234 lines
23 KiB
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234 lines
23 KiB
XML
<document ID-CLB-Dataset="21365" ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. & Hofman, Sebastian" docDate="2020" docId="1B2BA0EC1C6452D79A3D1249E5451D1B" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Imeretiopsis nakeralaensis Grego & Mumladze 2020, sp. nov." docType="treatment" docUuid="38835E75-8986-46B0-9040-F427F5E12A57" docUuidSource="ZooBank" docVersion="7" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1732936627699" updateUser="ExternalLinkService">
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<mods:mods id="B30343CF2B7C9D0C3C8243564B89B0D4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C2C09C2796E888F267F9A8DEEFB9D313">
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<mods:title id="6B80B58C8DE33CC0DB6A808715030605">Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
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</mods:titleInfo>
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<mods:name id="375CA0BA28132C2325CD730C9C48EA33" type="personal">
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<mods:role id="C2411EEBCB69145662420874445CA6FC">
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<mods:roleTerm id="67A183B457B3702AF3F45217B281E823">Author</mods:roleTerm>
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<mods:namePart id="2AB0C7726DEB91CA6FE5F1CD09D5C5D5">Grego, Jozef</mods:namePart>
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<mods:affiliation id="789FB40EA6ECD6D482BB022544E682DF">Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name id="0BBBCC1BA660AB8F7C053DD2F1E3110A" type="personal">
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<mods:role id="8562CA85F8051A572B9DA6A6C72897CC">
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<mods:roleTerm id="4C2D5F618EA27487E915D6C3C320CDB8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DAC71C978D403A85F04F18E02FB6D16C">Mumladze, Levan</mods:namePart>
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<mods:affiliation id="F3465B69BC90C8AEB9B85EC4EEC952E8">Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
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<mods:nameIdentifier id="1A0C4221AF86774197C32F60EA6B4A82" type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
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</mods:name>
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<mods:name id="527CF755238CCB95AE98B01B60D32EE4" type="personal">
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<mods:roleTerm id="6D7AB0B1813C319BD1B62FD6BC907F84">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87363F1586FEE9CC6C9458946054E133">Falniowski, Andrzej</mods:namePart>
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<mods:affiliation id="9549BA15D2B08CEBB42E53D352ED468D">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="B54D1B76B5B6E34E57875AAB8D2B73BA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C3EE13FBA332B16C5AE6881057B61899">Osikowski, Artur</mods:namePart>
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<mods:affiliation id="BA690316BF857488928DB3D6EE4F9CA6">Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
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<mods:nameIdentifier id="1539921D7013119FF719B1E95E34CDFC" type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
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<mods:nameIdentifier id="A3E02261B3CD8FDE8A4A1BC89377304F" type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
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<mods:roleTerm id="4D16D447FE178CDF9A4F4886A9B6C195">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6E56480819213A682A48A6D76F7DBFC">Rysiewska, Aleksandra</mods:namePart>
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<mods:affiliation id="42E929F71501A28904643BCF123FA410">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="F5459B5EB38AADADEC3BC861C20764FB">Palatov, Dimitry M.</mods:namePart>
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<mods:affiliation id="F0910648A780F06647C28E379A0D6C7E">Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
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</mods:name>
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<mods:name id="DEE021EA547180D8D7ED63749A162A92" type="personal">
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</mods:role>
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<mods:namePart id="68162E4F028D5297D2442022E0076122">Hofman, Sebastian</mods:namePart>
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<mods:affiliation id="96B9372BCBDCFA4940A3FB03C625711D">Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:typeOfResource id="57A4B2856D6A80E67F0F616212D02775">text</mods:typeOfResource>
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<mods:title id="BFD72B691D69E730D9D6EAB35B9BA1F8">ZooKeys</mods:title>
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<mods:part id="4594FCB87A3E470516F65AAD08FFD1B5">
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<mods:date id="AFCC4AA0E3ECC2F2FC8EBDE72BB0C964">2020</mods:date>
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<mods:detail id="6921328C5504265F4A8088E9C9FF40F9" type="volume">
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<mods:number id="4A48EE2B8CADAF89F1313A8F1CB38B49">955</mods:number>
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<mods:start id="D0C22098635607EB10098B2484C3873F">1</mods:start>
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<mods:end id="DC3F156340E0FD826BDB6F960F278D59">77</mods:end>
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<mods:url id="DBEE5A0E5A29DFF465383E550FCED1A3">http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
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<mods:classification id="A0069DA64F5636E09974B3F3C244DCF3">journal article</mods:classification>
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<mods:identifier id="6F148C27DAA5E9B3F03E73A90D9D2DA6" type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
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<mods:identifier id="D9F4F93DEFB83C7414C0F3B03938FCF8" type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
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<mods:identifier id="CCDBAF79BC9F9CD19472F779058CFBC9" type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
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<mods:identifier id="356F3B4CD31FC8AFD6EAC6AF6C4280E6" type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
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</mods:mods>
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<treatment id="1B2BA0EC1C6452D79A3D1249E5451D1B" ID-GBIF-Taxon="166096064" LSID="urn:lsid:zoobank.org:act:38835E75-8986-46B0-9040-F427F5E12A57" httpUri="http://treatment.plazi.org/id/1B2BA0EC1C6452D79A3D1249E5451D1B" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_phylum="Mollusca" scope_subFamily="Sadlerianinae" scope_superFamily="Truncatelloidea">
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<subSubSection id="FF27D31A83F8F615DB7FE5FAC3E66C2B" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="3962E0A367490762AFFD0677E6516A81" pageId="0" pageNumber="1">
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<taxonomicName id="A69FD724A8CC13D9456D70D93615F7AA" ID-CoL="7RQN8" LSID="1B2BA0EC-1C64-52D7-9A3D-1249E5451D1B" authority="Grego & Mumladze" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis nakeralaensis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nakeralaensis" status="sp. nov.">Imeretiopsis nakeralaensis Grego & Mumladze</taxonomicName>
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<taxonomicNameLabel id="65ED6285938BC9B0E682E8F027075700" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="964CB12CB5238281C88B3B2418F4F70F" captionStart="Plate 9" captionStartId="F21" captionText="Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego." figureDoi="10.3897/zookeys.955.51983.plate9" httpUri="https://binary.pensoft.net/fig/439172" pageId="0" pageNumber="1">Plate 9 (9)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="010AD02CBFF532E14B94CA31D89B2520" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="8D7EE1D8669ACECBA62DAEA51768F9CF" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="1D609103C09BE97202E85F2137C16EA3" pageId="0" pageNumber="1">
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Georgia • Imereti, Tkibuli, Tkibuli-Nikortsminda road to Nakerala Pass (ნაქერალას
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<normalizedToken id="08E62AEE5E462F0CD8270975F85D584D" originalValue="უღელტეხილი">უღელტეხილი</normalizedToken>
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);
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<geoCoordinate id="5CC8000BEAE204054F98361F19F4ABB7" degrees="42" direction="north" minutes="23" orientation="latitude" precision="15" seconds="00" value="42.383335">42°23'00"N</geoCoordinate>
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,
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<geoCoordinate id="2068C909C1170FEFF55042CED0414E63" degrees="42" direction="east" minutes="00" orientation="longitude" precision="15" seconds="45" value="42.0125">42°00'45"E</geoCoordinate>
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; 980 m a.s.l.; spring above left side of road with small travertine waterfall and a small spring cavern entrance.
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</paragraph>
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<caption id="6E09D9415E96A9EA0BF7D1356E186B5C" doi="10.3897/zookeys.955.51983.plate9" httpUri="https://binary.pensoft.net/fig/439172" pageId="0" pageNumber="1" start="Plate 9" startId="F21">
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<paragraph id="4593B6B6A276E13C8B0B04A79D7B4619" pageId="0" pageNumber="1">
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<emphasis id="DABF68DF345884CD0B5BE0C181EED9BA" bold="true" pageId="0" pageNumber="1">Plate 9.</emphasis>
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<emphasis id="FE1A8A157CB22007CCBC5DC8D2BABD64" bold="true" pageId="0" pageNumber="1">1</emphasis>
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<taxonomicName id="BCAEA5468995BB1A8C33C270F1EF6F1B" authorityName="Mumladze & Grego" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis olsavskyi" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="olsavskyi">
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<emphasis id="C00992F00AD150E01E173FF3B9EA3E67" italics="true" pageId="0" pageNumber="1">Caucasopsis olsavskyi</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype
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<emphasis id="54F66215434601AB9DEAE88DE75D7B23" bold="true" pageId="0" pageNumber="1">2</emphasis>
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<taxonomicName id="7CDCA00100DDFEF2FB5F638E61B727FD" lsidName="C. letsurtsume" pageId="0" pageNumber="1" rank="species" species="letsurtsume">
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<emphasis id="FB78DACA40DA1E3166098C8B41FE719B" italics="true" pageId="0" pageNumber="1">C. letsurtsume</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype
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<emphasis id="6D4F14C00ADA8347CD2D350B69F38321" bold="true" pageId="0" pageNumber="1">3</emphasis>
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<taxonomicName id="4CBFEB7FD0D26CB5A445333BBA5093A1" lsidName="C. letsurtsume" pageId="0" pageNumber="1" rank="species" species="letsurtsume">
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<emphasis id="C3A05F3222EF010E9DEBDD762CDE5771" italics="true" pageId="0" pageNumber="1">C. letsurtsume</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Chkhorotsku, Kachara Cave
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<emphasis id="B401186B40877459585D9E561F119D1B" bold="true" pageId="0" pageNumber="1">4</emphasis>
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<taxonomicName id="AE4796F2BC587B5C83956410FB460B8D" lsidName="C. egrisi" pageId="0" pageNumber="1" rank="species" species="egrisi">
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<emphasis id="1A070C7DD2F93B003CC38E98DEB50C88" italics="true" pageId="0" pageNumber="1">C. egrisi</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Pirveli Balda, Motena Cave form
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<emphasis id="11EB0436B46E2BC95EFA4F00689A2B77" bold="true" pageId="0" pageNumber="1">5, 6</emphasis>
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<taxonomicName id="DF831593B5AF0DD5FA15F2C1D0116C84" lsidName="C. egrisi" pageId="0" pageNumber="1" rank="species" species="egrisi">
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<emphasis id="9BADBFCAE3C2D2CD2B21D574B752B391" italics="true" pageId="0" pageNumber="1">C. egrisi</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Pirveli Balda, spring in village:
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<emphasis id="D4146B1D84F9FB35022B6441BA22AB95" bold="true" pageId="0" pageNumber="1">5</emphasis>
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holotype
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<emphasis id="EB355130166287B797A1ED42DCB0F648" bold="true" pageId="0" pageNumber="1">6</emphasis>
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paratype
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<emphasis id="3E1C57A598D19A370682C3F87E354A53" bold="true" pageId="0" pageNumber="1">7</emphasis>
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<taxonomicName id="464FDC68F2BEDBC831985A1D9CF2E1A2" authorityName="Grego & Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis prometheus" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prometheus">
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<emphasis id="80AA240BBFDB5A0ACCFF84A5EA622D69" italics="true" pageId="0" pageNumber="1">Imeretiopsis prometheus</emphasis>
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</taxonomicName>
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sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype
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<emphasis id="2D4735BAA63B7B9B7ABF17FAF838986E" bold="true" pageId="0" pageNumber="1">8</emphasis>
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<taxonomicName id="7ACDB3934E412C42A5454EC85A51E4A0" lsidName="I. cameroni" pageId="0" pageNumber="1" rank="species" species="cameroni">
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<emphasis id="F89B2553C8E10177D6E4BC2DB3B3AAC5" italics="true" pageId="0" pageNumber="1">I. cameroni</emphasis>
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</taxonomicName>
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sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype
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<emphasis id="CE710C17435E44611517EE22744A0628" bold="true" pageId="0" pageNumber="1">9</emphasis>
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<taxonomicName id="1AE93ED27C880B6267B9C7C0BC178183" lsidName="I. nakeralaensis" pageId="0" pageNumber="1" rank="species" species="nakeralaensis">
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<emphasis id="461BB98B0697C53393A8BCFE79AAD02E" italics="true" pageId="0" pageNumber="1">I. nakeralaensis</emphasis>
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</taxonomicName>
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sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype
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<emphasis id="F5003758F8240BC1C2B0B9753BBC869D" bold="true" pageId="0" pageNumber="1">10</emphasis>
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<taxonomicName id="BAA1638B80A0E1529EF26A5AD8889884" lsidName="C. letsurtsume" pageId="0" pageNumber="1" rank="species" species="letsurtsume">
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<emphasis id="2581BBD14009D34D0E351B4E15DF13AF" italics="true" pageId="0" pageNumber="1">C. letsurtsume</emphasis>
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</taxonomicName>
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sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="BDD8BEBE3B4FDEDC1459D6D366A03D9D" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="A80CC27BAE68E6701ACC8D55D6B798D4" pageId="0" pageNumber="1">Material.</paragraph>
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<paragraph id="14C458FAA961B4F016359E1DBDCD424D" pageId="0" pageNumber="1">
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<emphasis id="23426796B08683040E6CAD7E514E963B" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Georgia • 1 adult, dry; type locality; 04 May 2018; J. Grego, L. Mumladze and M.
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<normalizedToken id="DAC2229C4C9ACB8F3C91868D11A326D5" originalValue="Olšavský">Olsavsky</normalizedToken>
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leg.; ISU FM-T015-H.
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<emphasis id="1B8FDBBB61AAB2D01B5C8C7ADD265429" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Georgia • same as for holotype; ISU FM-T015-P1/10 dry, SBMNH 633110/2 dry, NHMW 113371/1 dry, NHMB 563969/2 dry, coll. JG F1009/10 dry, coll.
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<normalizedToken id="848D801D98A84A4785E814F0F1F2F194" originalValue="Glöer">Gloeer</normalizedToken>
|
|
/1 dry.
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</paragraph>
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|
</subSubSection>
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|
<subSubSection id="FBF33F85EB9FD672F9F92983EAB17B61" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="E1E41C6ACDC4E93E96E4B79C9A68E161" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="6B89C29723C413285E8671EDBB5F20C4" pageId="0" pageNumber="1">
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|
The new species differs from all the morphotypes with related shell shape in the region by its more robust oval shape, by the position of the aperture more right of the columella (to viewer; shell in apertural pose, apex up), and by the more open umbilicus.
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|
<taxonomicName id="4004F2961C375F63474206437E97B90C" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria shakuranica" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="shakuranica">
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|
<emphasis id="787639B2EE40726E7809F9F246960859" italics="true" pageId="0" pageNumber="1">Caucasogeyeria shakuranica</emphasis>
|
|
</taxonomicName>
|
|
(Starobogatov, 1962) from Abkhazia has similar but narrower shell shape with less inflated whorls and a proportionally smaller body whorl.
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|
<taxonomicName id="47E9BBF194170785600F468D83FFA5B7" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria letsurtsume" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="letsurtsume">
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|
<emphasis id="4B420A7C3795F0F63443F1C4CA10A8D8" italics="true" pageId="0" pageNumber="1">Caucasogeyeria letsurtsume</emphasis>
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|
</taxonomicName>
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|
sp. nov. has a smaller shell with more inflated whorls and more open umbilicus.
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</paragraph>
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|
</subSubSection>
|
|
<subSubSection id="0783C983A774955F3AFA73A6AB15AF75" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="48EC2F15249605FA0275938A7AA0DB7E" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="0081536A74308D09C6E98C10F2F5F467" pageId="0" pageNumber="1">
|
|
<emphasis id="14B18344AA512044D967B0896947660E" bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
: 2 mm high, elongate ovate-conical with pronounced protoconch, five tumid whorls and moderately deep suture. Shell surface whitish, translucent-glossy, covered by faint axial growth lines. Umbilicus open. Proportionally small aperture irregular, almost round, not expanded, with straight lateral and columellar labral profiles lacking sinuation. Protoconch surface densely and coarsely pitted.
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</paragraph>
|
|
<paragraph id="CB42A9883FF9220DC607DF078071AC78" pageId="0" pageNumber="1">
|
|
<emphasis id="B96F89CDCE9888B2164894695E5FBFAB" bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
<paragraph id="A6AB806D8919C8F178492E1EA95D84F6" pageId="0" pageNumber="1">
|
|
<emphasis id="911BE18173D7DAED2206F2C0D027CC15" bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
<paragraph id="A68C91F6AAACC470750D9513C96EAB02" pageId="0" pageNumber="1">
|
|
<emphasis id="5316EA0E989985DF5DA52F39565FBD7F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
|
|
: H-2.00 mm; W-1.09 mm; BH-1.02 mm; BW-0.85 mm; AH-0.64 mm; AW-0.53 mm; CA: 29°.
|
|
</paragraph>
|
|
<paragraph id="BB0BC4BC227BFEFD83E1A76E5F726748" pageId="0" pageNumber="1">
|
|
<emphasis id="29833450F5F8A9CC9B71F0D1D063D906" bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9366413D313DE8A18AF0A83A1DDF16B2" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="BCA9DB9AA042BD9F2FF75C3F78C8FBA4" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="E0AC491C9AF2AB047D371037E7F28F31" pageId="0" pageNumber="1">Name after the Nakerala Pass 1218 m alt. situated above the type locality north of Tikbuli along the road to Ambrolauri.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E79B095AC5DFC593604AC32BD98CDED3" pageId="0" pageNumber="1" type="habitat">
|
|
<paragraph id="C3C52AD189FBBED42A9014B4519B6963" pageId="0" pageNumber="1">Habitat.</paragraph>
|
|
<paragraph id="8E820E5E1D920D262566BCADDC08D73D" pageId="0" pageNumber="1">Stygobiotic species. The empty shells of the species were found at the foot of small travertine cascade formed by a small stream emerging from the very narrow cave spring (small entrance covered by moss and ivy). Only a few shells were found in sparse sediments accumulated near the cave walls. The shells were washed out from its subterranean habitat by the very small permanent stream.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B7448A1FCA829CF0EAC2ADDB629F0771" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="EC42F0C4E0A04D37D0C9517521FE5DF2" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="049F3DDFFD61C7041B6C787396B31FCC" pageId="0" pageNumber="1">Only known from the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9C208AB7942BF46553277710620A73C6" pageId="0" pageNumber="1" type="conservation status">
|
|
<paragraph id="40B7FAABECF778F014665DEED4544EFE" pageId="0" pageNumber="1">Conservation status.</paragraph>
|
|
<paragraph id="0A86813321171319E7306BB400A22E6C" pageId="0" pageNumber="1">The number of known locations (1) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="371E3EFEF859BABC3323A12FFC38FF4D" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="E911B6C3EACEA6AB2FD2D93C323E01E8" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="7D2CC5C7F5EF2028C619E5048269C335" pageId="0" pageNumber="1">
|
|
The assignment of the new species to the genus
|
|
<taxonomicName id="25CE04702E6CC2C7E12F014EC2057FB6" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="FEE31F2965EE3095B302F45D37A6834E" italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
|
|
</taxonomicName>
|
|
gen. nov. is only provisional, based on the resemblance if its shell to that of the type species. However, molecular data will be essential to confirm generic placement.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |