464 lines
68 KiB
XML
464 lines
68 KiB
XML
<document id="B1B7F1D7BACD5F9636B6E69DF6EC2E08" ID-DOI="10.11646/zootaxa.4048.1.2" ID-GBIF-Dataset="2f5da4b3-c0ee-4b70-b661-0e2ef9bacfa5" ID-ISSN="1175-5326" ID-Zenodo-Dep="290075" ID-ZooBank="97BB73FF-9D62-4A15-B09C-0F570E26E738" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461341944160" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2015" docId="1B6F879DE151BD6954A1FC91DEB18A65" docLanguage="en" docName="zt04048p056.pdf" docOrigin="Zootaxa 4048 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma amphinome De Man 1899" docType="treatment" docVersion="6" lastPageNumber="41" masterDocId="E756FFE5E150BD6D5436F955DF298F30" masterDocTitle="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species" masterLastPageNumber="56" masterPageNumber="37" pageNumber="38" updateTime="1698620270231" updateUser="plazi">
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<mods:title id="8FE4CD4780CE26E48A5C86C9F75029EA">Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species</mods:title>
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<mods:namePart id="45974BF72355BAF90579D807096B51F8">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B60FF109F3559E29778365033D62A4BA">Zootaxa</mods:title>
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<mods:date id="87DEC24711D05108DD22EFF19F33A49C">2015</mods:date>
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<mods:number id="BEC56B5B1C8412E495BDFA9D04480FE0">4048</mods:number>
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<treatment id="1B6F879DE151BD6954A1FC91DEB18A65" ID-DOI="http://doi.org/10.5281/zenodo.6121300" ID-GBIF-Taxon="127694320" ID-Zenodo-Dep="6121300" LSID="urn:lsid:plazi:treatment:1B6F879DE151BD6954A1FC91DEB18A65" httpUri="http://treatment.plazi.org/id/1B6F879DE151BD6954A1FC91DEB18A65" lastPageId="4" lastPageNumber="41" pageId="1" pageNumber="38">
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<subSubSection id="DBDC6500E151BD6C54A1FC91DE3B8930" pageId="1" pageNumber="38" type="nomenclature">
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<paragraph id="9379368BE151BD6C54A1FC91DDA08AEF" blockId="1.[151,649,1476,1537]" box="[151,649,1476,1503]" pageId="1" pageNumber="38">
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<heading id="C83181E7E151BD6C54A1FC91DDA08AEF" bold="true" box="[151,649,1476,1503]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
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<taxonomicName id="54C64D08E151BD6C54A1FC91DDA08AEF" ID-CoL="3FS8Q" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[151,649,1476,1503]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="amphinome">
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<emphasis id="A1B2EA99E151BD6C54A1FC91DDA08AEF" bold="true" box="[151,649,1476,1503]" pageId="1" pageNumber="38">
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<emphasis id="A1B2EA99E151BD6C54A1FC91DEE88AEE" bold="true" box="[151,449,1476,1502]" italics="true" pageId="1" pageNumber="38">Geosesarma amphinome</emphasis>
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(De
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<bibRefCitation id="F7574B7AE151BD6C55CAFC90DDA88AEF" author="Man" box="[508,641,1477,1503]" pageId="1" pageNumber="38" refString="Man, J. G. De (1899) Zoological Results of the Dutch Scientific Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="9379368BE151BD6C54A1FCBDDE3B8930" blockId="1.[151,649,1476,1537]" box="[151,274,1512,1537]" pageId="1" pageNumber="38">
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(
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<figureCitation id="0BFD2A0EE151BD6C54A9FCBDDFC58930" box="[159,236,1512,1537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="1" pageNumber="38">Figs. 1</figureCitation>
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,
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<figureCitation id="0BFD2A0EE151BD6C54CCFCBDDE208930" box="[250,265,1512,1536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="1" pageNumber="38">2</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DBDC6500E151BD6C54A1FF64DFD78992" pageId="1" pageNumber="38" type="reference_group">
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<paragraph id="9379368BE151BD6C54A1FF64DE5E8954" blockId="1.[151,1436,1584,1698]" pageId="1" pageNumber="38">
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<treatmentCitationGroup id="B3D611A5E151BD6C54A1FF64DE5E8954" pageId="1" pageNumber="38">
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<taxonomicName id="54C64D08E151BD6C54A1FF64DD998976" ID-CoL="4WYW3" authority="De Man, 1899: 133" authorityName="De Man" authorityPageNumber="133" authorityYear="1899" box="[151,688,1584,1606]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="amphinome" subGenus="Sesarma">
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<emphasis id="A1B2EA99E151BD6C54A1FF64DFC78976" box="[151,238,1585,1606]" italics="true" pageId="1" pageNumber="38">Sesarma</emphasis>
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(
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<emphasis id="A1B2EA99E151BD6C54C8FF64DE7C8976" box="[254,341,1585,1606]" italics="true" pageId="1" pageNumber="38">Sesarma</emphasis>
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)
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<emphasis id="A1B2EA99E151BD6C5550FF65DEF58976" box="[358,476,1584,1606]" italics="true" pageId="1" pageNumber="38">amphinome</emphasis>
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De
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<treatmentCitation id="1267109AE151BD6C563CFF64DD998976" author="Man" box="[522,688,1584,1606]" page="133" pageId="1" pageNumber="38" year="1899">
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<bibRefCitation id="F7574B7AE151BD6C563CFF64DD558976" author="Man" box="[522,636,1584,1606]" pageId="1" pageNumber="38" refString="Man, J. G. De (1899) Zoological Results of the Dutch Scientific Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
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: 133
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</treatmentCitation>
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</taxonomicName>
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, pl. 12 fig. 16; Tesch 1918: 129 (list);
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<treatmentCitation id="1267109AE151BD6C5072FF65DA398976" author="Serene" box="[1092,1296,1584,1606]" page="1087" pageId="1" pageNumber="38" year="1968">
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<bibRefCitation id="F7574B7AE151BD6C5072FF65DBFB8976" author="Serene" box="[1092,1234,1584,1606]" pageId="1" pageNumber="38" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène 1968a</bibRefCitation>
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: 1087
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</treatmentCitation>
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(list);
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<treatmentCitation id="1267109AE151BD6C5161FF65DE138954" author="Serene" page="106" pageId="1" pageNumber="38" year="1968">
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<bibRefCitation id="F7574B7AE151BD6C5161FF65DE238954" author="Serene" pageId="1" pageNumber="38" refString="Serene, R. (1968 b) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, 1, pp. 33 - 120." type="journal article" year="1968" yearSuffix="b">Serène 1968b</bibRefCitation>
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: 106
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</treatmentCitation>
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(list).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="9379368BE151BD6C54A1FF3BDFD78992" blockId="1.[151,1436,1584,1698]" pageId="1" pageNumber="38">
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||
<treatmentCitationGroup id="B3D611A5E151BD6C54A1FF3BDFD78992" pageId="1" pageNumber="38">
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||
<taxonomicName id="54C64D08E151BD6C54A1FF3BDEBA89B3" ID-CoL="3FS8Q" box="[151,403,1645,1667]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="amphinome">
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||
<emphasis id="A1B2EA99E151BD6C54A1FF3BDEBA89B3" box="[151,403,1645,1667]" italics="true" pageId="1" pageNumber="38">Geosesarma amphinome</emphasis>
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||
</taxonomicName>
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||
—
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<treatmentCitation id="1267109AE151BD6C559BFF38DD8589B3" author="Serene" box="[429,684,1645,1667]" page="402" pageId="1" pageNumber="38" year="1970">
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||
<bibRefCitation id="F7574B7AE151BD6C559BFF38DD5389B3" author="Serene" box="[429,634,1645,1667]" pageId="1" pageNumber="38" refString="Serene, R. & Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416, pls. 1 - 8." type="journal article" year="1970">Serène & Soh 1970</bibRefCitation>
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||
: 402
|
||
</treatmentCitation>
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||
(list); Ng 1988: 119 (list);
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||
<treatmentCitation id="1267109AE151BD6C57F1FF3BDBEC89B3" author="Fransen" box="[967,1221,1645,1667]" page="123" pageId="1" pageNumber="38" year="1997">
|
||
<bibRefCitation id="F7574B7AE151BD6C57F1FF3BDBB889B3" author="Fransen" box="[967,1169,1645,1667]" pageId="1" pageNumber="38" refString="Fransen, C. H. J. M., Holthuis, L. B. & Adema, J. P. H. M. (1997) Type-catalogue of the Decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologishe Verhandelingen, 311, i - xvi, 1 - 344, figs. 1 - 79." type="journal article" year="1997">
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Fransen
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<emphasis id="A1B2EA99E151BD6C5029FF3BDB7D89B3" box="[1055,1108,1645,1667]" italics="true" pageId="1" pageNumber="38">et al.</emphasis>
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1997
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</bibRefCitation>
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||
: 123
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||
</treatmentCitation>
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||
; Ng
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||
<emphasis id="A1B2EA99E151BD6C50CEFF3BDA0489B3" box="[1272,1325,1645,1667]" italics="true" pageId="1" pageNumber="38">et al.</emphasis>
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2008: 220 (list).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DBDC6500E151BD6C54A1FF9BDA3A883B" pageId="1" pageNumber="38" type="materials_examined">
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<paragraph id="9379368BE151BD6C54A1FF9BDA3A883B" blockId="1.[151,1437,1742,2019]" pageId="1" pageNumber="38">
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||
<emphasis id="A1B2EA99E151BD6C54A1FF9BDE5589D7" bold="true" box="[151,380,1742,1767]" pageId="1" pageNumber="38">Material examined</emphasis>
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.
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<typeStatus id="4C7D8829E151BD6C55BBFF85DD2889D8" box="[397,513,1744,1768]" pageId="1" pageNumber="38" type="lectotype">Lectotype</typeStatus>
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: female (11.1 ×
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<quantity id="543E9B6EE151BD6C56F7FF9BDC3289D7" box="[705,795,1742,1767]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="38" unit="mm" value="8.5">8.5 mm</quantity>
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||
) (
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<collectionCode id="F5D7AE4EE151BD6C5702FF9ADC5E89D7" box="[820,887,1743,1767]" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="38">NNM</collectionCode>
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1198), Sintang, Borneo, coll. M. Moret,
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<date id="E778104BE151BD6C5160FF9ADFFA883B" pageId="1" pageNumber="38" value="1893" valueMax="1894">1893– 1894</date>
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||
.
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||
<typeStatus id="4C7D8829E151BD6C54EBFFA7DEAC883B" box="[221,389,1778,1803]" pageId="1" pageNumber="38" type="paralectotype">Paralectotypes</typeStatus>
|
||
:
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||
<specimenCount id="85C0FD02E151BD6C55B9FFA6DECC883B" box="[399,485,1778,1803]" pageId="1" pageNumber="38" type="male">2 males</specimenCount>
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||
(larger 14.4 ×
|
||
<quantity id="543E9B6EE151BD6C56BAFFA6DDCF883B" box="[652,742,1779,1803]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="38" unit="mm" value="8.7">8.7 mm</quantity>
|
||
),
|
||
<specimenCount id="85C0FD02E151BD6C56CFFFA6DC73883B" box="[761,858,1778,1803]" pageId="1" pageNumber="38" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="F5D7AE4EE151BD6C575FFFA6DC85883B" box="[873,940,1779,1803]" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="38">NNM</collectionCode>
|
||
23283), same data as
|
||
<typeStatus id="4C7D8829E151BD6C5093FFA7DA39883B" box="[1189,1296,1778,1803]" pageId="1" pageNumber="38" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E151BD6F54F1FE43DE9E88FD" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="diagnosis">
|
||
<paragraph id="9379368BE151BD6F54F1FE43DC458E54" blockId="1.[151,1437,1742,2019]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
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||
<emphasis id="A1B2EA99E151BD6C54F1FE43DE13881F" bold="true" box="[199,314,1814,1839]" pageId="1" pageNumber="38">Diagnosis</emphasis>
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||
. Carapace subtrapezoidal, wider than long, width to length ratio 1.3–1.6, lateral margins gently diverging (
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||
<figureCitation id="0BFD2A0EE151BD6C5524FE6FDE748863" box="[274,349,1850,1875]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="1" pageNumber="38">Figs. 1</figureCitation>
|
||
A, B, 2A); dorsal surface with visible regions, anterior regions densely covered with low, small rounded, flattened granules (
|
||
<figureCitation id="0BFD2A0EE151BD6C55EEFE0BDD3E8847" box="[472,535,1886,1911]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
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||
A–D); front deflexed, frontal lobes broad with subtruncate margins; postfrontal, postorbital cristae low but distinct (
|
||
<figureCitation id="0BFD2A0EE151BD6C561CFED7DD4288AB" box="[554,619,1922,1947]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
|
||
C, D); external orbital tooth acutely triangular, directed obliquely, outer margin gently convex to almost straight, tip extending beyond lateral margin (
|
||
<figureCitation id="0BFD2A0EE151BD6C5016FEF3DB45888F" box="[1056,1132,1958,1983]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="1" pageNumber="38">Figs. 1</figureCitation>
|
||
A, B, 2A). Merus of third maxilliped subovate, subequal in length to ischium; exopod slender with long flagellum (
|
||
<figureCitation id="0BFD2A0EE151BD6C504DFE9FDB9688D3" box="[1147,1215,1994,2019]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="1" pageNumber="38">Fig. 2</figureCitation>
|
||
B). Outer surface of palm of adult male covered with small rounded granules; inner surface with transverse comb-like ridge; dorsal margin of dactylus with cluster of small rounded granules on proximal part, 3 or 4 slightly larger granules after it, unarmed on distal half (
|
||
<figureCitation id="0BFD2A0EE152BD6F5594F98ADEC28FC8" box="[418,491,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="2" pageNumber="39">Figs. 1</figureCitation>
|
||
F, 2F). Ambulatory legs with relatively slender merus, with sharp subdistal spine on dorsal margin, surfaces gently rugose (
|
||
<figureCitation id="0BFD2A0EE152BD6F5660F851DD8B8E2C" box="[598,674,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="2" pageNumber="39">Figs. 1</figureCitation>
|
||
A, B, 2D). Male abdomen triangular; somite 6 wide, with convex lateral margins; telson semicircular (
|
||
<figureCitation id="0BFD2A0EE152BD6F560FF872DDAC8E70" box="[569,645,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="2" pageNumber="39">Figs. 1</figureCitation>
|
||
E, 2C). G1 relatively slender, distal pectinated part elongated, gently curved, expanded, spatuliform, tip medially clefted (
|
||
<figureCitation id="0BFD2A0EE152BD6F56D4F819DC0C8E54" box="[738,805,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="2" pageNumber="39">Fig. 2</figureCitation>
|
||
G–K).
|
||
</paragraph>
|
||
<caption id="C7B96603E152BD6F54A1FE46DAAD8856" httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="2" pageNumber="39" targetBox="[151,1435,411,1761]" targetPageId="2">
|
||
<paragraph id="9379368BE152BD6F54A1FE46DAAD8856" blockId="2.[151,1436,1811,1894]" pageId="2" pageNumber="39">
|
||
<emphasis id="A1B2EA99E152BD6F54A1FE46DE258819" bold="true" box="[151,268,1811,1833]" pageId="2" pageNumber="39">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E152BD6F552EFE41DD978819" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[280,702,1811,1833]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E152BD6F552EFE41DD3A8819" box="[280,531,1811,1833]" italics="true" pageId="2" pageNumber="39">Geosesarma amphinome</emphasis>
|
||
(De Man, 1899)
|
||
</taxonomicName>
|
||
. A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B– F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B–F after De Man 1899: pl. 12 fig. 16).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9379368BE152BD6F54F1FEC5DE9E88FD" blockId="2.[199,439,1936,1997]" pageId="2" pageNumber="39">
|
||
<emphasis id="A1B2EA99E152BD6F54F1FEC5DE338899" bold="true" box="[199,282,1936,1961]" pageId="2" pageNumber="39">Colour</emphasis>
|
||
. Not known.
|
||
<emphasis id="A1B2EA99E152BD6F54F1FEE1DE0D88FD" bold="true" box="[199,292,1972,1997]" pageId="2" pageNumber="39">Ecology</emphasis>
|
||
. Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C7B96603E153BD6E54A1FC8FDB32895A" httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="3" pageNumber="40" targetBox="[157,1421,188,1471]" targetPageId="3">
|
||
<paragraph id="9379368BE153BD6E54A1FC8FDB32895A" blockId="3.[151,1436,1498,1642]" pageId="3" pageNumber="40">
|
||
<emphasis id="A1B2EA99E153BD6E54A1FC8FDE258AC0" bold="true" box="[151,268,1498,1520]" pageId="3" pageNumber="40">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="54C64D08E153BD6E552EFC8EDD958AC0" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[280,700,1498,1520]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E153BD6E552EFC8EDD3B8AC0" box="[280,530,1498,1520]" italics="true" pageId="3" pageNumber="40">Geosesarma amphinome</emphasis>
|
||
(De Man, 1899)
|
||
</taxonomicName>
|
||
, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G1 (ventral view); H, left G1 (dorsal view); I, distal part of left G1 (ventral view); J, distal part of left G1 (dorsal view); K, distal part of left G1 (subdorsal view); L, left G2. All structures denuded. Scales: A–D = 1.0 mm; E, F = 0.5 mm; G–L = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DBDC6500E153BD6E54F1FFC1DA7F88B5" pageId="3" pageNumber="40" type="discussion">
|
||
<paragraph id="9379368BE153BD6E54F1FFC1DBEB89C5" blockId="3.[151,1437,1684,2033]" pageId="3" pageNumber="40">
|
||
<emphasis id="A1B2EA99E153BD6E54F1FFC1DE1D899D" bold="true" box="[199,308,1684,1709]" pageId="3" pageNumber="40">Remarks</emphasis>
|
||
. The species was described on the basis of two male and three female specimens collected probably from Sintang, along the Kapuas River in western Kalimantan, Borneo (De
|
||
<bibRefCitation id="F7574B7AE153BD6E57D3FFECDB7589E1" author="Man" box="[997,1116,1721,1745]" pageId="3" pageNumber="40" refString="Man, J. G. De (1899) Zoological Results of the Dutch Scientific Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21, 53 - 144, pls. 5 - 12." type="journal article" year="1899">Man 1899</bibRefCitation>
|
||
: 133).
|
||
<bibRefCitation id="F7574B7AE153BD6E509CFFECDA6689E1" author="Serene" box="[1194,1359,1720,1745]" pageId="3" pageNumber="40" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène (1968a</bibRefCitation>
|
||
: 1089) identified a female specimen as the
|
||
<typeStatus id="4C7D8829E153BD6E561EFF88DDA689C6" box="[552,655,1757,1782]" pageId="3" pageNumber="40" type="lectotype">lectotype</typeStatus>
|
||
without discussion (see
|
||
<bibRefCitation id="F7574B7AE153BD6E5794FF8BDB5489C5" author="Fransen" box="[930,1149,1757,1781]" pageId="3" pageNumber="40" refString="Fransen, C. H. J. M., Holthuis, L. B. & Adema, J. P. H. M. (1997) Type-catalogue of the Decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologishe Verhandelingen, 311, i - xvi, 1 - 344, figs. 1 - 79." type="journal article" year="1997">
|
||
Fransen
|
||
<emphasis id="A1B2EA99E153BD6E5034FF8BDB1389C5" box="[1026,1082,1757,1781]" italics="true" pageId="3" pageNumber="40">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
: 123).
|
||
</paragraph>
|
||
<paragraph id="9379368BE153BD6E54F1FE55DA7F88B5" blockId="3.[151,1437,1684,2033]" pageId="3" pageNumber="40">
|
||
The
|
||
<typeStatus id="4C7D8829E153BD6E54CAFE55DE4A8829" box="[252,355,1792,1817]" pageId="3" pageNumber="40" type="lectotype">lectotype</typeStatus>
|
||
female is an adult; while the larger
|
||
<typeStatus id="4C7D8829E153BD6E5730FE55DCB48829" box="[774,925,1792,1817]" pageId="3" pageNumber="40" type="paralectotype">paralectotype</typeStatus>
|
||
male is mature, its chelae are still relatively small. Its male abdomen and G1, however, are fully formed. One characteristic feature of this species is that although the dorsal margin of the dactylus is mostly unarmed except for a few small granules on the proximal part (
|
||
<figureCitation id="0BFD2A0EE153BD6E54A9FE38DFC188B5" box="[159,232,1901,1926]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="3" pageNumber="40">Figs. 1</figureCitation>
|
||
F, 2F), there is a prominent transverse comb-like structure on the inner surface of the chela (
|
||
<figureCitation id="0BFD2A0EE153BD6E50C5FE38DA1E88B5" box="[1267,1335,1901,1926]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DBDC6500E153BD6954F1FEC4DEB18A65" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="reference_group">
|
||
<paragraph id="9379368BE153BD6954F1FEC4DBFE8E2D" blockId="3.[151,1437,1684,2033]" lastBlockId="4.[151,1437,151,1365]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
|
||
<taxonomicName id="54C64D08E153BD6E54F1FEC4DEF78899" ID-CoL="3FS8Q" box="[199,478,1937,1961]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E153BD6E54F1FEC4DEF78899" box="[199,478,1937,1961]" italics="true" pageId="3" pageNumber="40">Geosesarma amphinome</emphasis>
|
||
</taxonomicName>
|
||
is superficially similar to
|
||
<taxonomicName id="54C64D08E153BD6E573EFEC4DB4E8899" ID-CoL="6KBJ7" authority="Serene, 1968" authorityName="Serene" authorityYear="1968" box="[776,1127,1936,1961]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E153BD6E573EFEC4DC908899" box="[776,953,1937,1961]" italics="true" pageId="3" pageNumber="40">G. sarawakense</emphasis>
|
||
(Serène, 1968)
|
||
</taxonomicName>
|
||
(from central
|
||
<collectingCountry id="EBD1761BE153BD6E513AFEC5DA588899" box="[1292,1393,1936,1961]" name="Malaysia" pageId="3" pageNumber="40">Sarawak</collectingCountry>
|
||
),
|
||
<taxonomicName id="54C64D08E153BD6E51B2FEC4DE8388FD" ID-CoL="3FS9T" authority="Nobili, 1903" authorityName="Nobili" authorityYear="1903" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="A1B2EA99E153BD6E51B2FEC4DFD788FD" italics="true" pageId="3" pageNumber="40">G. peraccae</emphasis>
|
||
(Nobili, 1903)
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="EBD1761BE153BD6E55CBFEE0DD4688FE" box="[509,623,1973,1998]" name="Singapore" pageId="3" pageNumber="40">Singapore</collectingCountry>
|
||
and southern Peninsular
|
||
<collectingCountry id="EBD1761BE153BD6E57A3FEE0DB2988FE" box="[917,1024,1973,1998]" name="Malaysia" pageId="3" pageNumber="40">Malaysia</collectingCountry>
|
||
), and
|
||
<taxonomicName id="54C64D08E153BD6E507EFEE0DBC488FD" ID-CoL="3FS9W" box="[1096,1261,1973,1997]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E153BD6E507EFEE0DBC488FD" box="[1096,1261,1973,1997]" italics="true" pageId="3" pageNumber="40">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E153BD6E50C0FEE0DA7B88FE" bold="true" box="[1270,1362,1973,1998]" pageId="3" pageNumber="40">
|
||
<taxonomicNameLabel id="BA8157E2E153BD6E50C0FEE0DA7B88FE" box="[1270,1362,1973,1998]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(from Anjungan, Kalimantan Barat), with similar trapezoidal carapace shapes and the exopod of their third maxillipeds possess a long flagellum (
|
||
<figureCitation id="0BFD2A0EE154BD695580F9C2DED68F80" box="[438,511,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
|
||
A, B, 2A, B, 3A, B, 4A, 5A, B, 6A, B, 7A;
|
||
<bibRefCitation id="F7574B7AE154BD6957D5F9CDDB578F80" author="Serene" box="[995,1150,151,176]" pageId="4" pageNumber="41" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène 1968a</bibRefCitation>
|
||
: pl. 1 fig. 3; Ng 1988: fig. 56A, C).
|
||
<taxonomicName id="54C64D08E154BD695537F9E9DD3E8FE4" ID-CoL="3FS8Q" box="[257,535,188,212]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD695537F9E9DD3E8FE4" box="[257,535,188,212]" italics="true" pageId="4" pageNumber="41">Geosesarma amphinome</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="54C64D08E154BD695610F9E9DD828FE4" ID-CoL="3FS9T" box="[550,683,188,212]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="A1B2EA99E154BD695610F9E9DD828FE4" box="[550,683,188,212]" italics="true" pageId="4" pageNumber="41">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E154BD6956D2F9E9DCAC8FE4" ID-CoL="3FS9W" box="[740,901,188,212]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD6956D2F9E9DCAC8FE4" box="[740,901,188,212]" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD6957B8F9E9DCC18FE5" bold="true" box="[910,1000,188,213]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD6957B8F9E9DCC18FE5" box="[910,1000,188,213]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possess a prominent transverse comblike structure on the inner surface of the chela (
|
||
<figureCitation id="0BFD2A0EE154BD695692F98ADDD88FC8" box="[676,753,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="4" pageNumber="41">Figs. 2</figureCitation>
|
||
F, 3F, 5F; Ng 1988: fig. 56A). The chela of the
|
||
<typeStatus id="4C7D8829E154BD6950C8F98ADA498FC8" box="[1278,1376,223,248]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="54C64D08E154BD6951B2F9B5DE018E2C" ID-CoL="6KBJ7" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E154BD6951B2F9B5DE018E2C" italics="true" pageId="4" pageNumber="41">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
is small (cf.
|
||
<bibRefCitation id="F7574B7AE154BD69558FF851DD7C8E2D" author="Serene" box="[441,597,260,285]" pageId="4" pageNumber="41" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène 1968a</bibRefCitation>
|
||
: pl. 1 fig. 3) but also has a low transverse ridge (
|
||
<figureCitation id="0BFD2A0EE154BD695043F851DB918E2D" box="[1141,1208,260,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[151,1435,193,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Geosesarma sarawakense (Serène, 1968), holotype male (11.8 × 10.3 mm) (ZRC 1965.7.29.6), Sibu, Sarawak, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, outer view of right chela; F, inner view of right chela." httpUri="https://zenodo.org/record/290081/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<paragraph id="9379368BE154BD6954F1F87DDE068D50" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="41">
|
||
<taxonomicName id="54C64D08E154BD6954F1F87DDEF58E70" ID-CoL="3FS8Q" box="[199,476,296,320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD6954F1F87DDEF58E70" box="[199,476,296,320]" italics="true" pageId="4" pageNumber="41">Geosesarma amphinome</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="54C64D08E154BD69573CF87DDC938E70" ID-CoL="6KBJ7" box="[778,954,296,320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E154BD69573CF87DDC938E70" box="[778,954,296,320]" italics="true" pageId="4" pageNumber="41">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
by the dorsal surface of the anterior part of the carapace being more prominently granular (
|
||
<figureCitation id="0BFD2A0EE154BD695699F819DDDE8E54" box="[687,759,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
|
||
A–C, 2A versus
|
||
<figureCitation id="0BFD2A0EE154BD69578EF819DCD28E55" box="[952,1019,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[151,1435,193,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURE 6. Geosesarma sarawakense (Serène, 1968), holotype male (11.8 × 10.3 mm) (ZRC 1965.7.29.6), Sibu, Sarawak, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, outer view of right chela; F, inner view of right chela." httpUri="https://zenodo.org/record/290081/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
|
||
A–C;
|
||
<bibRefCitation id="F7574B7AE154BD695009F819DBF28E55" author="Serene" box="[1087,1243,332,357]" pageId="4" pageNumber="41" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène 1968a</bibRefCitation>
|
||
: pl. 1 fig. 3), and the distal pectinated part of the G1 is more curved, spatuliform and relatively longer (
|
||
<figureCitation id="0BFD2A0EE154BD6950A0F83ADBF58EB8" box="[1174,1244,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
|
||
G–K) (the distal pectinated part of the G1 is straight and relatively shorter in
|
||
<taxonomicName id="54C64D08E154BD695776F8C1DCD98E9C" ID-CoL="6KBJ7" box="[832,1008,404,428]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E154BD695776F8C1DCD98E9C" box="[832,1008,404,428]" italics="true" pageId="4" pageNumber="41">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="0BFD2A0EE154BD6957C8F8C1DB168E9C" box="[1022,1087,404,429]" captionStart="FIGURE 7" captionStartId="9.[151,250,1172,1194]" captionTargetBox="[165,1423,208,1139]" captionTargetId="figure@9.[151,1436,193,1151]" captionTargetPageId="9" captionText="FIGURE 7. Geosesarma sarawakense (Serène, 1968), holotype male (11.8 × 10.3 mm) (ZRC 1965.7.29.6), Sibu, Sarawak, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, left last second ambulatory leg; D, left G 1 (subventral view); E, left G 1 (ventral view); F, left G 1 (dorsal view). All structures denuded. Scales: A – C = 1.0 mm; D – F = 0.5 mm." httpUri="https://zenodo.org/record/290082/files/figure.png" pageId="4" pageNumber="41">Fig. 7</figureCitation>
|
||
D–F;
|
||
<bibRefCitation id="F7574B7AE154BD6950B7F8C1DA348E9D" author="Serene" box="[1153,1309,404,429]" pageId="4" pageNumber="41" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="journal article" year="1968" yearSuffix="a">Serène 1968a</bibRefCitation>
|
||
: figs. 7, 8). The dorsal surface of the carapace, male abdomen and G1 of
|
||
<taxonomicName id="54C64D08E154BD695709F8EDDCED8EE0" ID-CoL="3FS9T" box="[831,964,440,464]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="A1B2EA99E154BD695709F8EDDCED8EE0" box="[831,964,440,464]" italics="true" pageId="4" pageNumber="41">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
is distinctly different from
|
||
<taxonomicName id="54C64D08E154BD6950C2F8EDDABC8EE0" ID-CoL="3FS8Q" box="[1268,1429,440,464]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD6950C2F8EDDABC8EE0" box="[1268,1429,440,464]" italics="true" pageId="4" pageNumber="41">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
, with the dorsal surface prominently more granular, somite 6 proportionately broader and the G1 being very slender, with the distal pectinated part almost straight (
|
||
<figureCitation id="0BFD2A0EE154BD6956A9F8AADDC98D28" box="[671,736,511,536]" captionStart="FIGURE 5" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[151,1435,193,1832]" captionTargetId="figure@7.[151,1436,193,1833]" captionTargetPageId="7" captionText="FIGURE 5. Geosesarma peraccae (Nobili, 1903), male (14.2 × 12.0 mm) (ZRC 2010.324), Nee Soon Swamp Forest, Singapore. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax showing thoracic sternum and abdomen; E, outer view of left chela; F, inner view of left chela." httpUri="https://zenodo.org/record/290080/files/figure.png" pageId="4" pageNumber="41">Fig. 5</figureCitation>
|
||
A, B, D; Ng 1988: fig. 56D–F) (the carapace less granular, the abdominal somite 6 less broad, and the distal pectinated part wider and gently curved in
|
||
<taxonomicName id="54C64D08E154BD69504BFB71DA368D0C" ID-CoL="3FS8Q" box="[1149,1311,548,572]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD69504BFB71DA368D0C" box="[1149,1311,548,572]" italics="true" pageId="4" pageNumber="41">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="0BFD2A0EE154BD69511BFB71DA5C8D0C" box="[1325,1397,548,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
|
||
A– D, 2C, G–K).
|
||
</paragraph>
|
||
<paragraph id="9379368BE154BD6954F1FB39DE368CF8" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="41">
|
||
Although both
|
||
<taxonomicName id="54C64D08E154BD69554FFB39DD378DB4" ID-CoL="3FS8Q" box="[377,542,620,644]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD69554FFB39DD378DB4" box="[377,542,620,644]" italics="true" pageId="4" pageNumber="41">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E154BD69566BFB39DC2A8DB4" ID-CoL="3FS9W" box="[605,771,620,644]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD69566BFB39DC2A8DB4" box="[605,771,620,644]" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD69573BFB39DC428DB5" bold="true" box="[781,875,620,645]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD69573BFB39DC428DB5" box="[781,875,620,645]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occur in Kalimantan Barat, their
|
||
<typeStatus id="4C7D8829E154BD6950CAFB38DA058DB5" box="[1276,1324,621,645]" pageId="4" pageNumber="41">type</typeStatus>
|
||
localities (Sintang and Anjungan, respectively) are almost
|
||
<quantity id="543E9B6EE154BD6956F6FBC5DC3F8D98" box="[704,790,655,680]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="41" unit="km" value="200.0">200 km</quantity>
|
||
apart.
|
||
<taxonomicName id="54C64D08E154BD695753FBC5DB558D98" ID-CoL="3FS8Q" box="[869,1148,656,680]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD695753FBC5DB558D98" box="[869,1148,656,680]" italics="true" pageId="4" pageNumber="41">Geosesarma amphinome</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="54C64D08E154BD6951B2FBC5DE338DFC" ID-CoL="3FS9W" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD6951B2FBC5DE338DFC" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD695517FBE1DE528DFD" bold="true" box="[289,379,692,717]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD695517FBE1DE528DFD" box="[289,379,692,717]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having the anterior half of the dorsal surface of the carapace relatively more granular (
|
||
<figureCitation id="0BFD2A0EE154BD695158FBE1DF898DC0" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
|
||
A–C versus
|
||
<figureCitation id="0BFD2A0EE154BD695503FB82DE508DC0" box="[309,377,727,752]" captionStart="FIGURE 3" captionStartId="5.[151,250,1926,1948]" captionTargetBox="[151,1435,265,1904]" captionTargetId="figure@5.[151,1436,265,1905]" captionTargetPageId="5" captionText="FIGURE 3. Geosesarma pylaemenes sp. nov., holotype male (11.6 × 10.6 mm) (MZB Cru 4394), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, dorsal view of right chela; E, outer view of right chela; F, inner view of right chela." httpUri="https://zenodo.org/record/290078/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
A, B), the external orbital tooth is more acutely triangular (
|
||
<figureCitation id="0BFD2A0EE154BD695018FB82DB508DC0" box="[1070,1145,727,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
|
||
A–C, 2A) (more broadly triangular in
|
||
<taxonomicName id="54C64D08E154BD695511FBA9DEE18C24" ID-CoL="3FS9W" box="[295,456,764,788]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD695511FBA9DEE18C24" box="[295,456,764,788]" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD6955F8FBA9DD0E8C25" bold="true" box="[462,551,764,789]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD6955F8FBA9DD0E8C25" box="[462,551,764,789]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE154BD695603FBA9DD5C8C24" box="[565,629,764,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1926,1948]" captionTargetBox="[151,1435,265,1904]" captionTargetId="figure@5.[151,1436,265,1905]" captionTargetPageId="5" captionText="FIGURE 3. Geosesarma pylaemenes sp. nov., holotype male (11.6 × 10.6 mm) (MZB Cru 4394), Anjungan, Kalimantan Barat, Borneo. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, dorsal view of right chela; E, outer view of right chela; F, inner view of right chela." httpUri="https://zenodo.org/record/290078/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
A, B), abdominal somite 6 is proportionately less broad and the telson is not recessed into its distal margin (
|
||
<figureCitation id="0BFD2A0EE154BD69562BFA4ADD4E8C08" box="[541,615,799,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1833]" captionTargetBox="[151,1435,411,1761]" captionTargetId="figure@2.[151,1436,411,1786]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma amphinome (De Man, 1899). A, lectotype female (11.1 × 8.5 mm) (NNM 1198), Sintang, Borneo; B – F, paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, B, overall habitus; C, dorsal view of anterior part of carapace; D, frontal view of carapace; E, male abdomen; F, outer view of left chela. (B – F after De Man 1899: pl. 12 fig. 16)." httpUri="https://zenodo.org/record/290076/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
|
||
E, 2C) (somite 6 broader and the telson is partially recessed into its distal margin in
|
||
<taxonomicName id="54C64D08E154BD695523FA11DE958C6C" ID-CoL="3FS9W" box="[277,444,836,860]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD695523FA11DE958C6C" box="[277,444,836,860]" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD6955FFFA11DD008C6D" bold="true" box="[457,553,836,861]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD6955FFFA11DD008C6D" box="[457,553,836,861]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE154BD69560BFA11DDAF8C6C" box="[573,646,836,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1126,1148]" captionTargetBox="[155,1423,204,1086]" captionTargetId="figure@6.[150,1436,193,1105]" captionTargetPageId="6" captionText="FIGURE 4. Geosesarma pylaemenes sp. nov., holotype male (11.6 × 10.6 mm) (MZB Cru 4394), Anjungan, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, right last second ambulatory leg; D, left G 1 (dorsal view); E, distal part of left G 1 (dorsal view); F, left G 1 (subventral view); G, left G 1 (ventral view). All structures denuded. Scales: A – C = 1.0 mm; D – G = 0.5 mm." httpUri="https://zenodo.org/record/290079/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
|
||
B), and the G1 is relatively shorter with the distal pectinated part proportionately broader and more spatuliform, and with the median part deeply clefted (
|
||
<figureCitation id="0BFD2A0EE154BD695040FA32DB9E8CB0" box="[1142,1207,871,896]" captionStart="FIGURE 2" captionStartId="3.[151,250,1498,1520]" captionTargetBox="[157,1421,188,1471]" captionTargetId="figure@3.[150,1436,177,1477]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma amphinome (De Man, 1899), paralectotype male (14.4 × 8.7 mm) (NNM 23283), Sintang, Borneo. A, dorsal view of carapace; B, left third maxilliped; C, male abdominal somites 5, 6 and telson; D, right fourth ambulatory leg (setae not drawn); E, dorsal view of left carpus; F, inner surface view of left chela; G, left G 1 (ventral view); H, left G 1 (dorsal view); I, distal part of left G 1 (ventral view); J, distal part of left G 1 (dorsal view); K, distal part of left G 1 (subdorsal view); L, left G 2. All structures denuded. Scales: A – D = 1.0 mm; E, F = 0.5 mm; G – L = 0.25 mm." httpUri="https://zenodo.org/record/290077/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
|
||
G–K) (G1 relatively more slender with the distal pectinated part narrower and not strongly clefted medially in
|
||
<taxonomicName id="54C64D08E154BD6950A6FAD9DA1A8C94" ID-CoL="3FS9W" box="[1168,1331,908,932]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pylaemenes" status="sp. nov.">
|
||
<emphasis id="A1B2EA99E154BD6950A6FAD9DA1A8C94" box="[1168,1331,908,932]" italics="true" pageId="4" pageNumber="41">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A1B2EA99E154BD69510DFAD9DABC8C95" bold="true" box="[1339,1429,908,933]" pageId="4" pageNumber="41">
|
||
<taxonomicNameLabel id="BA8157E2E154BD69510DFAD9DABC8C95" box="[1339,1429,908,933]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="0BFD2A0EE154BD6954A1FAFADFF08CF8" box="[151,217,943,968]" captionStart="FIGURE 4" captionStartId="6.[151,250,1126,1148]" captionTargetBox="[155,1423,204,1086]" captionTargetId="figure@6.[150,1436,193,1105]" captionTargetPageId="6" captionText="FIGURE 4. Geosesarma pylaemenes sp. nov., holotype male (11.6 × 10.6 mm) (MZB Cru 4394), Anjungan, Kalimantan Barat, Borneo. A, left third maxilliped; B, male abdominal somites 1 – 6 and telson; C, right last second ambulatory leg; D, left G 1 (dorsal view); E, distal part of left G 1 (dorsal view); F, left G 1 (subventral view); G, left G 1 (ventral view). All structures denuded. Scales: A – C = 1.0 mm; D – G = 0.5 mm." httpUri="https://zenodo.org/record/290079/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
|
||
D–G).
|
||
</paragraph>
|
||
<paragraph id="9379368BE154BD6954F1FA81DA358BF4" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="41">
|
||
<taxonomicName id="54C64D08E154BD6954F1FA81DED98CDC" ID-CoL="6KBJ7" box="[199,496,980,1004]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E154BD6954F1FA81DED98CDC" box="[199,496,980,1004]" italics="true" pageId="4" pageNumber="41">Geosesarma sarawakense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="54C64D08E154BD695619FA81DD908CDC" ID-CoL="3FS9T" box="[559,697,980,1004]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="A1B2EA99E154BD695619FA81DD908CDC" box="[559,697,980,1004]" italics="true" pageId="4" pageNumber="41">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
are both lowland species, typically associated with freshwater swamps or muddy forested habitats. Considering their similarity with
|
||
<taxonomicName id="54C64D08E154BD69578AFAADDB498B20" ID-CoL="3FS8Q" box="[956,1120,1016,1040]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD69578AFAADDB498B20" box="[956,1120,1016,1040]" italics="true" pageId="4" pageNumber="41">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
, the latter species probably lives in similar habitats.
|
||
<taxonomicName id="54C64D08E154BD6955F3FD49DDEF8B04" ID-CoL="3FS9T" box="[453,710,1052,1076]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="A1B2EA99E154BD6955F3FD49DDEF8B04" box="[453,710,1052,1076]" italics="true" pageId="4" pageNumber="41">Geosesarma peraccae</emphasis>
|
||
</taxonomicName>
|
||
has also been observed to forage in cups of pitcher plants (Nepentheceae) (
|
||
<bibRefCitation id="F7574B7AE154BD69556FFD15DD308B68" author="Tan" box="[345,537,1088,1112]" pageId="4" pageNumber="41" refString="Tan, H. H. & Ng, P. K. L. (2008) First record in Singapore of a nepenthiphilous crab, Geosesarma perracae (Crustacea: Decapoda: Sesarmidae). Nature in Singapore, 1, 201 - 205." type="journal article" year="2008">Tan & Ng 2008</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="54C64D08E154BD695655FD15DC218B68" ID-CoL="3FS8Q" box="[611,776,1088,1112]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="A1B2EA99E154BD695655FD15DC218B68" box="[611,776,1088,1112]" italics="true" pageId="4" pageNumber="41">G. amphinome</emphasis>
|
||
</taxonomicName>
|
||
may have similar phytotelmic habits. In any case, other species of
|
||
<taxonomicName id="54C64D08E154BD69552FFD31DE8C8B4B" ID-CoL="7P3H8" box="[281,421,1124,1147]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A1B2EA99E154BD69552FFD31DE8C8B4B" box="[281,421,1124,1147]" italics="true" pageId="4" pageNumber="41">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
have been reported from this habitat:
|
||
<taxonomicName id="54C64D08E154BD69574EFD31DB868B4D" ID-CoL="3FS9L" authority="Ng & Lim" authorityName="Ng & Lim" box="[888,1199,1124,1149]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="malayanum">
|
||
<emphasis id="A1B2EA99E154BD69574EFD31DB098B4C" box="[888,1056,1124,1148]" italics="true" pageId="4" pageNumber="41">G. malayanum</emphasis>
|
||
Ng & Lim
|
||
</taxonomicName>
|
||
, in Ng, 1986, from Peninsular
|
||
<collectingCountry id="EBD1761BE154BD695521FDD2DE578B90" box="[279,382,1159,1184]" name="Malaysia" pageId="4" pageNumber="41">Malaysia</collectingCountry>
|
||
(Ng &
|
||
<bibRefCitation id="F7574B7AE154BD6955E8FDD2DD7A8B90" author="Lim" box="[478,595,1159,1184]" pageId="4" pageNumber="41" refString="Ng, P. K. L. & Lim, R. P. (1987) The taxonomy and biology of the nepenthiphilous sesarmine freshwater crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea, Decapoda, Brachyura, Grapsidae) from Peninsular Malaysia. Malayan Nature Journal, 41, 393 - 402." type="journal article" year="1987">Lim 1987</bibRefCitation>
|
||
);
|
||
<taxonomicName id="54C64D08E154BD69565CFDDDDC748B90" ID-CoL="3FS9N" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" box="[618,861,1159,1185]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="nemesis">
|
||
<emphasis id="A1B2EA99E154BD69565CFDDDDDCC8BAF" box="[618,741,1160,1183]" italics="true" pageId="4" pageNumber="41">G. nemesis</emphasis>
|
||
Ng, 1986
|
||
</taxonomicName>
|
||
, from southern Peninsular
|
||
<collectingCountry id="EBD1761BE154BD6950A0FDD2DA298B90" box="[1174,1280,1159,1184]" name="Malaysia" pageId="4" pageNumber="41">Malaysia</collectingCountry>
|
||
;
|
||
<emphasis id="A1B2EA99E154BD695139FDDDDA718B90" box="[1295,1368,1160,1184]" italics="true" pageId="4" pageNumber="41">G. b a u</emphasis>
|
||
from
|
||
<collectingCountry id="EBD1761BE154BD6954A1FDF9DFDE8BF5" box="[151,247,1196,1221]" name="Malaysia" pageId="4" pageNumber="41">Sarawak</collectingCountry>
|
||
(see comparative material examined) and
|
||
<taxonomicName id="54C64D08E154BD6956E5FDF9DC528BF4" ID-CoL="3FS99" box="[723,891,1196,1220]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="A1B2EA99E154BD6956E5FDF9DC528BF4" box="[723,891,1196,1220]" italics="true" pageId="4" pageNumber="41">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="EBD1761BE154BD695789FDF9DB228BF5" box="[959,1035,1196,1221]" name="Brunei" pageId="4" pageNumber="41">Brunei</collectingCountry>
|
||
(S. Choy, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="9379368BE154BD6954F1FD85DEB18A65" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="41">
|
||
It is noteworthy that the
|
||
<typeStatus id="4C7D8829E154BD695633FD9ADD4E8BD8" box="[517,615,1231,1256]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="54C64D08E154BD6956A8FD85DC7F8BD8" ID-CoL="6KBJ7" box="[670,854,1232,1256]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="A1B2EA99E154BD6956A8FD85DC7F8BD8" box="[670,854,1232,1256]" italics="true" pageId="4" pageNumber="41">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
was actually collected with another species of
|
||
<taxonomicName id="54C64D08E154BD6954A1FDA1DE0A8A3B" ID-CoL="7P3H8" box="[151,291,1268,1291]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A1B2EA99E154BD6954A1FDA1DE0A8A3B" box="[151,291,1268,1291]" italics="true" pageId="4" pageNumber="41">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
. This specimen, an ovigerous female (11.2 ×
|
||
<quantity id="543E9B6EE154BD69571AFDA1DCBB8A3C" box="[812,914,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="4" pageNumber="41" unit="mm" value="10.9">10.9 mm</quantity>
|
||
) (ZRC 1965.7.29.5), has a quadrate carapace, longer ambulatory legs and is clearly a separate species. In the absence of a male, it is provisionally referred to
|
||
<taxonomicName id="54C64D08E154BD6951B2FC4DDFE88A64" ID-CoL="3FS8V" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="A1B2EA99E154BD6951B2FC4DDFE88A64" italics="true" pageId="4" pageNumber="41">G. bau</emphasis>
|
||
</taxonomicName>
|
||
for the time being.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |