93DC5C89EE" box="[737,885,1734,1758]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="aurantium">
|
||
<emphasis id="A1B2EA99E15EBD6356D7FF93DC5C89EE" box="[737,885,1734,1758]" italics="true" pageId="14" pageNumber="51">G. aurantium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="54C64D08E15EBD6357B5FF93DB0989EE" box="[899,1056,1734,1758]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="danumense">
|
||
<emphasis id="A1B2EA99E15EBD6357B5FF93DB0989EE" box="[899,1056,1734,1758]" italics="true" pageId="14" pageNumber="51">G. danumense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD635061FF93DBF289EE" box="[1111,1243,1734,1758]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="sabanum">
|
||
<emphasis id="A1B2EA99E15EBD635061FF93DBF289EE" box="[1111,1243,1734,1758]" italics="true" pageId="14" pageNumber="51">G. sabanum</emphasis>
|
||
</taxonomicName>
|
||
are distinct from
|
||
<taxonomicName id="54C64D08E15EBD6354A1FFBDDE1B89CF" box="[151,306,1767,1791]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6354A1FFBDDE1B89CF" box="[151,306,1767,1791]" italics="true" pageId="14" pageNumber="51">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63550CFFB2DEBC8830" box="[314,405,1767,1792]" pageId="14" pageNumber="51" rank="species">
|
||
<emphasis id="A1B2EA99E15EBD63550CFFB2DEBC8830" bold="true" box="[314,405,1767,1792]" pageId="14" pageNumber="51">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD6355F8FFBDDD7789CF" box="[462,606,1767,1791]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6355F8FFBDDD7789CF" box="[462,606,1767,1791]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635650FFB2DDE88830" bold="true" box="[614,705,1767,1792]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635650FFB2DDE88830" box="[614,705,1767,1792]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in that the dorsal margin of the dactylus of the adult male chela has only low proximal granules and not a row of tubercles (cf. Ng 1992: fig. 1D; Ng 1995a: fig. 1C; Ng 2002: fig. 2b). In
|
||
<taxonomicName id="54C64D08E15EBD6354DCFE7FDEAD8872" box="[234,388,1834,1858]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6354DCFE7FDEAD8872" box="[234,388,1834,1858]" italics="true" pageId="14" pageNumber="51">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD6355BAFE7CDECF8872" bold="true" box="[396,486,1833,1858]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD6355BAFE7CDECF8872" box="[396,486,1833,1858]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the dactylus has 11 or 12 such tubercles on the dorsal margin (
|
||
<figureCitation id="0BFD2A0EE15EBD635080FE7CDBD18872" box="[1206,1272,1833,1858]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="14" pageNumber="51">Fig. 8</figureCitation>
|
||
E, F), whereas
|
||
<taxonomicName id="54C64D08E15EBD6354A1FE1EDE0C8853" box="[151,293,1867,1891]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6354A1FE1EDE0C8853" box="[151,293,1867,1891]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63551AFE1EDEAC8854" bold="true" box="[300,389,1867,1892]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63551AFE1EDEAC8854" box="[300,389,1867,1892]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has 12 or 13 tubercles (
|
||
<figureCitation id="0BFD2A0EE15EBD6356A6FE1EDDF78853" box="[656,734,1867,1892]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="14" pageNumber="51">Fig. 11</figureCitation>
|
||
E, F).
|
||
<taxonomicName id="54C64D08E15EBD635712FE1EDB028853" box="[804,1067,1867,1891]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="aurantium">
|
||
<emphasis id="A1B2EA99E15EBD635712FE1EDB028853" box="[804,1067,1867,1891]" italics="true" pageId="14" pageNumber="51">Geosesarma aurantium</emphasis>
|
||
</taxonomicName>
|
||
(southern Sabah) differs from
|
||
<taxonomicName id="54C64D08E15EBD6351B2FE1EDE3B88B5" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD6351B2FE1EDE3B88B5" italics="true" pageId="14" pageNumber="51">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD63552CFE39DE5C88B5" bold="true" box="[282,373,1900,1925]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD63552CFE39DE5C88B5" box="[282,373,1900,1925]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E15EBD635598FE38DD1688B5" box="[430,575,1901,1925]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15EBD635598FE38DD1688B5" box="[430,575,1901,1925]" italics="true" pageId="14" pageNumber="51">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15EBD635671FE39DD8A88B5" bold="true" box="[583,675,1900,1925]" pageId="14" pageNumber="51">
|
||
<taxonomicNameLabel id="BA8157E2E15EBD635671FE39DD8A88B5" box="[583,675,1900,1925]" pageId="14" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a less developed frontal margin with the external orbital teeth more acutely triangular in shape (cf. Ng 1995a: fig. 1A), anterior parts of the dorsal surface of the carapace is relatively less granular (cf.
|
||
<bibRefCitation id="F7574B7AE15EBD6355E3FEFADDEC88F7" author="Ebin" box="[469,709,1967,1992]" pageId="14" pageNumber="51" refString="Ebin, J. & Chung, A. Y. C. (2012) Revisiting the elegant Bornean endemic crab of Mt Silam, Sabah. Malayan Naturalist, 65 (4), 50 - 51." type="journal article" year="2012">Ebin & Chung 2012</bibRefCitation>
|
||
), the ambulatory legs are proportionately much shorter (cf. Ng 1995a: fig. 1E, F), the male telson is recessed in the broad distal margin of somite 6 (cf. Ng 1995a: fig. 1H), and the distal pectinated part of the G1 is spatuliform and flared (cf. Ng 1995a: fig. 3).
|
||
<taxonomicName id="54C64D08E15FBD62501DF9CDDA148F80" box="[1067,1341,152,176]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="danumense">
|
||
<emphasis id="A1B2EA99E15FBD62501DF9CDDA148F80" box="[1067,1341,152,176]" italics="true" pageId="15" pageNumber="52">Geosesarma danumense</emphasis>
|
||
</taxonomicName>
|
||
(central Sabah) differs from
|
||
<taxonomicName id="54C64D08E15FBD625549F9ECDD328FE1" box="[383,539,185,209]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15FBD625549F9ECDD328FE1" box="[383,539,185,209]" italics="true" pageId="15" pageNumber="52">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15FBD625612F9ECDDA98FE2" bold="true" box="[548,640,185,210]" pageId="15" pageNumber="52">
|
||
<taxonomicNameLabel id="BA8157E2E15FBD625612F9ECDDA98FE2" box="[548,640,185,210]" pageId="15" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having the anterior dorsal carapace surface distinctly less strongly granular (cf. Ng 2002: fig. 1a, b), an external orbital tooth that is broadly triangular (cf. Ng 2002: fig. 1a, b, 3A), the ambulatory meri are more slender and relatively longer (cf. Ng 2002: fig. 1a, 3B), the male telson is recessed in the distal margin of somite 6 (cf. Ng 2002: fig. 2c, 3C), and the G1 is proportionately more slender, with the distal pectinated part spatuliform with a subdistal projection (cf. Ng 2002: fig. 3D–H).
|
||
<taxonomicName id="54C64D08E15FBD625002F86ADA118E66" box="[1076,1336,318,342]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15FBD625002F86ADA118E66" box="[1076,1336,318,342]" italics="true" pageId="15" pageNumber="52">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15FBD625177F86BDAB58E67" bold="true" box="[1345,1436,318,343]" pageId="15" pageNumber="52">
|
||
<taxonomicNameLabel id="BA8157E2E15FBD625177F86BDAB58E67" box="[1345,1436,318,343]" pageId="15" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="54C64D08E15FBD625514F835DE948E48" box="[290,445,352,376]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="danumense">
|
||
<emphasis id="A1B2EA99E15FBD625514F835DE948E48" box="[290,445,352,376]" italics="true" pageId="15" pageNumber="52">G. danumense</emphasis>
|
||
</taxonomicName>
|
||
in all these characters as well, except that the ambulatory legs are similar in length (
|
||
<figureCitation id="0BFD2A0EE15FBD625153F80ADF998EA9" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="15" pageNumber="52">Figs. 11</figureCitation>
|
||
A, 13C versus Ng 2002: figs. 1a, 3B).
|
||
<taxonomicName id="54C64D08E15FBD62567FF8D4DC1F8EA9" box="[585,822,385,409]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="sabanum">
|
||
<emphasis id="A1B2EA99E15FBD62567FF8D4DC1F8EA9" box="[585,822,385,409]" italics="true" pageId="15" pageNumber="52">Geosesarma sabanum</emphasis>
|
||
</taxonomicName>
|
||
(from eastern Sabah) differs from
|
||
<taxonomicName id="54C64D08E15FBD62509EF8D4DA138EA9" box="[1192,1338,385,409]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15FBD62509EF8D4DA138EA9" box="[1192,1338,385,409]" italics="true" pageId="15" pageNumber="52">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E15FBD625177F8D4DABF8EAA" bold="true" box="[1345,1430,385,410]" pageId="15" pageNumber="52">
|
||
<taxonomicNameLabel id="BA8157E2E15FBD625177F8D4DABF8EAA" box="[1345,1430,385,410]" pageId="15" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="C7B96603E15FBD6254A1FE17DD5788C1" httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="15" pageNumber="52" targetBox="[161,1422,463,1843]" targetPageId="15">
|
||
<paragraph id="9379368BE15FBD6254A1FE17DD5788C1" blockId="15.[151,1436,1858,2033]" pageId="15" pageNumber="52">
|
||
<emphasis id="A1B2EA99E15FBD6254A1FE17DE328868" bold="true" box="[151,283,1858,1880]" pageId="15" pageNumber="52">FIGURE 13</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E15FBD62551CFE16DD3F8868" box="[298,534,1858,1880]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="52" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E15FBD62551CFE16DD3F8868" box="[298,534,1858,1880]" italics="true" pageId="15" pageNumber="52">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E15FBD625629FE17DD5B8868" box="[543,626,1858,1880]" pageId="15" pageNumber="52" rank="species">
|
||
<emphasis id="A1B2EA99E15FBD625629FE17DD5B8868" bold="true" box="[543,626,1858,1880]" pageId="15" pageNumber="52">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Anjungan, Kalimantan Barat, Borneo. A–I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G1 (ventral view); E, left G1 (dorsal view); F, K, M, distal part of left G1 (ventral view); G, distal part of left G1 (dorsal view); H, distal part of left G1 (subdorsal view); I, left G2. All structures (except C) denuded. Scales: A– C, D, E, I, J, L = 1.0 mm, F–H, K, M = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C7B96603E140BD7D54A1FEC6DBBD88D9" httpUri="https://zenodo.org/record/290089/files/figure.png" pageId="16" pageNumber="53" targetBox="[150,1436,179,1916]" targetPageId="16">
|
||
<paragraph id="9379368BE140BD7D54A1FEC6DBBD88D9" blockId="16.[151,1436,1939,2025]" pageId="16" pageNumber="53">
|
||
<emphasis id="A1B2EA99E140BD7D54A1FEC6DE0A889A" bold="true" box="[151,291,1939,1964]" pageId="16" pageNumber="53">FIGURE 14</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E140BD7D5505FEC0DDC2889A" authority="De Man, 1902" authorityName="De Man" authorityYear="1902" box="[307,747,1940,1962]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="53" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E140BD7D5505FEC0DD1E889A" box="[307,567,1940,1962]" italics="true" pageId="16" pageNumber="53">Geosesarma gracillimum</emphasis>
|
||
(De Man, 1902)
|
||
</taxonomicName>
|
||
, male (12.1 × 11.8 mm) (ZRC 2007.262), Lambir National Park, Sarawak, Malaysia Sarawak, Malaysia. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9379368BE141BD7C54A1F9C2DC408EB8" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="54">
|
||
in having an external orbital tooth that is more acutely triangular and separated from rest of the margin by a low indentation, not by a cleft (cf. Ng 1992: fig. 1A), and the G1 has a short subdistal projection (cf. Ng 1992: fig. 1G, I, J).
|
||
<taxonomicName id="54C64D08E141BD7C54E2F9B5DEF08FC8" box="[212,473,224,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C54E2F9B5DEF08FC8" box="[212,473,224,248]" italics="true" pageId="17" pageNumber="54">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C55D7F98ADD148FC8" bold="true" box="[481,573,223,248]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C55D7F98ADD148FC8" box="[481,573,223,248]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also differs from
|
||
<taxonomicName id="54C64D08E141BD7C573AF9B5DCB88FC8" box="[780,913,224,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="sabanum">
|
||
<emphasis id="A1B2EA99E141BD7C573AF9B5DCB88FC8" box="[780,913,224,248]" italics="true" pageId="17" pageNumber="54">G. sabanum</emphasis>
|
||
</taxonomicName>
|
||
in these characters except that its ambulatory legs are proportionately longer (
|
||
<figureCitation id="0BFD2A0EE141BD7C5637F851DD738E2C" box="[513,602,260,285]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Figs. 11</figureCitation>
|
||
A, 13C versus Ng 1992: fig. 1C). The colour of
|
||
<taxonomicName id="54C64D08E141BD7C504DF851DA268E2C" box="[1147,1295,260,284]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="aurantium">
|
||
<emphasis id="A1B2EA99E141BD7C504DF851DA268E2C" box="[1147,1295,260,284]" italics="true" pageId="17" pageNumber="54">G. aurantium</emphasis>
|
||
</taxonomicName>
|
||
is strikingly different from
|
||
<taxonomicName id="54C64D08E141BD7C550AF87DDEFE8E70" box="[316,471,296,320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C550AF87DDEFE8E70" box="[316,471,296,320]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C55E8F872DD108E70" bold="true" box="[478,569,295,320]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C55E8F872DD108E70" box="[478,569,295,320]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, being bright yellowish-orange in life (Ng 1995a;
|
||
<bibRefCitation id="F7574B7AE141BD7C5042F872DA748E70" author="Ebin" box="[1140,1373,295,320]" pageId="17" pageNumber="54" refString="Ebin, J. & Chung, A. Y. C. (2012) Revisiting the elegant Bornean endemic crab of Mt Silam, Sabah. Malayan Naturalist, 65 (4), 50 - 51." type="journal article" year="2012">Ebin & Chung 2012</bibRefCitation>
|
||
). The carapace of
|
||
<taxonomicName id="54C64D08E141BD7C5519F819DEFC8E54" box="[303,469,332,356]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="danumense">
|
||
<emphasis id="A1B2EA99E141BD7C5519F819DEFC8E54" box="[303,469,332,356]" italics="true" pageId="17" pageNumber="54">G. danumense</emphasis>
|
||
</taxonomicName>
|
||
is orangish-brown in life, with the legs purplish brown and the chelae white (unpublished data). The live colour of
|
||
<taxonomicName id="54C64D08E141BD7C5670F825DDE38EB8" box="[582,714,368,392]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="sabanum">
|
||
<emphasis id="A1B2EA99E141BD7C5670F825DDE38EB8" box="[582,714,368,392]" italics="true" pageId="17" pageNumber="54">G. sabanum</emphasis>
|
||
</taxonomicName>
|
||
is not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E141BD7F54F1F8C1DB84891C" lastPageId="18" lastPageNumber="55" pageId="17" pageNumber="54" type="reference_group">
|
||
<paragraph id="9379368BE141BD7C54F1F8C1DD598A65" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="54">
|
||
Like
|
||
<taxonomicName id="54C64D08E141BD7C5530F8C1DE8D8E9C" ID-CoL="3FS8P" box="[262,420,404,428]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5530F8C1DE8D8E9C" box="[262,420,404,428]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5599F8C1DD248E9D" bold="true" box="[431,525,404,429]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5599F8C1DD248E9D" box="[431,525,404,429]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE141BD7C5617F8C1DD4E8E9C" box="[545,615,404,429]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="17" pageNumber="54">Fig. 8</figureCitation>
|
||
E, F) and
|
||
<taxonomicName id="54C64D08E141BD7C56D2F8C1DC5E8E9C" ID-CoL="6KBJ5" box="[740,887,404,428]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C56D2F8C1DC5E8E9C" box="[740,887,404,428]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C57B5F8C1DCC88E9D" bold="true" box="[899,993,404,429]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C57B5F8C1DCC88E9D" box="[899,993,404,429]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE141BD7C57C3F8C1DB618E9C" box="[1013,1096,404,429]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Fig. 11</figureCitation>
|
||
E, F), the three species from
|
||
<collectingCountry id="EBD1761BE141BD7C54A1F8E2DFD38EE0" box="[151,250,439,464]" name="Malaysia" pageId="17" pageNumber="54">Sarawak</collectingCountry>
|
||
,
|
||
<emphasis id="A1B2EA99E141BD7C5533F8EDDE628EE0" box="[261,331,440,464]" italics="true" pageId="17" pageNumber="54">G. b a u</emphasis>
|
||
,
|
||
<taxonomicName id="54C64D08E141BD7C556BF8EDDD2C8EE0" ID-CoL="3FS99" box="[349,517,440,464]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E141BD7C556BF8EDDD2C8EE0" box="[349,517,440,464]" italics="true" pageId="17" pageNumber="54">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E141BD7C560AF8EDDD848EE0" ID-CoL="8YX9B" box="[572,685,440,464]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="A1B2EA99E141BD7C560AF8EDDD848EE0" box="[572,685,440,464]" italics="true" pageId="17" pageNumber="54">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
all have 9–11 tubercles on the dorsal margin of the dactylus of the male chela (
|
||
<figureCitation id="0BFD2A0EE141BD7C5516F889DE588EC4" box="[288,369,476,501]" captionStart="FIGURE 14" captionStartId="16.[151,256,1939,1964]" captionTargetBox="[150,1436,179,1916]" captionTargetId="figure@16.[150,1437,179,1918]" captionTargetPageId="16" captionText="FIGURE 14. Geosesarma gracillimum (De Man, 1902), male (12.1 × 11.8 mm) (ZRC 2007.262), Lambir National Park, Sarawak, Malaysia Sarawak, Malaysia. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290089/files/figure.png" pageId="17" pageNumber="54">Fig. 14</figureCitation>
|
||
E, F;
|
||
<bibRefCitation id="F7574B7AE141BD7C5599F888DDBF8EC4" author="Ng" box="[431,662,476,501]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, 59, New Series No. 80, Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: 320; Ng 1995b: figs. 11C, D, 13A).
|
||
<taxonomicName id="54C64D08E141BD7C500EF889DBDF8EC4" ID-CoL="3FS8V" box="[1080,1270,476,500]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="A1B2EA99E141BD7C500EF889DBDF8EC4" box="[1080,1270,476,500]" italics="true" pageId="17" pageNumber="54">Geosesarma bau</emphasis>
|
||
</taxonomicName>
|
||
(from western
|
||
<collectingCountry id="EBD1761BE141BD7C54A1F8AADFD08D28" box="[151,249,511,536]" name="Malaysia" pageId="17" pageNumber="54">Sarawak</collectingCountry>
|
||
) differs from
|
||
<taxonomicName id="54C64D08E141BD7C55AEFB55DD1A8D28" ID-CoL="3FS8P" box="[408,563,512,536]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C55AEFB55DD1A8D28" box="[408,563,512,536]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C560DF8AADDBC8D28" bold="true" box="[571,661,511,536]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C560DF8AADDBC8D28" box="[571,661,511,536]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having ambulatory legs that are proportionately longer (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C516AFB55DE1E8D0C" author="Ng" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, 59, New Series No. 80, Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 8A, B), and the G1 is proportionately more slender without any subdistal hump, with the distal pectinated part only gently bent and subcylindrical in shape (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C57BEFB1DDB5E8D50" author="Ng" box="[904,1143,583,608]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, 59, New Series No. 80, Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 9D–F).
|
||
<taxonomicName id="54C64D08E141BD7C5126FB1DDE2E8DB4" ID-CoL="6KBJ5" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5126FB1DDE2E8DB4" italics="true" pageId="17" pageNumber="54">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5527FB39DE478DB5" bold="true" box="[273,366,620,645]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5527FB39DE478DB5" box="[273,366,620,645]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<emphasis id="A1B2EA99E141BD7C55C5FB39DD158DB4" box="[499,572,620,644]" italics="true" pageId="17" pageNumber="54">G. b a u</emphasis>
|
||
in the proportions of the ambulatory legs, but its external orbital tooth is directed anteriorly with the outer margins curving anteriorly (
|
||
<figureCitation id="0BFD2A0EE141BD7C5759FBDADCEB8D98" box="[879,962,655,680]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Fig. 11</figureCitation>
|
||
A, B) (directed obliquely with the outer margin gently convex in
|
||
<emphasis id="A1B2EA99E141BD7C558EFBE1DD298DFC" box="[440,512,692,716]" italics="true" pageId="17" pageNumber="54">G. b a u</emphasis>
|
||
,
|
||
<bibRefCitation id="F7574B7AE141BD7C5622FBE0DC298DFC" author="Ng" box="[532,768,692,717]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, 59, New Series No. 80, Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 9A, B); and the G1 is relatively much stouter with a distinct subdistal hump (
|
||
<figureCitation id="0BFD2A0EE141BD7C559BFB82DED58DC0" box="[429,508,727,752]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="17" pageNumber="54">Fig. 13</figureCitation>
|
||
D–H, J–M) (slender without a subdistal hump in
|
||
<taxonomicName id="54C64D08E141BD7C501DFB8DDA438DC0" ID-CoL="3FS8V" authority="Ng & Grinang 2004" authorityName="Ng & Grinang" authorityYear="2004" box="[1067,1386,727,752]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="A1B2EA99E141BD7C501DFB8DDB5D8DC0" box="[1067,1140,728,752]" italics="true" pageId="17" pageNumber="54">G. bau</emphasis>
|
||
,
|
||
<bibRefCitation id="F7574B7AE141BD7C50B5FB8DDA438DC0" author="Ng" box="[1155,1386,727,752]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity, Sarawak Museum Journal, 59, New Series No. 80, Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 9D–F).
|
||
<taxonomicName id="54C64D08E141BD7C54D8FBA9DD238C24" ID-CoL="3FS99" box="[238,522,764,788]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E141BD7C54D8FBA9DD238C24" box="[238,522,764,788]" italics="true" pageId="17" pageNumber="54">Geosesarma gracillimum</emphasis>
|
||
</taxonomicName>
|
||
(northern
|
||
<collectingCountry id="EBD1761BE141BD7C56B4FBA9DDCB8C25" box="[642,738,764,789]" name="Malaysia" pageId="17" pageNumber="54">Sarawak</collectingCountry>
|
||
and
|
||
<collectingCountry id="EBD1761BE141BD7C572FFBA9DC4E8C25" box="[793,871,764,789]" name="Brunei" pageId="17" pageNumber="54">Brunei</collectingCountry>
|
||
) can be distinguished from
|
||
<taxonomicName id="54C64D08E141BD7C5097FBA9DA128C24" ID-CoL="3FS8P" box="[1185,1339,764,788]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5097FBA9DA128C24" box="[1185,1339,764,788]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5174FBA9DAB58C25" bold="true" box="[1346,1436,764,789]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5174FBA9DAB58C25" box="[1346,1436,764,789]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a more distinctly quadrate carapace, the ambulatory legs are proportionately longer and male abdominal somite 6 is relatively wider (
|
||
<figureCitation id="0BFD2A0EE141BD7C55E3FA11DD0C8C6C" box="[469,549,836,861]" captionStart="FIGURE 14" captionStartId="16.[151,256,1939,1964]" captionTargetBox="[150,1436,179,1916]" captionTargetId="figure@16.[150,1437,179,1918]" captionTargetPageId="16" captionText="FIGURE 14. Geosesarma gracillimum (De Man, 1902), male (12.1 × 11.8 mm) (ZRC 2007.262), Lambir National Park, Sarawak, Malaysia Sarawak, Malaysia. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290089/files/figure.png" pageId="17" pageNumber="54">Fig. 14</figureCitation>
|
||
A, B, D). Most significantly, the G1 of
|
||
<taxonomicName id="54C64D08E141BD7C57ECFA11DBA88C6C" ID-CoL="3FS99" box="[986,1153,836,860]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E141BD7C57ECFA11DBA88C6C" box="[986,1153,836,860]" italics="true" pageId="17" pageNumber="54">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
differs markedly from
|
||
<taxonomicName id="54C64D08E141BD7C51B2FA11DE3B8CB0" ID-CoL="3FS8P" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C51B2FA11DE3B8CB0" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C552AFA32DE508CB0" bold="true" box="[284,377,871,896]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C552AFA32DE508CB0" box="[284,377,871,896]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E141BD7C5581FA3DDD638CB0" ID-CoL="6KBJ5" box="[439,586,872,896]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5581FA3DDD638CB0" box="[439,586,872,896]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5662FA32DD9B8CB0" bold="true" box="[596,690,871,896]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5662FA32DD9B8CB0" box="[596,690,871,896]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in that it has no subdistal hump, with the distal pectinated part subcylindrical and tapering to subtruncate tip (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C56E7FAD9DC5F8C94" author="Holthuis" box="[721,886,908,933]" pageId="17" pageNumber="54" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 3 - 47, figs. 1 - 9, pls. 1 - 8." type="journal article" year="1979">Holthuis 1979</bibRefCitation>
|
||
: fig. 9b, c; Ng 1995b: fig. 13B–E).
|
||
<taxonomicName id="54C64D08E141BD7C5126FAD9DFC18CF8" ID-CoL="8YX9B" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="A1B2EA99E141BD7C5126FAD9DFC18CF8" italics="true" pageId="17" pageNumber="54">Geosesarma katibas</emphasis>
|
||
</taxonomicName>
|
||
(Lanjak-Lantimau, central Borneo) can be separated from
|
||
<taxonomicName id="54C64D08E141BD7C57A1FAE5DB1D8CF8" ID-CoL="3FS8P" box="[919,1076,944,968]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C57A1FAE5DB1D8CF8" box="[919,1076,944,968]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5008FAFADBB28CF8" bold="true" box="[1086,1179,943,968]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5008FAFADBB28CF8" box="[1086,1179,943,968]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having an external orbital tooth that is more acutely triangular in shape (cf. Ng 1995b: fig. 9, 11A), the anterior carapace regions are more prominently granular (cf. Ng 1995b: fig. 9), the ambulatory meri and propodi are longer (cf. Ng 1995b: fig. 9, 11F), the merus of the third maxilliped is shorter than the ischium (cf. Ng 1995b: fig. 11B), and the G1 has no subdistal hump, with the distal pectinated part subcylindrical and tapering (cf. Ng 1995b: fig. 12A–E). These differences are also applicable for
|
||
<taxonomicName id="54C64D08E141BD7C5601FD31DDE58B4C" ID-CoL="6KBJ5" box="[567,716,1124,1148]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5601FD31DDE58B4C" box="[567,716,1124,1148]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C56EFFD31DC108B4D" bold="true" box="[729,825,1124,1149]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C56EFFD31DC108B4D" box="[729,825,1124,1149]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, except that their ambulatory legs do not differ in proportions (
|
||
<figureCitation id="0BFD2A0EE141BD7C551CFDD2DEAF8B90" box="[298,390,1159,1184]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Figs. 11</figureCitation>
|
||
A, 13C; Ng 1995b: fig. 9, 11F), and in
|
||
<taxonomicName id="54C64D08E141BD7C5765FDDDDCEC8B90" ID-CoL="8YX9B" box="[851,965,1160,1184]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="A1B2EA99E141BD7C5765FDDDDCEC8B90" box="[851,965,1160,1184]" italics="true" pageId="17" pageNumber="54">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
, the male abdomen, notably somite 6, is proportionately narrower (cf. Ng 1995b: fig. 11G versus
|
||
<figureCitation id="0BFD2A0EE141BD7C5739FDF9DC408BF4" box="[783,873,1196,1221]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="17" pageNumber="54">Figs. 10</figureCitation>
|
||
D, 12B). The live colour of
|
||
<taxonomicName id="54C64D08E141BD7C50ABFDF9DA6C8BF4" ID-CoL="3FS99" box="[1181,1349,1196,1220]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E141BD7C50ABFDF9DA6C8BF4" box="[1181,1349,1196,1220]" italics="true" pageId="17" pageNumber="54">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
is quite different from that of
|
||
<taxonomicName id="54C64D08E141BD7C55BFFD85DD0B8BD8" ID-CoL="3FS8P" box="[393,546,1232,1256]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C55BFFD85DD0B8BD8" box="[393,546,1232,1256]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C561FFD9ADDAB8BD8" bold="true" box="[553,642,1231,1256]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C561FFD9ADDAB8BD8" box="[553,642,1231,1256]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E141BD7C5681FD85DC6F8BD8" ID-CoL="6KBJ5" box="[695,838,1232,1256]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5681FD85DC6F8BD8" box="[695,838,1232,1256]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C577AFD9ADC8C8BD8" bold="true" box="[844,933,1231,1256]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C577AFD9ADC8C8BD8" box="[844,933,1231,1256]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, being a relatively uniform reddish-brown to red, including the chelae (unpublished data), whereas
|
||
<taxonomicName id="54C64D08E141BD7C56CAFDA1DC448A3C" ID-CoL="8YX9B" box="[764,877,1268,1292]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="A1B2EA99E141BD7C56CAFDA1DC448A3C" box="[764,877,1268,1292]" italics="true" pageId="17" pageNumber="54">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
has a reddish-brown carapace, with the palms of the chela purple and the fingers orange (cf. Ng 1995b: fig. 10). Live
|
||
<taxonomicName id="54C64D08E141BD7C57A4FC4DDCF38A00" ID-CoL="3FS8V" box="[914,986,1304,1328]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="A1B2EA99E141BD7C57A4FC4DDCF38A00" box="[914,986,1304,1328]" italics="true" pageId="17" pageNumber="54">G. bau</emphasis>
|
||
</taxonomicName>
|
||
have an orangish-red carapace, with the chelae and legs brown (unpublished data).
|
||
</paragraph>
|
||
<paragraph id="9379368BE141BD7F54F1FC35DBA88D50" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,1581]" lastPageId="18" lastPageNumber="55" pageId="17" pageNumber="54">
|
||
<taxonomicName id="54C64D08E141BD7C54F1FC35DEFC8A48" ID-CoL="3FS8P" box="[199,469,1376,1400]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C54F1FC35DEFC8A48" box="[199,469,1376,1400]" italics="true" pageId="17" pageNumber="54">Geosesarma ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C55EAFC0ADD1C8A48" bold="true" box="[476,565,1375,1400]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C55EAFC0ADD1C8A48" box="[476,565,1375,1400]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E141BD7C565DFC35DDD38A48" ID-CoL="6KBJ5" box="[619,762,1376,1400]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C565DFC35DDD38A48" box="[619,762,1376,1400]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5737FC0ADC728A48" bold="true" box="[769,859,1375,1400]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5737FC0ADC728A48" box="[769,859,1375,1400]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also resemble
|
||
<taxonomicName id="54C64D08E141BD7C5034FC35DA418A48" ID-CoL="3FS9Q" authority="Ng & Tan, 1995" authorityName="Ng & Tan" authorityYear="1995" box="[1026,1384,1375,1400]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E141BD7C5034FC35DB808A48" box="[1026,1193,1376,1400]" italics="true" pageId="17" pageNumber="54">G. notophorum</emphasis>
|
||
Ng &
|
||
<bibRefCitation id="F7574B7AE141BD7C50C0FC35DA418A48" author="Tan" box="[1270,1384,1375,1400]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="A1B2EA99E141BD7C54A1FCD1DFFC8AAC" box="[151,213,1412,1436]" italics="true" pageId="17" pageNumber="54">G. r a j</emphasis>
|
||
<bibRefCitation id="F7574B7AE141BD7C54D4FCD1DEF28AAC" author="Schubart" box="[226,475,1412,1437]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng, 2014</bibRefCitation>
|
||
, from
|
||
<collectingCountry id="EBD1761BE141BD7C5611FCD1DD4E8AAD" box="[551,615,1412,1437]" name="Iceland" pageId="17" pageNumber="54">Pulau</collectingCountry>
|
||
Lingga and
|
||
<collectingCountry id="EBD1761BE141BD7C56CDFCD1DC128AAD" box="[763,827,1412,1437]" name="Iceland" pageId="17" pageNumber="54">Pulau</collectingCountry>
|
||
Bintan in the Riau Islands off Sumatra, respectively. Both species also lack a flagellum on the exopod of the third maxilliped.
|
||
<taxonomicName id="54C64D08E141BD7C5039FCFDDBE08A8F" ID-CoL="3FS9X" box="[1039,1225,1448,1471]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="raj">
|
||
<emphasis id="A1B2EA99E141BD7C5039FCFDDBE08A8F" box="[1039,1225,1448,1471]" italics="true" pageId="17" pageNumber="54">Geosesarma raj</emphasis>
|
||
</taxonomicName>
|
||
, however, can be separated from
|
||
<taxonomicName id="54C64D08E141BD7C5572FC99DEF78AD4" ID-CoL="3FS8P" box="[324,478,1484,1508]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5572FC99DEF78AD4" box="[324,478,1484,1508]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C55D2FC99DD178AD5" bold="true" box="[484,574,1484,1509]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C55D2FC99DD178AD5" box="[484,574,1484,1509]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having frontal lobes that are more truncated in shape from the dorsal view (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C54F3FCBADE9E8938" author="Schubart" box="[197,439,1519,1544]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 1B, D), the external orbital tooth is more acutely triangular and anteriorly directed (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C54F3FF41DE9C891C" author="Schubart" box="[197,437,1556,1581]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 1B, D), the dorsal margin of the dactylus of the chela has only 8 or 9 tubercles (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C54A1FF62DEA48960" author="Schubart" box="[151,397,1591,1616]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 2D), the ambulatory merus is relatively longer and more slender (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C50DBFF62DFFD8944" author="Schubart" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 1B, 3D), and the distal pectinated part of the G1 is subcylindrical and tapering (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C509AFF09DAB18944" author="Schubart" box="[1196,1432,1628,1653]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 3J–M). Its live colour also differs slightly, with the carapace being greyish and the chelae are uniformly reddish-orange (
|
||
<bibRefCitation id="F7574B7AE141BD7C557BFFF1DDAC898C" author="Schubart" box="[333,645,1700,1725]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014: fig. 4</bibRefCitation>
|
||
).
|
||
<taxonomicName id="54C64D08E141BD7C56AFFFF1DCB5898C" ID-CoL="6KBJ5" box="[665,924,1700,1724]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C56AFFFF1DCB5898C" box="[665,924,1700,1724]" italics="true" pageId="17" pageNumber="54">Geosesarma pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5795FFF1DCD4898D" bold="true" box="[931,1021,1700,1725]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5795FFF1DCD4898D" box="[931,1021,1700,1725]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<emphasis id="A1B2EA99E141BD7C50B9FFF1DBE2898C" box="[1167,1227,1700,1724]" italics="true" pageId="17" pageNumber="54">G. r a j</emphasis>
|
||
in the structure of the external orbital teeth and ambulatory leg proportions (
|
||
<figureCitation id="0BFD2A0EE141BD7C5717FF92DC5989D0" box="[801,880,1735,1760]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Fig. 11</figureCitation>
|
||
A, B, 13C) but the carapace of
|
||
<emphasis id="A1B2EA99E141BD7C50E7FF9DDA2489EF" box="[1233,1293,1736,1760]" italics="true" pageId="17" pageNumber="54">G. r a j</emphasis>
|
||
is relatively broader (width to length usually 1.07–1.10) with the postorbital and postfrontal cristae not as strong (cf.
|
||
<bibRefCitation id="F7574B7AE141BD7C510FFFB9DE368818" author="Schubart" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014</bibRefCitation>
|
||
: fig. 1B, D); male abdominal somite 6 is proportionately less broad (
|
||
<bibRefCitation id="F7574B7AE141BD7C5077FE5ADABD8818" author="Schubart" box="[1089,1428,1807,1833]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014: fig. 3F</bibRefCitation>
|
||
) (much wider in
|
||
<taxonomicName id="54C64D08E141BD7C557AFE61DEF5887C" ID-CoL="6KBJ5" box="[332,476,1844,1868]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C557AFE61DEF5887C" box="[332,476,1844,1868]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C55D2FE61DD17887D" bold="true" box="[484,574,1844,1869]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C55D2FE61DD17887D" box="[484,574,1844,1869]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE141BD7C567BFE61DD81887C" box="[589,680,1844,1869]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="17" pageNumber="54">Figs. 11</figureCitation>
|
||
D, 12, 13B); and the subterminal hump of the G1 is gently sloping (
|
||
<bibRefCitation id="F7574B7AE141BD7C54A8FE02DD2D8840" author="Schubart" box="[158,516,1879,1905]" pageId="17" pageNumber="54" refString="Schubart, C. D. & Ng, P. K. L. (2014) Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia. Raffles Bulletin of Zoology, 62, 615 - 619." type="journal article" year="2014">Schubart & Ng 2014: fig. 3J–M</bibRefCitation>
|
||
) (at right angles
|
||
<taxonomicName id="54C64D08E141BD7C56F7FE0DDC668840" ID-CoL="6KBJ5" box="[705,847,1880,1904]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C56F7FE0DDC668840" box="[705,847,1880,1904]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5760FE02DC868840" bold="true" box="[854,943,1879,1904]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5760FE02DC868840" box="[854,943,1879,1904]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE141BD7C578BFE02DB238840" box="[957,1034,1879,1904]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="17" pageNumber="54">Fig. 13</figureCitation>
|
||
D–H, J, K, M). The carapace shape, proportions, structures of the postfrontal and postorbital cristae, as well as the external orbital tooth of
|
||
<taxonomicName id="54C64D08E141BD7C51B2FE29DE098888" ID-CoL="3FS9Q" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E141BD7C51B2FE29DE098888" italics="true" pageId="17" pageNumber="54">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
(cf. Ng &
|
||
<bibRefCitation id="F7574B7AE141BD7C55AFFEF4DD2C8889" author="Tan" box="[409,517,1952,1977]" pageId="17" pageNumber="54" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: pl. 1, fig. 1A) are similar to those of
|
||
<taxonomicName id="54C64D08E141BD7C5791FEF4DB688888" ID-CoL="3FS8P" box="[935,1089,1952,1976]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5791FEF4DB688888" box="[935,1089,1952,1976]" italics="true" pageId="17" pageNumber="54">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C5071FEF5DB888889" bold="true" box="[1095,1185,1952,1977]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C5071FEF5DB888889" box="[1095,1185,1952,1977]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE141BD7C5086FEF5DBD88888" box="[1200,1265,1952,1977]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="17" pageNumber="54">Fig. 8</figureCitation>
|
||
A, B) (width to length ratios 1.05-1.10 and 1.06–1.08, respectively). This is in contrast to the carapace of
|
||
<taxonomicName id="54C64D08E141BD7C5096FE91DA1988EC" ID-CoL="6KBJ5" box="[1184,1328,1988,2012]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E141BD7C5096FE91DA1988EC" box="[1184,1328,1988,2012]" italics="true" pageId="17" pageNumber="54">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E141BD7C510FFE91DAB588ED" bold="true" box="[1337,1436,1988,2013]" pageId="17" pageNumber="54">
|
||
<taxonomicNameLabel id="BA8157E2E141BD7C510FFE91DABF88ED" box="[1337,1430,1988,2013]" pageId="17" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
which is distinctly more quadrate (width to length ratio 1.01–1.02), the postfrontal and postorbital cristae are much sharper and more prominent, and the external orbital tooth is broader and directed anteriorly (
|
||
<figureCitation id="0BFD2A0EE142BD7F50F8F9E9DA368FE4" box="[1230,1311,188,213]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="18" pageNumber="55">Fig. 11</figureCitation>
|
||
A, B). The ambulatory legs are relatively short in
|
||
<taxonomicName id="54C64D08E142BD7F5666F9B5DDC28FC8" ID-CoL="3FS8P" box="[592,747,224,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5666F9B5DDC28FC8" box="[592,747,224,248]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F56C5F98ADC678FC8" bold="true" box="[755,846,223,248]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F56C5F98ADC678FC8" box="[755,846,223,248]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE142BD7F5769F98ADC858FC8" box="[863,940,223,248]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="18" pageNumber="55">Figs. 8</figureCitation>
|
||
A, 9C), but are proportionately longer in
|
||
<taxonomicName id="54C64D08E142BD7F51B2F9B5DE098E2C" ID-CoL="3FS9Q" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F51B2F9B5DE098E2C" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
(cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F55ABF850DD238E2D" author="Tan" box="[413,522,260,285]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: pl. 1, fig. 1F) and
|
||
<taxonomicName id="54C64D08E142BD7F56D5F851DC5A8E2C" ID-CoL="6KBJ5" box="[739,883,260,284]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F56D5F851DC5A8E2C" box="[739,883,260,284]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F574CF851DCFC8E2D" bold="true" box="[890,981,260,285]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F574CF851DCFC8E2D" box="[890,981,260,285]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE142BD7F57D2F851DB178E2C" box="[996,1086,260,285]" captionStart="FIGURE 11" captionStartId="13.[151,250,1936,1958]" captionTargetBox="[168,1422,181,1924]" captionTargetId="figure@13.[168,1423,181,1925]" captionTargetPageId="13" captionText="FIGURE 11. Geosesarma pontianak sp. nov., holotype male (13.4 × 13.2 mm) (MZB Cru 4392), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of left chela." httpUri="https://zenodo.org/record/290086/files/figure.png" pageId="18" pageNumber="55">Figs. 11</figureCitation>
|
||
A, 13C). The male telson of
|
||
<taxonomicName id="54C64D08E142BD7F51B2F851DE098E70" ID-CoL="3FS9Q" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F51B2F851DE098E70" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
is distinctly more triangular (cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F56D1F87DDC7F8E70" author="Tan" box="[743,854,295,320]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: fig. 1C) than that of
|
||
<taxonomicName id="54C64D08E142BD7F5067F87DDBC48E70" ID-CoL="3FS8P" box="[1105,1261,296,320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5067F87DDBC48E70" box="[1105,1261,296,320]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F50C0F872DA7B8E70" bold="true" box="[1270,1362,295,320]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F50C0F872DA7B8E70" box="[1270,1362,295,320]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(or
|
||
<taxonomicName id="54C64D08E142BD7F51B2F87DDE2E8E54" ID-CoL="6KBJ5" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F51B2F87DDE2E8E54" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5524F819DE598E55" bold="true" box="[274,368,332,357]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5524F819DE598E55" box="[274,368,332,357]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(relatively broader and more rounded in these two species,
|
||
<figureCitation id="0BFD2A0EE142BD7F500CF819DBA38E54" box="[1082,1162,332,357]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="18" pageNumber="55">Figs. 8</figureCitation>
|
||
D, 9B, 11D, 13B). The proportions of the male abdominal somite 6 of
|
||
<taxonomicName id="54C64D08E142BD7F5694F825DC638EB8" ID-CoL="3FS9Q" box="[674,842,368,392]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F5694F825DC638EB8" box="[674,842,368,392]" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
(cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F57F4F825DB078EB8" author="Tan" box="[962,1070,367,392]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: fig. 1C), however, are similar to that of
|
||
<taxonomicName id="54C64D08E142BD7F54DCF8C1DEAC8E9C" ID-CoL="3FS8P" box="[234,389,404,428]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F54DCF8C1DEAC8E9C" box="[234,389,404,428]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F55BBF8C1DEC08E9D" bold="true" box="[397,489,404,429]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F55BBF8C1DEC08E9D" box="[397,489,404,429]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE142BD7F55CCF8C1DD168E9C" box="[506,575,404,429]" captionStart="FIGURE 9" captionStartId="11.[151,250,1514,1536]" captionTargetBox="[167,1418,200,1477]" captionTargetId="figure@11.[150,1436,193,1493]" captionTargetPageId="11" captionText="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." httpUri="https://zenodo.org/record/290084/files/figure.png" pageId="18" pageNumber="55">Fig. 9</figureCitation>
|
||
B), with both being proportionately less broad than that of
|
||
<taxonomicName id="54C64D08E142BD7F50D7F8C1DA5B8E9C" ID-CoL="6KBJ5" box="[1249,1394,404,428]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F50D7F8C1DA5B8E9C" box="[1249,1394,404,428]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F514CF8C1DFE18EE0" bold="true" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F514CF8C1DFE18EE0" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="0BFD2A0EE142BD7F54E1F8E2DE008EE0" box="[215,297,439,464]" captionStart="FIGURE 13" captionStartId="15.[151,250,1858,1880]" captionTargetBox="[161,1422,463,1843]" captionTargetId="figure@15.[151,1436,459,1849]" captionTargetPageId="15" captionText="FIGURE 13. Geosesarma pontianak sp. nov., Anjungan, Kalimantan Barat, Borneo. A – I, holotype male (13.4 × 13.2 mm) (MZB Cru 4392); J, K, paratype male (13.2 × 13.0 mm) (ZRC 2015.0454 a); L, paratype male (11.4 × 10.0 mm) (ZRC 2015.0454 b); M, paratype male (11.1 × 10.8 mm) (ZRC 2015.0454 c). A, left third maxilliped; B, male abdomen; C, left second ambulatory leg; D, J, L, left G 1 (ventral view); E, left G 1 (dorsal view); F, K, M, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, left G 2. All structures (except C) denuded. Scales: A – C, D, E, I, J, L = 1.0 mm, F – H, K, M = 0.5 mm." httpUri="https://zenodo.org/record/290088/files/figure.png" pageId="18" pageNumber="55">Fig. 13</figureCitation>
|
||
B). Most significantly, the G1 of
|
||
<taxonomicName id="54C64D08E142BD7F56AAF8EDDC6A8EE0" ID-CoL="3FS9Q" box="[668,835,440,464]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F56AAF8EDDC6A8EE0" box="[668,835,440,464]" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
does not have a distinct subdistal hump, being gently sloping and the distal pectinated part is more tapering (cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F575BF888DCF08EC5" author="Tan" box="[877,985,476,501]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: fig. 1H–J), in sharp contrast to those of
|
||
<taxonomicName id="54C64D08E142BD7F54A1FB55DE198D28" ID-CoL="3FS8P" box="[151,304,512,536]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F54A1FB55DE198D28" box="[151,304,512,536]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5501F8AADEB88D28" bold="true" box="[311,401,511,536]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5501F8AADEB88D28" box="[311,401,511,536]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E142BD7F55F1FB55DD7F8D28" ID-CoL="6KBJ5" box="[455,598,512,536]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F55F1FB55DD7F8D28" box="[455,598,512,536]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F566BF8AADD978D28" bold="true" box="[605,702,511,536]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F566BF8AADD918D28" box="[605,696,511,536]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
which have a distinct hump, being usually distinctly right-angled in the latter species (
|
||
<figureCitation id="0BFD2A0EE142BD7F55BDFB71DEF38D0C" box="[395,474,548,573]" captionStart="FIGURE 9" captionStartId="11.[151,250,1514,1536]" captionTargetBox="[167,1418,200,1477]" captionTargetId="figure@11.[150,1436,193,1493]" captionTargetPageId="11" captionText="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." httpUri="https://zenodo.org/record/290084/files/figure.png" pageId="18" pageNumber="55">Figs. 9</figureCitation>
|
||
D–H, 13D–H, J, K, M). The carapace
|
||
<taxonomicName id="54C64D08E142BD7F57ABFB71DB618D0C" ID-CoL="3FS9Q" box="[925,1096,548,572]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F57ABFB71DB618D0C" box="[925,1096,548,572]" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
is also evenly pale brown to purplish-brown in life, with the posterior half usually darker (Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F57A6FB1DDCD58D50" author="Tan" box="[912,1020,583,608]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
: 394, pl. 1).
|
||
</paragraph>
|
||
<paragraph id="9379368BE142BD7F54F1FB38DF938CDC" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="55">
|
||
In live coloration,
|
||
<taxonomicName id="54C64D08E142BD7F55ABFB39DD108DB4" ID-CoL="3FS8P" box="[413,569,620,644]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F55ABFB39DD108DB4" box="[413,569,620,644]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5675FB39DDB68DB5" bold="true" box="[579,671,620,645]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5675FB39DDB68DB5" box="[579,671,620,645]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E142BD7F56EDFB39DC458DB4" ID-CoL="6KBJ5" box="[731,876,620,644]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F56EDFB39DC458DB4" box="[731,876,620,644]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5743FB39DCFB8DB5" bold="true" box="[885,978,620,645]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5743FB39DCFB8DB5" box="[885,978,620,645]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resemble
|
||
<taxonomicName id="54C64D08E142BD7F507CFB39DAB58DB5" ID-CoL="3FS8W" authority="Ng & Davie, 1995" authorityName="Ng & Davie" authorityYear="1995" box="[1098,1436,620,645]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="A1B2EA99E142BD7F507CFB39DB958DB4" box="[1098,1212,620,644]" italics="true" pageId="18" pageNumber="55">G. bicolor</emphasis>
|
||
Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F5126FB39DAB58DB5" author="Davie" box="[1296,1436,620,645]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
(western Java), which was described as bright purple on the anterior third of the carapace and ambulatory legs, with the posterior parts of the carapace bluish-grey; the chelae red and eyes bright yellow (Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F509CFBE1DA188DFD" author="Davie" box="[1194,1329,692,717]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie 1995</bibRefCitation>
|
||
: 32). The ambulatory legs of
|
||
<emphasis id="A1B2EA99E142BD7F554FFB8DDECE8DC0" box="[377,487,728,752]" italics="true" pageId="18" pageNumber="55">G. b i c o l o r</emphasis>
|
||
, however, are distinctively shorter than either species (Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F5081FB82DA698DC0" author="Davie" box="[1207,1344,727,752]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie 1995</bibRefCitation>
|
||
: fig. 1). While the carapace of
|
||
<emphasis id="A1B2EA99E142BD7F5596FBA9DD268C24" box="[416,527,764,788]" italics="true" pageId="18" pageNumber="55">G. b i co l o r</emphasis>
|
||
superficially resembles
|
||
<taxonomicName id="54C64D08E142BD7F5718FBA9DCE28C24" ID-CoL="3FS8P" box="[814,971,764,788]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5718FBA9DCE28C24" box="[814,971,764,788]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F57E3FBA9DB1B8C25" bold="true" box="[981,1074,764,789]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F57E3FBA9DB1B8C25" box="[981,1074,764,789]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="54C64D08E142BD7F5059FBA9DA288C24" ID-CoL="6KBJ5" box="[1135,1281,764,788]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5059FBA9DA288C24" box="[1135,1281,764,788]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F513AFBA9DA418C25" bold="true" box="[1292,1384,764,789]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F513AFBA9DA418C25" box="[1292,1384,764,789]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the exopod of the third maxilliped of
|
||
<emphasis id="A1B2EA99E142BD7F562EFA75DDAC8C08" box="[536,645,800,824]" italics="true" pageId="18" pageNumber="55">G. b i co l o r</emphasis>
|
||
has an elongated flagellum (cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F5076FA4ADBE18C08" author="Davie" box="[1088,1224,799,824]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie 1995</bibRefCitation>
|
||
: fig. 2C), the male abdomen is proportionately broader (cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F569FFA11DC198C6D" author="Davie" box="[681,816,836,861]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie 1995</bibRefCitation>
|
||
: fig. 2F; Ng
|
||
<emphasis id="A1B2EA99E142BD7F57F4FA10DCD58C6C" box="[962,1020,836,860]" italics="true" pageId="18" pageNumber="55">et al.</emphasis>
|
||
2015: fig. 2J), and the G1 is slender with an elongated pectinated distal part (cf. Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F56F2FA32DC608CB0" author="Davie" box="[708,841,871,896]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Davie, P. J. F. (1995) The terrestrial sesarmine crabs of the genera Metasesarma and Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) of Ujung Kulon, West Jawa, Indonesia. Tropical Biodiversity, 3, 29 - 43." type="journal article" year="1995">Davie 1995</bibRefCitation>
|
||
: fig. 2G, I–M; Ng
|
||
<emphasis id="A1B2EA99E142BD7F5025FA3CDB628CB0" box="[1043,1099,872,896]" italics="true" pageId="18" pageNumber="55">et al.</emphasis>
|
||
2015: fig. 2F–I).
|
||
<taxonomicName id="54C64D08E142BD7F5126FA3DDD498C95" ID-CoL="3FS95" authority="Ng, Schubart & Lukhaup, 2015" authorityName="Ng, Schubart & Lukhaup" authorityYear="2015" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="dennerle">
|
||
<emphasis id="A1B2EA99E142BD7F5126FA3DDFD08C94" italics="true" pageId="18" pageNumber="55">Geosesarma dennerle</emphasis>
|
||
Ng,
|
||
<bibRefCitation id="F7574B7AE142BD7F5506FAD9DD498C95" author="Schubart" box="[304,608,908,933]" pageId="18" pageNumber="55" refString="Ng, P. K. L., Schubart, C. D. & Lukhaup, C. (2015) New species of " vampire crabs " (Geosesarma De Man, 1892) from central Java, Indonesia, and the identity of Sesarma (Geosesarma) nodulifera De Man, 1892 (Crustacea, Brachyura, Thoracotremata, Sesarmidae). Raffles Bulletin of Zoology, 63, 3 - 13." type="journal article" year="2015">Schubart & Lukhaup, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
(central Java) has a bi-coloured carapace, but in this species, the carapace colour is a much brighter purple and the chelae are also bright purple, not red or orange (cf. Ng
|
||
<emphasis id="A1B2EA99E142BD7F508DFAE4DBDA8CF8" box="[1211,1267,944,968]" italics="true" pageId="18" pageNumber="55">et al.</emphasis>
|
||
2015: fig. 6A– C).
|
||
</paragraph>
|
||
<paragraph id="9379368BE142BD7F54F1FAA2DFC68A64" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="55">
|
||
The aquarium trade has been exporting many species of
|
||
<taxonomicName id="54C64D08E142BD7F570DFAADDCEE8B3F" ID-CoL="7P3H8" box="[827,967,1016,1039]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A1B2EA99E142BD7F570DFAADDCEE8B3F" box="[827,967,1016,1039]" italics="true" pageId="18" pageNumber="55">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
to Europe, but the identities of these have always been dubious, partly because of their unknown provenance (see
|
||
<bibRefCitation id="F7574B7AE142BD7F57EDFD49DA718B04" author="Rademacher" box="[987,1368,1052,1077]" pageId="18" pageNumber="55" refString="Rademacher, M. & Mengedoht, O. (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, 95 pp." type="book" year="2011">Rademacher & Mengedoht 2011</bibRefCitation>
|
||
). One species worth noting is what has been called “
|
||
<taxonomicName id="54C64D08E142BD7F56AEFD15DB5F8B68" ID-CoL="3FS9Q" authority="Mandarinkrabbe" authorityName="Mandarinkrabbe" box="[664,1142,1087,1112]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F56AEFD15DC9A8B68" box="[664,947,1088,1112]" italics="true" pageId="18" pageNumber="55">Geosesarma notophorum</emphasis>
|
||
Mandarinkrabbe
|
||
</taxonomicName>
|
||
” by the
|
||
<collectingCountry id="EBD1761BE142BD7F50E7FD15DA028B68" box="[1233,1323,1088,1112]" name="Germany" pageId="18" pageNumber="55">German</collectingCountry>
|
||
aquraium trade (
|
||
<bibRefCitation id="F7574B7AE142BD7F54EAFD31DD518B4C" author="Rademacher" box="[220,632,1124,1149]" pageId="18" pageNumber="55" refString="Rademacher, M. & Mengedoht, O. (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, 95 pp." type="book" year="2011">Rademacher & Mengedoht 2011: 42</bibRefCitation>
|
||
). These specimens, which have been imported into
|
||
<collectingCountry id="EBD1761BE142BD7F5080FD31DA378B4C" box="[1206,1310,1124,1148]" name="Germany" pageId="18" pageNumber="55">Germany</collectingCountry>
|
||
since 2006 (possibly earlier), bear a close resemblance in their colour pattern to what is here described as
|
||
<taxonomicName id="54C64D08E142BD7F5098FDDDDA158B90" ID-CoL="6KBJ5" box="[1198,1340,1160,1184]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5098FDDDDA158B90" box="[1198,1340,1160,1184]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5175FDD2DAB58B90" bold="true" box="[1347,1436,1159,1184]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5175FDD2DAB58B90" box="[1347,1436,1159,1184]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in their colour pattern. Although they were reportedly collected from the
|
||
<typeStatus id="4C7D8829E142BD7F57E7FDF8DB288BF5" box="[977,1025,1197,1221]" pageId="18" pageNumber="55">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="EBD1761BE142BD7F50B4FDF9DBEB8BF5" box="[1154,1218,1196,1221]" name="Iceland" pageId="18" pageNumber="55">Pulau</collectingCountry>
|
||
Lingga, this seems unlikely as this location is not easily accessible.
|
||
<taxonomicName id="54C64D08E142BD7F5687FD9ADC0D8BD8" ID-CoL="6KBJ5" box="[689,804,1231,1256]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, on the other hand has several major aquarium dealers in the city, and they regularly export freshwater fish from western Borneo. It seems more likely that the collectors in
|
||
<taxonomicName id="54C64D08E142BD7F54A1FC42DE238A00" ID-CoL="6KBJ5" box="[151,266,1303,1328]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
|
||
, who obtained the present specimens from Anjungan for the trade, had and continue to export material to Europe.
|
||
</paragraph>
|
||
<paragraph id="9379368BE142BD7F54F1FC35DB84891C" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="55">
|
||
<taxonomicName id="54C64D08E142BD7F54F1FC35DECF8A48" ID-CoL="3FS9Q" box="[199,486,1376,1400]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F54F1FC35DECF8A48" box="[199,486,1376,1400]" italics="true" pageId="18" pageNumber="55">Geosesarma notophorum</emphasis>
|
||
</taxonomicName>
|
||
is known to possess a novel method of caring for its newly hatched juveniles by carrying them on its carapace (Ng &
|
||
<bibRefCitation id="F7574B7AE142BD7F5600FCD0DD8A8AAD" author="Tan" box="[566,675,1412,1437]" pageId="18" pageNumber="55" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. http: // dx. doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Tan 1995</bibRefCitation>
|
||
). This behaviour is now known for several species that also practice direct development. The eggs of
|
||
<taxonomicName id="54C64D08E142BD7F5631FCFDDD888AF0" ID-CoL="3FS8P" box="[519,673,1448,1472]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F5631FCFDDD888AF0" box="[519,673,1448,1472]" italics="true" pageId="18" pageNumber="55">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F569EFCF2DC2B8AF0" bold="true" box="[680,770,1447,1472]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F569EFCF2DC2B8AF0" box="[680,770,1447,1472]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are large, and indicate that they hatch directly into juvenile crabs, and possibly behave the same way as
|
||
<taxonomicName id="54C64D08E142BD7F56B2FC99DC028AD4" ID-CoL="3FS9Q" box="[644,811,1484,1508]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F56B2FC99DC028AD4" box="[644,811,1484,1508]" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
. If the material exported into
|
||
<collectingCountry id="EBD1761BE142BD7F5041FC99DBF68AD4" box="[1143,1247,1484,1508]" name="Germany" pageId="18" pageNumber="55">Germany</collectingCountry>
|
||
as “
|
||
<taxonomicName id="54C64D08E142BD7F5126FC99DECC8938" ID-CoL="3FS9Q" authority="Mandarinkrabbe" authorityName="Mandarinkrabbe" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F5126FC99DE098938" italics="true" pageId="18" pageNumber="55">Geosesarma notophorum</emphasis>
|
||
Mandarinkrabbe
|
||
</taxonomicName>
|
||
” is indeed what is here described as
|
||
<taxonomicName id="54C64D08E142BD7F57B8FCA5DB378938" ID-CoL="6KBJ5" box="[910,1054,1520,1544]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E142BD7F57B8FCA5DB378938" box="[910,1054,1520,1544]" italics="true" pageId="18" pageNumber="55">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E142BD7F5011FCBADBAB8938" bold="true" box="[1063,1154,1519,1544]" pageId="18" pageNumber="55">
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F5011FCBADBAB8938" box="[1063,1154,1519,1544]" pageId="18" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above), then it also possesses the same brood care method as
|
||
<taxonomicName id="54C64D08E142BD7F565CFF41DC3B891C" ID-CoL="3FS9Q" box="[618,786,1556,1580]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" sensu="stricto" species="notophorum">
|
||
<emphasis id="A1B2EA99E142BD7F565CFF41DC3B891C" box="[618,786,1556,1580]" italics="true" pageId="18" pageNumber="55">G. notophorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA8157E2E142BD7F572FFF40DC7A891C" box="[793,851,1557,1580]" pageId="18" pageNumber="55" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
(O. Mengedoht, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |