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<document id="37F3746482AD99224C8799DFA5714E71" ID-DOI="10.11646/zootaxa.4048.1.2" ID-GBIF-Dataset="2f5da4b3-c0ee-4b70-b661-0e2ef9bacfa5" ID-ISSN="1175-5326" ID-Zenodo-Dep="290075" ID-ZooBank="97BB73FF-9D62-4A15-B09C-0F570E26E738" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461341944160" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2015" docId="1B6F879DE156BD6454A1FF16DC668856" docLanguage="en" docName="zt04048p056.pdf" docOrigin="Zootaxa 4048 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma ambawang Ng, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="46" masterDocId="E756FFE5E150BD6D5436F955DF298F30" masterDocTitle="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species" masterLastPageNumber="56" masterPageNumber="37" pageNumber="43" updateTime="1698620270231" updateUser="plazi">
<mods:mods id="A6B66793A4FA7237530B901AC00D4D1C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="615ED5D990292DB1EC75FD0635A17ABF">Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="2FDDA834281FBBBC95B975A2077B7813" type="personal">
<mods:role id="37A12C4CE3D993A98992C8B3EE69497F">
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<mods:namePart id="F821FEFB784FCF64BA3269AFC4EF071F">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="C645790F68E3082600165FF5F7C603C4">text</mods:typeOfResource>
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<mods:title id="32BA4C745A0AD2CB67704C31D7F09E97">Zootaxa</mods:title>
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<mods:part id="47117F4D91F875F654C211AF9108638A">
<mods:date id="7994AE78EA60C6BF4D59A0C267A5807F">2015</mods:date>
<mods:detail id="A395922B69A70E6C85EB9DAF770E0E51" type="volume">
<mods:number id="F1D67EE0FD37825676CDE96AF4F96C87">4048</mods:number>
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<mods:classification id="DB86D28784353F5670435072C2E4F13A">journal article</mods:classification>
<mods:identifier id="4AAB19B221C8847496CDEED512EFF406" type="DOI">10.11646/zootaxa.4048.1.2</mods:identifier>
<mods:identifier id="7900D85D027F6CC11C05B0406C48F021" type="GBIF-Dataset">2f5da4b3-c0ee-4b70-b661-0e2ef9bacfa5</mods:identifier>
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<mods:identifier id="CFB2790A868131BDE364AAC48CE9BE66" type="Zenodo-Dep">290075</mods:identifier>
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<treatment id="1B6F879DE156BD6454A1FF16DC668856" ID-DOI="http://doi.org/10.5281/zenodo.6121304" ID-GBIF-Taxon="127694317" ID-Zenodo-Dep="6121304" LSID="urn:lsid:plazi:treatment:1B6F879DE156BD6454A1FF16DC668856" httpUri="http://treatment.plazi.org/id/1B6F879DE156BD6454A1FF16DC668856" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43">
<subSubSection id="DBDC6500E156BD6B54A1FF16DE36894F" pageId="6" pageNumber="43" type="nomenclature">
<paragraph id="9379368BE156BD6B54A1FF16DD32896E" blockId="6.[151,539,1603,1663]" box="[151,539,1603,1630]" pageId="6" pageNumber="43">
<heading id="C83181E7E156BD6B54A1FF16DD32896E" bold="true" box="[151,539,1603,1630]" fontSize="11" level="1" pageId="6" pageNumber="43" reason="1">
<emphasis id="A1B2EA99E156BD6B54A1FF16DD32896E" bold="true" box="[151,539,1603,1630]" pageId="6" pageNumber="43">
<taxonomicName id="54C64D08E156BD6B54A1FF16DE9F896D" ID-CoL="3FS8P" box="[151,438,1603,1629]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">
<emphasis id="A1B2EA99E156BD6B54A1FF16DE9F896D" bold="true" box="[151,438,1603,1629]" italics="true" pageId="6" pageNumber="43">Geosesarma ambawang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA8157E2E156BD6B558BFF11DD32896E" box="[445,539,1604,1630]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9379368BE156BD6B54A1FF33DE36894F" blockId="6.[151,539,1603,1663]" box="[151,287,1638,1663]" pageId="6" pageNumber="43">
(
<figureCitation id="0BFD2A0EE156BD6B54A9FF33DE3E894F" box="[159,279,1638,1663]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="10.[151,250,1932,1954]" captionStartId-1="11.[151,250,1514,1536]" captionStartId-2="12.[151,250,1518,1540]" captionTargetBox-0="[167,1422,179,1913]" captionTargetBox-1="[167,1418,200,1477]" captionTargetBox-2="[161,1424,395,1502]" captionTargetId-0="figure@10.[167,1422,169,1913]" captionTargetId-1="figure@11.[150,1436,193,1493]" captionTargetId-2="figure@12.[161,1426,395,1502]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." captionText-1="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." captionText-2="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri-0="https://zenodo.org/record/290083/files/figure.png" httpUri-1="https://zenodo.org/record/290084/files/figure.png" httpUri-2="https://zenodo.org/record/290085/files/figure.png" pageId="6" pageNumber="43">Figs. 8–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DBDC6500E156BD6B54A1FFFBDB518803" pageId="6" pageNumber="43" type="materials_examined">
<paragraph id="9379368BE156BD6B54A1FFFBDB518803" blockId="6.[151,1437,1710,2013]" pageId="6" pageNumber="43">
<emphasis id="A1B2EA99E156BD6B54A1FFFBDE5389F7" bold="true" box="[151,378,1710,1735]" pageId="6" pageNumber="43">Material examined</emphasis>
.
<typeStatus id="4C7D8829E156BD6B55BFFFFBDEDF89F7" box="[393,502,1710,1735]" pageId="6" pageNumber="43" type="holotype">Holotype</typeStatus>
: male (11.5 ×
<quantity id="543E9B6EE156BD6B56AEFFFADDD689F7" box="[664,767,1711,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="43" unit="mm" value="10.7">10.7 mm</quantity>
) (
<collectionCode id="F5D7AE4EE156BD6B5723FFFADC7A89F7" box="[789,851,1711,1735]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="6" pageNumber="43">MZB</collectionCode>
Cru 4390), in hill forest with sandy soil,
<quantity id="543E9B6EE156BD6B511CFFFBDAB589F7" box="[1322,1436,1710,1735]" metricMagnitude="2" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="6" pageNumber="43" unit="m" value="115.0" valueMax="150.0" valueMin="80.0">80–150 m</quantity>
asl, Bukit Wangkan, northeast of Gunung
<taxonomicName id="54C64D08E156BD6B56B4FF87DC2F89DB" box="[642,774,1746,1771]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">Ambawang</taxonomicName>
, south of
<taxonomicName id="54C64D08E156BD6B574BFF87DCD989DB" box="[893,1008,1746,1771]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">Pontianak</taxonomicName>
,
<geoCoordinate id="F6F2504CE156BD6B57C9FF86DB6189DB" box="[1023,1096,1747,1771]" direction="south" orientation="latitude" pageId="6" pageNumber="43" precision="925" value="-0.36666667">0°22'S</geoCoordinate>
<geoCoordinate id="F6F2504CE156BD6B5065FF86DB9489DB" box="[1107,1213,1747,1771]" direction="east" orientation="longitude" pageId="6" pageNumber="43" precision="925" value="109.28333">109°17'E</geoCoordinate>
, Kalimantan Barat,
<collectingCountry id="EBD1761BE156BD6B54A1FFA3DE2E883F" box="[151,263,1782,1807]" name="Indonesia" pageId="6" pageNumber="43">Indonesia</collectingCountry>
, coll. Y. Kho,
<date id="E778104BE156BD6B559AFFA3DD11883F" box="[428,568,1782,1807]" pageId="6" pageNumber="43" value="2015-03">March 2015</date>
.
<typeStatus id="4C7D8829E156BD6B5672FFADDD9F8820" box="[580,694,1784,1808]" pageId="6" pageNumber="43" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="85C0FD02E156BD6B56F7FFA2DC0C883F" box="[705,805,1782,1807]" pageId="6" pageNumber="43" type="female">1 female</specimenCount>
(9.4 ×
<quantity id="543E9B6EE156BD6B574FFFA2DCFB883F" box="[889,978,1783,1807]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="6" pageNumber="43" unit="mm" value="8.7">8.7 mm</quantity>
) (
<collectionCode id="F5D7AE4EE156BD6B57DFFFA2DB0E883F" box="[1001,1063,1783,1807]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="6" pageNumber="43">MZB</collectionCode>
Cru 4391),
<specimenCount id="85C0FD02E156BD6B5084FFA2DA23883F" box="[1202,1290,1782,1807]" pageId="6" pageNumber="43" type="male">2 males</specimenCount>
(11.9 × 11.0 mm, 9.3 ×
<quantity id="543E9B6EE156BD6B5527FE4EDE428803" box="[273,363,1819,1843]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="43" unit="mm" value="8.8">8.8 mm</quantity>
),
<specimenCount id="85C0FD02E156BD6B5548FE4EDEF68803" box="[382,479,1818,1843]" pageId="6" pageNumber="43" type="female">1 female</specimenCount>
(10.5 ×
<quantity id="543E9B6EE156BD6B5608FE4EDDBC8803" box="[574,661,1819,1843]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="6" pageNumber="43" unit="mm" value="9.9">9.9 mm</quantity>
) (
<collectionCode id="F5D7AE4EE156BD6B569AFE4EDDCD8803" box="[684,740,1819,1843]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="43">ZRC</collectionCode>
2015.0453), same data as
<typeStatus id="4C7D8829E156BD6B5038FE4FDB5D8803" box="[1038,1140,1818,1843]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="DBDC6500E156BD6454F1FE6BDABA8981" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43" type="diagnosis">
<paragraph id="9379368BE156BD6454F1FE6BDDF28911" blockId="6.[151,1437,1710,2013]" lastBlockId="9.[151,1437,1328,1894]" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43">
<emphasis id="A1B2EA99E156BD6B54F1FE6BDE138867" bold="true" box="[199,314,1854,1879]" pageId="6" pageNumber="43">Diagnosis</emphasis>
. Carapace quadrate, slightly wider than long, width to length ratio 1.06–1.08, lateral margins subparallel (
<figureCitation id="0BFD2A0EE156BD6B5513FE35DE438848" box="[293,362,1888,1913]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="6" pageNumber="43">Fig. 8</figureCitation>
A, B); dorsal surface with well-defined regions, anterior regions densely covered with small rounded, flattened granules (
<figureCitation id="0BFD2A0EE156BD6B55ECFED4DD3488AA" box="[474,541,1921,1946]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="6" pageNumber="43">Fig. 8</figureCitation>
B); front deflexed, frontal lobes broad with gently convex margins; postfrontal, postorbital cristae prominent (
<figureCitation id="0BFD2A0EE156BD6B55DCFEF7DD04888B" box="[490,557,1954,1979]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="6" pageNumber="43">Fig. 8</figureCitation>
B, C); external orbital tooth triangular, directed obliquely, outer margin gently convex, tip extending beyond lateral margin (
<figureCitation id="0BFD2A0EE156BD6B56A2FE91DDFE88EC" box="[660,727,1988,2013]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="6" pageNumber="43">Fig. 8</figureCitation>
B). Merus of third maxilliped subovate, subequal in length to ischium; exopod without trace of flagellum (
<figureCitation id="0BFD2A0EE159BD6456A8FC65DDCC8A79" box="[670,741,1328,1353]" captionStart="FIGURE 9" captionStartId="11.[151,250,1514,1536]" captionTargetBox="[167,1418,200,1477]" captionTargetId="figure@11.[150,1436,193,1493]" captionTargetPageId="11" captionText="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." httpUri="https://zenodo.org/record/290084/files/figure.png" pageId="9" pageNumber="46">Fig. 9</figureCitation>
A). Outer surface of palm of adult male covered with small rounded granules; dorsal margin of dactylus with 11 or 12 tubercles (distal one smallest), each with pectinated tip (
<figureCitation id="0BFD2A0EE159BD6454A9FC2CDFCB8AA1" box="[159,226,1401,1426]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="9" pageNumber="46">Fig. 8</figureCitation>
E, F). Ambulatory legs with relatively slender merus with sharp subdistal spine on dorsal margin, surfaces slightly rugose (
<figureCitation id="0BFD2A0EE159BD645579FCC8DEB48A85" box="[335,413,1437,1462]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="9" pageNumber="46">Figs. 8</figureCitation>
A, 9C). Male abdomen triangular; somite 6 not wide, with gently convex lateral margins; telson semicircular (
<figureCitation id="0BFD2A0EE159BD645548FC94DEE58AE9" box="[382,460,1473,1498]" captionStart="FIGURE 8" captionStartId="10.[151,250,1932,1954]" captionTargetBox="[167,1422,179,1913]" captionTargetId="figure@10.[167,1422,169,1913]" captionTargetPageId="10" captionText="FIGURE 8. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, F, outer view of right chela." httpUri="https://zenodo.org/record/290083/files/figure.png" pageId="9" pageNumber="46">Figs. 8</figureCitation>
D, 9B). G1 relatively stout, subdistal part of outer margin with distinct angled humplike arch, just before distal pectinated part; pectinated distal part elongate from lateral view, subspatuliform from marginal view, with small median cleft (
<figureCitation id="0BFD2A0EE159BD64566CFF5CDDB48911" box="[602,669,1545,1570]" captionStart="FIGURE 9" captionStartId="11.[151,250,1514,1536]" captionTargetBox="[167,1418,200,1477]" captionTargetId="figure@11.[150,1436,193,1493]" captionTargetPageId="11" captionText="FIGURE 9. Geosesarma ambawang sp. nov., holotype male (11.5 × 10.7 mm) (MZB Cru 4390), Pontianak, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left second ambulatory leg; D, left G 1 (ventral view); E, left G 1 (ventral view); F, distal part of left G 1 (ventral view); G, distal part of left G 1 (dorsal view); H, distal part of left G 1 (subdorsal view); I, J, distal part of left G 1 (mesial view); K, left G 2. All structures (except C) denuded. Scales: A – E, K = 1.0 mm; F – J = 0.5 mm." httpUri="https://zenodo.org/record/290084/files/figure.png" pageId="9" pageNumber="46">Fig. 9</figureCitation>
D–J).
</paragraph>
<caption id="C7B96603E157BD6A54A1FE6BDB7F88A1" httpUri="https://zenodo.org/record/290080/files/figure.png" pageId="7" pageNumber="44" targetBox="[151,1435,193,1832]" targetPageId="7">
<paragraph id="9379368BE157BD6A54A1FE6BDB7F88A1" blockId="7.[151,1436,1854,1937]" pageId="7" pageNumber="44">
<emphasis id="A1B2EA99E157BD6A54A1FE6BDE3B8864" bold="true" box="[151,274,1854,1876]" pageId="7" pageNumber="44">FIGURE 5</emphasis>
.
<taxonomicName id="54C64D08E157BD6A5512FE6ADD998864" authority="Nobili, 1903" authorityName="Nobili" authorityYear="1903" box="[292,688,1854,1876]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="peraccae">
<emphasis id="A1B2EA99E157BD6A5512FE6ADD238864" box="[292,522,1855,1876]" italics="true" pageId="7" pageNumber="44">Geosesarma peraccae</emphasis>
(Nobili, 1903)
</taxonomicName>
, male (14.2 × 12.0 mm) (ZRC 2010.324), Nee Soon Swamp Forest, Singapore. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, outer view of left chela; F, inner view of left chela.
</paragraph>
</caption>
<caption id="C7B96603E158BD6554A1FE25DB5A88F3" httpUri="https://zenodo.org/record/290081/files/figure.png" pageId="8" pageNumber="45" targetBox="[151,1435,193,1882]" targetPageId="8">
<paragraph id="9379368BE158BD6554A1FE25DB5A88F3" blockId="8.[151,1436,1904,1987]" pageId="8" pageNumber="45">
<emphasis id="A1B2EA99E158BD6554A1FE25DE2688B6" bold="true" box="[151,271,1904,1926]" pageId="8" pageNumber="45">FIGURE 6</emphasis>
.
<taxonomicName id="54C64D08E158BD65552BFE24DDE088B6" authority="Serene, 1968" authorityName="Serene" authorityYear="1968" box="[285,713,1904,1926]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="sarawakense">
<emphasis id="A1B2EA99E158BD65552BFE24DD0E88B6" box="[285,551,1904,1926]" italics="true" pageId="8" pageNumber="45">Geosesarma sarawakense</emphasis>
(Serène, 1968)
</taxonomicName>
, holotype male (11.8 × 10.3 mm) (ZRC 1965.7.29.6), Sibu, Sarawak, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, outer view of right chela; F, inner view of right chela.
</paragraph>
</caption>
<caption id="C7B96603E159BD6454A1FDC1DFEF8A37" httpUri="https://zenodo.org/record/290082/files/figure.png" pageId="9" pageNumber="46" targetBox="[165,1423,208,1139]" targetPageId="9">
<paragraph id="9379368BE159BD6454A1FDC1DFEF8A37" blockId="9.[151,1436,1172,1287]" pageId="9" pageNumber="46">
<emphasis id="A1B2EA99E159BD6454A1FDC1DE268B9A" bold="true" box="[151,271,1172,1194]" pageId="9" pageNumber="46">FIGURE 7</emphasis>
.
<taxonomicName id="54C64D08E159BD64552BFDC0DDE08B9A" authority="Serene, 1968" authorityName="Serene" authorityYear="1968" box="[285,713,1172,1194]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="sarawakense">
<emphasis id="A1B2EA99E159BD64552BFDC0DD0E8B9A" box="[285,551,1172,1194]" italics="true" pageId="9" pageNumber="46">Geosesarma sarawakense</emphasis>
(Serène, 1968)
</taxonomicName>
, holotype male (11.8 × 10.3 mm) (ZRC 1965.7.29.6), Sibu, Sarawak, Borneo. A, left third maxilliped; B, male abdominal somites 1–6 and telson; C, left last second ambulatory leg; D, left G1 (subventral view); E, left G1 (ventral view); F, left G1 (dorsal view). All structures denuded. Scales: A–C = 1.0 mm; D–F = 0.5 mm.
</paragraph>
</caption>
<paragraph id="9379368BE159BD6454F1FF78DABA8981" blockId="9.[151,1437,1328,1894]" pageId="9" pageNumber="46">
<emphasis id="A1B2EA99E159BD6454F1FF78DE338976" bold="true" box="[199,282,1581,1606]" pageId="9" pageNumber="46">Colour</emphasis>
. Adult males have the anterior half of the dorsal surface of the carapace dark brown to reddish-brown in life, with the posterior half cream or grey; with the chelipeds bright red and the ambulatory legs reddish-brown (
<figureCitation id="0BFD2A0EE159BD6454A9FF20DFD889BD" box="[159,241,1653,1678]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="9" pageNumber="46">Fig. 10</figureCitation>
A, B). The ventral surfaces are orangish-yellow to red (
<figureCitation id="0BFD2A0EE159BD64575AFF20DCE989BD" box="[876,960,1653,1678]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="9" pageNumber="46">Fig. 10</figureCitation>
B). In smaller specimens and females, the demarcation of the two colours on the dorsal surface of the carapace is less distinct and more irregular (
<figureCitation id="0BFD2A0EE159BD645128FFCCDA468981" box="[1310,1391,1689,1714]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="9" pageNumber="46">Fig. 10</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="DBDC6500E159BD6454F1FFE8DC388871" pageId="9" pageNumber="46" type="etymology">
<paragraph id="9379368BE159BD6454F1FFE8DE3A89CA" blockId="9.[151,1437,1328,1894]" pageId="9" pageNumber="46">
<emphasis id="A1B2EA99E159BD6454F1FFE8DE6F89E6" bold="true" box="[199,326,1725,1750]" pageId="9" pageNumber="46">Etymology</emphasis>
. The species is named after the
<typeStatus id="4C7D8829E159BD645680FFEBDDCF89E6" box="[694,742,1726,1750]" pageId="9" pageNumber="46">type</typeStatus>
locality, Gunung
<taxonomicName id="54C64D08E159BD645784FFE8DB1E89E6" box="[946,1079,1725,1750]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="ambawang" status="sp. nov.">Ambawang</taxonomicName>
. The name is used as a noun in apposition.
</paragraph>
<paragraph id="9379368BE159BD6454F1FE50DC388871" blockId="9.[151,1437,1328,1894]" pageId="9" pageNumber="46">
<emphasis id="A1B2EA99E159BD6454F1FE50DE0D882E" bold="true" box="[199,292,1797,1822]" pageId="9" pageNumber="46">Ecology</emphasis>
. The species is found in the forest floor near streams. It has also been observed to climbing shrubs and small trees at night (
<figureCitation id="0BFD2A0EE159BD64554CFE7CDEE58871" box="[378,460,1833,1858]" captionStart="FIGURE 10" captionStartId="12.[151,250,1518,1540]" captionTargetBox="[161,1424,395,1502]" captionTargetId="figure@12.[161,1426,395,1502]" captionTargetPageId="12" captionText="FIGURE 10. Geosesarma ambawang sp. nov., colours in life. A, B, non-type male (12.5 × 11.8 mm); C, D, specimens climbing shrubs in situ. Specimens not preserved, photographs by Y. Kho." httpUri="https://zenodo.org/record/290085/files/figure.png" pageId="9" pageNumber="46">Fig. 10</figureCitation>
C, D) (Y. Kho, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="DBDC6500E159BD6454F1FE18DC668856" box="[199,847,1869,1894]" pageId="9" pageNumber="46" type="discussion">
<paragraph id="9379368BE159BD6454F1FE18DC668856" blockId="9.[151,1437,1328,1894]" box="[199,847,1869,1894]" pageId="9" pageNumber="46">
<emphasis id="A1B2EA99E159BD6454F1FE18DE1D8856" bold="true" box="[199,308,1869,1894]" pageId="9" pageNumber="46">Remarks</emphasis>
. See remarks for
<taxonomicName id="54C64D08E159BD6455CCFE18DDA08855" box="[506,649,1869,1893]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="pontianak" status="sp. nov.">
<emphasis id="A1B2EA99E159BD6455CCFE18DDA08855" box="[506,649,1869,1893]" italics="true" pageId="9" pageNumber="46">G. pontianak</emphasis>
</taxonomicName>
<emphasis id="A1B2EA99E159BD6456B9FE18DDC08856" bold="true" box="[655,745,1869,1894]" pageId="9" pageNumber="46">
<taxonomicNameLabel id="BA8157E2E159BD6456B9FE18DDC08856" box="[655,745,1869,1894]" pageId="9" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(below).
</paragraph>
</subSubSection>
</treatment>
</document>
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