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<document id="0A4246F3C006502ED03C854A88B45B9C" ID="10.11646/zootaxa.4269.1.1SLASH11087" ID-CLB-Dataset="33014" ID-DOI="10.11646/zootaxa.4269.1.1" ID-GBIF-Dataset="0b7fd4db-9f34-47ef-b7e8-33363155d866" ID-ISSN="1175-5326" ID-Zenodo-Dep="581259" ID-ZooBank="2AFC0D53-B439-4BBB-BC22-A0C90C08D65D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495182755873" checkinUser="plazi" docAuthor="Lee, Chi-Feng" docDate="2017" docId="0A03A20CFFBBFF9FFF24FF25537AEDB6" docLanguage="en" docName="zootaxa.4269.1.1SLASH11087.pdf" docOrigin="Zootaxa 4269 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Doryscus wangi Lee, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="F63ADA74FF93FFB7FFB3FFB25110EB24" masterDocTitle="Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="41" updateTime="1698439733711" updateUser="ExternalLinkService">
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<mods:title id="B8CD9B104E92B906A4506F932B3617DC">Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
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<mods:name id="EFD0CC2B7ADD2A4A1D6F6FF40088E6E2" type="personal">
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<mods:namePart id="164EB772076FDC16CC629626F0638482">Lee, Chi-Feng</mods:namePart>
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<mods:title id="8F3072302363DE4D440A2F0430D5BAA7">Zootaxa</mods:title>
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<mods:part id="10C37EE3F3B338D01825CA38B23C7865">
<mods:date id="D0B9B7B3C802692489B5C7E70DDCCCB4">2017</mods:date>
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<mods:identifier id="3A4259F1BB4DDD22EB599480EDF2B2DF" type="DOI">10.11646/zootaxa.4269.1.1</mods:identifier>
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<treatment id="0A03A20CFFBBFF9FFF24FF25537AEDB6" ID-DOI="http://doi.org/10.5281/zenodo.6003769" ID-GBIF-Taxon="130810378" ID-Zenodo-Dep="6003769" LSID="urn:lsid:plazi:treatment:0A03A20CFFBBFF9FFF24FF25537AEDB6" httpUri="http://treatment.plazi.org/id/0A03A20CFFBBFF9FFF24FF25537AEDB6" lastPageNumber="41" pageId="40" pageNumber="41">
<subSubSection id="CAB04091FFBBFF9FFF24FF255066EBF7" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8215131AFFBBFF9FFF24FF255066EBF7" blockId="40.[151,441,151,211]" pageId="40" pageNumber="41">
<taxonomicName id="45AA6899FFBBFF9FFF24FF255047EB95" box="[151,343,151,177]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="wangi" status="sp. nov.">
<emphasis id="B0DECF08FFBBFF9FFF24FF255047EB95" bold="true" box="[151,343,151,177]" italics="true" pageId="40" pageNumber="41">Doryscus wangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABED7273FFBBFF9FFEECFF2A50A9EB95" box="[351,441,152,177]" pageId="40" pageNumber="41" rank="species">
<emphasis id="B0DECF08FFBBFF9FFEECFF2A50A9EB95" bold="true" box="[351,441,152,177]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="1A910F9FFFBBFF9FFF2CFF0851E4EBF7" box="[159,244,186,211]" captionStart="FIGURE 20" captionStartId="35.[151,250,1500,1522]" captionTargetBox="[155,1432,193,1478]" captionTargetId="figure@35.[155,1432,193,1478]" captionTargetPageId="35" captionText="FIGURE 20. Habitus of Doryscus species, dorsal view. 20 A. D. sumatrensis Lee, sp. nov.; 20 B. D. scapus Mohamedsaid, male; 20 C. D. scapus Mohamedsaid, female; 20 D. D. wangi Lee, sp. nov.; 20 E. D. wangi Lee, sp. nov., color variation." httpUri="https://zenodo.org/record/581279/files/figure.png" pageId="40" pageNumber="41">Figs 20</figureCitation>
D20E, 23)
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FFBBFF9FFF24FEB15324EA81" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="8215131AFFBBFF9FFF24FEB15234EA3C" blockId="40.[151,804,258,280]" box="[151,804,258,280]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="A2BA3434FFBBFF9FFF24FEB15234EA3C" box="[151,804,258,280]" pageId="40" pageNumber="41">
<taxonomicName id="45AA6899FFBBFF9FFF24FEB1537CEA3C" authority="Mohamedsaid, 1999: 124" authorityName="Mohamedsaid" authorityPageNumber="124" authorityYear="1999" box="[151,620,258,280]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FFBBFF9FFF24FEB1504AEA3C" box="[151,346,259,280]" italics="true" pageId="40" pageNumber="41">Doryscus testaceus</emphasis>
:
<treatmentCitation id="030B350BFFBBFF9FFED4FEB0537CEA3C" author="Mohamedsaid" box="[359,620,258,280]" page="124" pageId="40" pageNumber="41" year="1999">
<bibRefCitation id="E63B6EEBFFBBFF9FFED4FEB05329EA3C" author="Mohamedsaid" box="[359,569,258,280]" pageId="40" pageNumber="41" refString="Mohamedsaid, M. S. (1999) The Galerucinae from Taman Kinabalu Sabah, Malaysia (Coleoptera: Chrysomelidae). Serangga, 4, 87 - 145." type="journal article" year="1999">Mohamedsaid, 1999</bibRefCitation>
: 124
</treatmentCitation>
</taxonomicName>
. Misidentification
</treatmentCitationGroup>
</paragraph>
<paragraph id="8215131AFFBBFF9FFF24FEF75324EA81" blockId="40.[151,1437,325,1682]" pageId="40" pageNumber="41">
<materialsCitation id="32C21947FFBBFF9FFF24FEF755C5EA7A" ID-GBIF-Occurrence="1503377149" box="[151,1237,325,350]" collectingDate="2015-03-20" collectionCode="TARI" collectorName="T. Wang" country="Malaysia" location="Trusmadi" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B0DECF08FFBBFF9FFF24FEF7502AEA7A" bold="true" box="[151,314,325,350]" pageId="40" pageNumber="41">
<typeStatus id="5D11ADB8FFBBFF9FFF24FEF751CCEA7A" box="[151,220,325,350]" pageId="40" pageNumber="41">Types</typeStatus>
(n= 5).
</emphasis>
<typeStatus id="5D11ADB8FFBBFF9FFEF6FEF750BDEA7A" box="[325,429,325,350]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
<specimenCount id="94ACD893FFBBFF9FFE0BFEF450DDEA79" box="[440,461,326,349]" pageId="40" pageNumber="41" type="male"></specimenCount>
(
<collectionCode id="E4BB8BDFFFBBFF9FFE53FEF45335EA7A" box="[480,549,326,350]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="40" pageNumber="41">TARI</collectionCode>
),
<collectingCountry id="FABD538AFFBBFF9FFD8EFEF753B8EA7A" box="[573,680,325,350]" name="Malaysia" pageId="40" pageNumber="41">Malaysia</collectingCountry>
:
<collectingRegion id="406EDDF8FFBBFF9FFD05FEF753EDEA7A" box="[694,765,325,350]" country="Malaysia" name="Sabah" pageId="40" pageNumber="41">Sabah</collectingRegion>
,
<location id="877545C1FFBBFF9FFCBEFEF7526AEA7A" LSID="urn:lsid:plazi:treatment:0A03A20CFFBBFF9FFF24FF25537AEDB6:877545C1FFBBFF9FFCBEFEF7526AEA7A" box="[781,890,325,350]" country="Malaysia" name="Trusmadi" pageId="40" pageNumber="41" stateProvince="Sabah">Trusmadi</location>
,
<collectingDate id="E650CC32FFBBFF9FFC39FEF7551BEA7A" box="[906,1035,325,350]" pageId="40" pageNumber="41" value="2015-03-20">
<date id="F61435DAFFBBFF9FFC39FEF7551BEA7A" box="[906,1035,325,350]" pageId="40" pageNumber="41" value="2015-03-20">20.III.2015</date>
</collectingDate>
, leg. Y.-
<collectorName id="2F5F76CCFFBBFF9FFBC3FEF455C5EA7A" box="[1136,1237,326,350]" pageId="40" pageNumber="41">T. Wang</collectorName>
</materialsCitation>
. Paratypes: 3♀♀ (TARI), same as holotype; 1♀ (TARI), same but with “
<date id="F61435DAFFBBFF9FFD4EFEDB5269EAA5" box="[765,889,361,385]" pageId="40" pageNumber="41" value="2014-10-02">02.X.2014</date>
”;
<materialsCitation id="32C21947FFBBFF9FFC23FEDB5324EA81" ID-GBIF-Occurrence="1503377157" collectingDate="1997-10-09" collectionCode="BMNH" collectorName="K. Liston &amp; Dryobalonops" country="Malaysia" location="Danum Valley" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="94ACD893FFBBFF9FFC23FEDB52A3EAA5" box="[912,947,361,385]" pageId="40" pageNumber="41" type="female">1♀</specimenCount>
(
<collectionCode id="E4BB8BDFFFBBFF9FFC71FEDB550BEAA5" box="[962,1051,361,385]" pageId="40" pageNumber="41">BMNH</collectionCode>
),
<location id="877545C1FFBBFF9FFB9DFED855C0EAA6" LSID="urn:lsid:plazi:treatment:0A03A20CFFBBFF9FFF24FF25537AEDB6:877545C1FFBBFF9FFB9DFED855C0EAA6" box="[1070,1232,361,386]" country="Malaysia" name="Danum Valley" pageId="40" pageNumber="41" stateProvince="Sabah">Danum Valley</location>
,
<collectingDate id="E650CC32FFBBFF9FFB68FEDB5456EAA5" box="[1243,1350,361,385]" pageId="40" pageNumber="41" value="1997-10-09">
<date id="F61435DAFFBBFF9FFB68FEDB5456EAA5" box="[1243,1350,361,385]" pageId="40" pageNumber="41" value="1997-10-09">9.X.1997</date>
</collectingDate>
, leg.
<collectorName id="2F5F76CCFFBBFF9FFA32FED851F2EA82" pageId="40" pageNumber="41">K. Liston</collectorName>
, on
<emphasis id="B0DECF08FFBBFF9FFEBCFE3F533CEA82" box="[271,556,397,422]" italics="true" pageId="40" pageNumber="41">
<collectorName id="2F5F76CCFFBBFF9FFEBCFE3F50BFEA82" box="[271,431,397,422]" pageId="40" pageNumber="41">Dryobalonops</collectorName>
lanceolata
</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FFBBFF9FFF74FE0352BAEE22" pageId="40" pageNumber="41" type="description">
<paragraph id="8215131AFFBBFF9FFF74FE035245EFBE" blockId="40.[151,1437,325,1682]" pageId="40" pageNumber="41">
<emphasis id="B0DECF08FFBBFF9FFF74FE035049EAEE" bold="true" box="[199,345,433,458]" pageId="40" pageNumber="41">Description.</emphasis>
Length
<quantity id="4552BEFFFFBBFF9FFE0CFE03535DEAED" box="[447,589,433,458]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="40" pageNumber="41" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.15.1 mm</quantity>
, width
<quantity id="4552BEFFFFBBFF9FFD1AFE035228EAED" box="[681,824,433,457]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="40" pageNumber="41" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
. General color brown (
<figureCitation id="1A910F9FFFBBFF9FFBFEFE0355B6EAED" box="[1101,1190,433,458]" captionStart="FIGURE 20" captionStartId="35.[151,250,1500,1522]" captionTargetBox="[155,1432,193,1478]" captionTargetId="figure@35.[155,1432,193,1478]" captionTargetPageId="35" captionText="FIGURE 20. Habitus of Doryscus species, dorsal view. 20 A. D. sumatrensis Lee, sp. nov.; 20 B. D. scapus Mohamedsaid, male; 20 C. D. scapus Mohamedsaid, female; 20 D. D. wangi Lee, sp. nov.; 20 E. D. wangi Lee, sp. nov., color variation." httpUri="https://zenodo.org/record/581279/files/figure.png" pageId="40" pageNumber="41">Fig. 20</figureCitation>
D). Antenna filiform, antennomere I not modified in males (
<figureCitation id="1A910F9FFFBBFF9FFDF0FE675385EACA" box="[579,661,469,494]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
A), length ratios of antennomeres IXI 1.0: 0.4: 0.7: 0.8: 0.9: 0.9: 0.9: 0.9: 0.9: 0.9: 1.0, length to width ratios of antennomeres IXI 3.5: 2.2: 3.5: 4.4: 4.5: 4.5: 4.7: 4.9: 4.9: 4.9: 1.6; narrower in females (
<figureCitation id="1A910F9FFFBBFF9FFE5EFDAF532FE912" box="[493,575,541,566]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
B), length ratios of antennomeres IXI 1.0: 0.3: 0.7: 0.8: 0.9: 0.9: 0.9: 0.9: 0.8: 0.8: 1.0; length to width ratios of antennomeres IXI 3.8: 1.5: 3.2: 4.1: 4.1: 4.5: 4.8: 4.8: 4.6: 4.4: 5.0. Elytra 1.7 x longer than wide, disc with erect, brown, dense, elongate setae; and fine punctures. Apical margin of last abdominal ventrite with two well developed incisions in male (
<figureCitation id="1A910F9FFFBBFF9FFC35FD3B52C8E986" box="[902,984,649,674]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
G; widely rounded in female (
<figureCitation id="1A910F9FFFBBFF9FFA9BFD3B546AE986" box="[1320,1402,649,674]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
H). Penis (
<figureCitation id="1A910F9FFFBBFF9FFF5FFD1F505AE9E2" box="[236,330,685,710]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Figs 23</figureCitation>
C23D) extremely slender, gradually narrowed towards apex; apex of dorsal surface with moderately deep; notch of apex of ventral surface shallow; weakly curved in lateral view, apex with groove between dorsal and ventral surface; ventral surface smooth. Internal sac with median endophallic spiculae elongate, about 0.8 x as long as penis, apices lobed, basally conjoined and narrowed, with one pair of lateral processes near apex, apically tapering; one apical endophallic spiculum between two median endophalic spiculae, apex truncate and basally tapering, curved downwards; with one curved process at basal 1/
<quantity id="4552BEFFFFBBFF9FFC45FCD05531E85E" box="[1014,1057,865,890]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="40" pageNumber="41" unit="in" value="3.0">3 in</quantity>
lateral view; three pairs of dorsal spiculae at basal 1/3 of median endophallic spiculae, inner pair longest and straight, outer pair shortest and hornlike; one pair of ventral endophallic spiculae long, separated, apices bifurcate. Gonocoxae (
<figureCitation id="1A910F9FFFBBFF9FFB06FC1B5418E8E6" box="[1205,1288,937,962]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
E) extremely slender, 6.8 x longer than wide, sides slightly narrowed at apical 1/10; apices narrowed rounded; inner margins narrowly approximate from apex to apical 2/5, apex with nine setae; extremely slender from midpoint to base but abruptly widened at base. Ventrite VIII (
<figureCitation id="1A910F9FFFBBFF9FFDECFBA753A0EF0A" box="[607,688,1045,1070]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
F) well sclerotized; spiculum elongate; apex square, apical margin truncate, with dense short setae; disc with several long setae at sides. Receptacle of spermatheca (
<figureCitation id="1A910F9FFFBBFF9FFB64FB8B5438EF76" box="[1239,1320,1081,1106]" captionStart="FIGURE 23" captionStartId="39.[151,250,1828,1850]" captionTargetBox="[161,1426,205,1798]" captionTargetId="figure@39.[151,1436,193,1807]" captionTargetPageId="39" captionText="FIGURE 23. Diagnostic characters of Doryscus wangi Lee sp. nov. 23 A. Antenna, male; 23 B. Antenna, female; 23 C. Penis, dorsal view; 23 D. Penis, lateral view; 23 E. Gonocoxae; 23 F. Abdominal ventrite VIII. 23 G. Apical margin of abdominal ventrite V, male; 23 H. Apical margin of abdominal ventrite V, female; 23 I. Spermatheca." httpUri="https://zenodo.org/record/581282/files/figure.png" pageId="40" pageNumber="41">
Fig.
<date id="F61435DAFFBBFF9FFAB8FB885438EF76" box="[1291,1320,1082,1106]" pageId="40" pageNumber="41" value="1923-01">23</date>
</figureCitation>
I) strongly swollen; pump much narrower and strongly curved, apically tapering; proximal spermathecal duct wide but much narrower than receptacle, deeply projecting into receptaculum.
</paragraph>
<paragraph id="8215131AFFBBFF9FFF74FB1752BAEE22" blockId="40.[151,1437,325,1682]" pageId="40" pageNumber="41">
<emphasis id="B0DECF08FFBBFF9FFF74FB175084EF9A" bold="true" box="[199,404,1189,1214]" pageId="40" pageNumber="41">Color variability.</emphasis>
One female specimen possesses dark brown frons and anterior area of vertex. Another has a yellowish brown body but outer margins of the pronotum, outer margins and suture of the elytra are black, and the head is black but the labium, clypeus, and frons are yellow (
<figureCitation id="1A910F9FFFBBFF9FFC8BFB5F529BEE22" box="[824,907,1261,1286]" captionStart="FIGURE 20" captionStartId="35.[151,250,1500,1522]" captionTargetBox="[155,1432,193,1478]" captionTargetId="figure@35.[155,1432,193,1478]" captionTargetPageId="35" captionText="FIGURE 20. Habitus of Doryscus species, dorsal view. 20 A. D. sumatrensis Lee, sp. nov.; 20 B. D. scapus Mohamedsaid, male; 20 C. D. scapus Mohamedsaid, female; 20 D. D. wangi Lee, sp. nov.; 20 E. D. wangi Lee, sp. nov., color variation." httpUri="https://zenodo.org/record/581279/files/figure.png" pageId="40" pageNumber="41">Fig. 20</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FFBBFF9FFF74FAA353FBED25" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8215131AFFBBFF9FFF74FAA353FBED25" blockId="40.[151,1437,325,1682]" pageId="40" pageNumber="41">
<emphasis id="B0DECF08FFBBFF9FFF74FAA350DEEE0E" bold="true" box="[199,462,1297,1322]" pageId="40" pageNumber="41">Differential diagnosis.</emphasis>
Males of
<taxonomicName id="45AA6899FFBBFF9FFD8CFAA053E4EE0D" box="[575,756,1298,1321]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="wangi" status="sp. nov.">
<emphasis id="B0DECF08FFBBFF9FFD8CFAA053E4EE0D" box="[575,756,1298,1321]" italics="true" pageId="40" pageNumber="41">Doryscus wangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABED7273FFBBFF9FFD4FFAA35246EE0E" box="[764,854,1297,1322]" pageId="40" pageNumber="41" rank="species">
<emphasis id="B0DECF08FFBBFF9FFD4FFAA35246EE0E" bold="true" box="[764,854,1297,1322]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
are easily distinguished from those of
<taxonomicName id="45AA6899FFBBFF9FFAB9FAA05469EE0D" box="[1290,1401,1298,1321]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="scapus">
<emphasis id="B0DECF08FFBBFF9FFAB9FAA05469EE0D" box="[1290,1401,1298,1321]" italics="true" pageId="40" pageNumber="41">D. scapus</emphasis>
</taxonomicName>
by the normal first antennomeres (strongly swollen first antennomeres in
<taxonomicName id="45AA6899FFBBFF9FFC70FA845524EE69" box="[963,1076,1334,1357]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="scapus">
<emphasis id="B0DECF08FFBBFF9FFC70FA845524EE69" box="[963,1076,1334,1357]" italics="true" pageId="40" pageNumber="41">D. scapus</emphasis>
</taxonomicName>
) and characteristic structure of endophallic sclerites (
<figureCitation id="1A910F9FFFBBFF9FFE3EFAEB50F4EE56" box="[397,484,1369,1394]" captionStart="FIGURE 22" captionStartId="38.[151,250,1828,1850]" captionTargetBox="[160,1420,278,1803]" captionTargetId="figure@38.[151,1436,267,1806]" captionTargetPageId="38" captionText="FIGURE 22. Diagnostic characters of Doryscus scapus Mohamedsaid. 22 A. Antenna, male; 22 B. Antenna, female; 22 C. Penis, dorsal view; 21 D. Penis, lateral view; 22 E. Gonocoxae; 22 F. Abdominal ventrite VIII. 22 G. Apical margin of abdominal ventrite V, male; 22 H. Apical margin of abdominal ventrite V, female; 22 I. Spermatheca." httpUri="https://zenodo.org/record/581281/files/figure.png" pageId="40" pageNumber="41">Figs 22</figureCitation>
C22D, 23C23D). Females of
<taxonomicName id="45AA6899FFBBFF9FFCFFFAE852A2EE55" box="[844,946,1370,1393]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="wangi" status="sp. nov.">
<emphasis id="B0DECF08FFBBFF9FFCFFFAE852A2EE55" box="[844,946,1370,1393]" italics="true" pageId="40" pageNumber="41">D. wangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABED7273FFBBFF9FFC08FAEB5506EE56" box="[955,1046,1369,1394]" pageId="40" pageNumber="41" rank="species">
<emphasis id="B0DECF08FFBBFF9FFC08FAEB5506EE56" bold="true" box="[955,1046,1369,1394]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated from those of
<taxonomicName id="45AA6899FFBBFF9FFA32FAE851F3EEB1" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="scapus">
<emphasis id="B0DECF08FFBBFF9FFA32FAE851F3EEB1" italics="true" pageId="40" pageNumber="41">D. scapus</emphasis>
</taxonomicName>
by lacking transverse dark bands on elytral bases, swollen receptacle and wide proximal spermathecal duct of the spermatheca, and the comparatively shorter first antennomere (only slightly longer than others). These are in contrast to the narrow receptacle and slender proximal spermathecal duct, and the comparatively longer first antennomere (much longer than others) of
<taxonomicName id="45AA6899FFBBFF9FFDC5FA5853F4ED25" box="[630,740,1514,1537]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="scapus">
<emphasis id="B0DECF08FFBBFF9FFDC5FA5853F4ED25" box="[630,740,1514,1537]" italics="true" pageId="40" pageNumber="41">D. scapus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FFBBFF9FFF74F9BF5232ED4A" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8215131AFFBBFF9FFF74F9BF55E2ED02" blockId="40.[151,1437,325,1682]" box="[199,1266,1549,1574]" pageId="40" pageNumber="41">
<emphasis id="B0DECF08FFBBFF9FFF74F9BF505EED02" bold="true" box="[199,334,1549,1574]" pageId="40" pageNumber="41">Etymology.</emphasis>
This new species is dedicated to Yu-Tang Wang who collected the
<typeStatus id="5D11ADB8FFBBFF9FFB8CF9BC557FED02" box="[1087,1135,1550,1574]" pageId="40" pageNumber="41">type</typeStatus>
specimens.
</paragraph>
<paragraph id="8215131AFFBBFF9FFF74F9835232ED4A" blockId="40.[151,1437,325,1682]" pageId="40" pageNumber="41">
Remarks. Males of East
<collectingCountry id="FABD538AFFBBFF9FFE59F9835340ED6E" box="[490,592,1585,1610]" name="Malaysia" pageId="40" pageNumber="41">Malaysia</collectingCountry>
with normal antennae identified as
<taxonomicName id="45AA6899FFBBFF9FFC48F98055C7ED6D" box="[1019,1239,1586,1609]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FFBBFF9FFC48F98055C7ED6D" box="[1019,1239,1586,1609]" italics="true" pageId="40" pageNumber="41">Doryscus testaceus</emphasis>
</taxonomicName>
(not studied) by
<bibRefCitation id="E63B6EEBFFBBFF9FFF24F9E75097ED4A" author="Mohamedsaid" box="[151,391,1621,1646]" pageId="40" pageNumber="41" refString="Mohamedsaid, M. S. (1999) The Galerucinae from Taman Kinabalu Sabah, Malaysia (Coleoptera: Chrysomelidae). Serangga, 4, 87 - 145." type="journal article" year="1999">Mohamedsaid (1999)</bibRefCitation>
are supposed to be this new species.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FFBBFF9FFF74F9CB537AEDB6" box="[199,618,1657,1682]" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8215131AFFBBFF9FFF74F9CB537AEDB6" blockId="40.[151,1437,325,1682]" box="[199,618,1657,1682]" pageId="40" pageNumber="41">
<emphasis id="B0DECF08FFBBFF9FFF74F9CB5070EDB6" bold="true" box="[199,352,1657,1682]" pageId="40" pageNumber="41">Distribution.</emphasis>
East
<collectingCountry id="FABD538AFFBBFF9FFE2DF9CB5315EDB6" box="[414,517,1657,1682]" name="Malaysia" pageId="40" pageNumber="41">Malaysia</collectingCountry>
(
<collectingRegion id="406EDDF8FFBBFF9FFDA7F9CB534EEDB6" box="[532,606,1657,1682]" country="Malaysia" name="Sabah" pageId="40" pageNumber="41">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>