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<document id="83129E9CA2224DD529D334B086F9B2D8" ID-CLB-Dataset="6233" ID-DOI="10.11646/zootaxa.4974.1.7" ID-GBIF-Dataset="59461c15-34d6-454e-9613-e03fe6332d23" ID-ISSN="1175-5326" ID-Zenodo-Dep="4772319" ID-ZooBank="DB053133-4F39-4E2C-B918-54DFD0B11D6A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621416828918" checkinUser="plazi" docAuthor="Heppner, John B., Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2021" docId="03F98C68FF93063959C3296DFAA4F8BE" docLanguage="en" docName="zootaxa.4974.1.7.pdf" docOrigin="Zootaxa 4974 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tisis siantara Heppner &amp; Park &amp; Bae 2021, new species" docType="treatment" docUuid="7833A61B-E91A-465A-96B1-507E6C0DEC56" docUuidSource="ZooBank" docVersion="9" lastPageNumber="167" masterDocId="FFC0F410FF92063B59542802FFCCFFE2" masterDocTitle="A new species of Tisis Walker from North Sumatra, Indonesia (Lepidoptera Lecithoceridae)" masterLastPageNumber="168" masterPageNumber="165" pageNumber="166" updateTime="1698940085860" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9EBE0E3CF925BFA6ED16265A0DA8F65F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0ABE7B7769872BAD355AE7CEA6B41814">
<mods:title id="E9D4725A9FDF66728955E805821EA08A">A new species of Tisis Walker from North Sumatra, Indonesia (Lepidoptera Lecithoceridae)</mods:title>
</mods:titleInfo>
<mods:name id="24E78278451346DC3AEF6B2DA11FC3B9" type="personal">
<mods:role id="F6611C93BFBD62B072CF3A56027A696E">
<mods:roleTerm id="D8B61389397C461BB6D4DFD931372D74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A65D11FD6872A8B9144C181D21B16CEF">Heppner, John B.</mods:namePart>
<mods:affiliation id="78C7448B8E9A5A918A3CB0950E45F296">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA. &amp; jheppner @ flmnh. ufl. edu</mods:affiliation>
</mods:name>
<mods:name id="3B1BEE80F9C5A3E9EAF8A921320797BF" type="personal">
<mods:role id="F4C39D59105E9AF62FD30E30D54D3C6E">
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</mods:role>
<mods:namePart id="F8A22175B69B9EED653C6FC07298A723">Park, Kyu-Tek</mods:namePart>
<mods:nameIdentifier id="6FEF810DBF9007D6DE55FE14297A9808" type="ORCID">0000-0001-9933-4497</mods:nameIdentifier>
<mods:affiliation id="AEAC585110A1F3AE6E81D7C13849D20F">Bio ˗ Resource and Environmental Center, Incheon National University, Songdo ˗ dong, Incheon 22012, South Korea. &amp; ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
<mods:nameIdentifier id="35AC5F36464F9856C5201CB87E54B64B" type="email">ktpark02@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3929454711AE42EA5878DF180F0D02A3" type="personal">
<mods:role id="341DF0DD2A6081982358CA4A6C69FD75">
<mods:roleTerm id="38C2802A1B3544FBDF1FC6FC6DE082CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="137798A488A8AE349A092E8BCF046BA4">Bae, Yang-Seop</mods:namePart>
<mods:affiliation id="F49CDF8561F5D755DCE812BE05DCA3DA">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, South Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, Songdo ˗ dong, Incheon 22012, South Korea.</mods:affiliation>
</mods:name>
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<mods:title id="429460259CFA68A0025EC7A763F84BFA">Zootaxa</mods:title>
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<mods:part id="DDFDAF5E5A73839E4159BD05B2A314C5">
<mods:date id="4818EF0BA4B2559D82402C96D693A816">2021</mods:date>
<mods:detail id="285FF94B28AB1D7B702EC8FE6A8B06E7" type="pubDate">
<mods:number id="7CC85D97A688E9B696F367C26F88E49A">2021-05-19</mods:number>
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<mods:number id="1B3C04AFBD28D7B9100EA41900DE578D">4974</mods:number>
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<mods:identifier id="B66F55435D1DB24349371F532598A317" type="DOI">10.11646/zootaxa.4974.1.7</mods:identifier>
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<treatment id="03F98C68FF93063959C3296DFAA4F8BE" ID-DOI="http://doi.org/10.5281/zenodo.4917246" ID-GBIF-Taxon="182641401" ID-Zenodo-Dep="4917246" LSID="urn:lsid:zoobank.org:act:7833A61B-E91A-465A-96B1-507E6C0DEC56" httpUri="http://treatment.plazi.org/id/03F98C68FF93063959C3296DFAA4F8BE" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166">
<subSubSection id="C34A6EF5FF93063A59C3296DFD7FFE6A" box="[151,691,366,393]" pageId="1" pageNumber="166" type="nomenclature">
<paragraph id="8BEF3D7EFF93063A59C3296DFD7FFE6A" blockId="1.[151,691,366,465]" box="[151,691,366,393]" pageId="1" pageNumber="166">
<heading id="D0A78A12FF93063A59C3296DFD7FFE6A" bold="true" box="[151,691,366,393]" fontSize="11" level="1" pageId="1" pageNumber="166" reason="1">
<emphasis id="B924E16CFF93063A59C3296DFD7FFE6A" bold="true" box="[151,691,366,393]" pageId="1" pageNumber="166">
<taxonomicName id="4C5046FDFF93063A59C3296DFDDBFE6B" ID-CoL="9FPVJ" authority="Heppner &amp; Park" authorityName="Heppner &amp; Park &amp; Bae" authorityYear="2021" box="[151,535,366,393]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="siantara" status="sp. nov.">
<emphasis id="B924E16CFF93063A59C3296DFEF5FE6B" bold="true" box="[151,313,367,393]" italics="true" pageId="1" pageNumber="166">Tisis siantara</emphasis>
Heppner &amp; Park
</taxonomicName>
,
<taxonomicNameLabel id="A2175C17FF93063A5B77296DFD7FFE6A" box="[547,691,366,393]" pageId="1" pageNumber="166" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF93063A59C32991FF19FE33" pageId="1" pageNumber="166" type="description">
<paragraph id="8BEF3D7EFF93063A59C32991FF27FE4F" blockId="1.[151,691,366,465]" box="[151,235,403,429]" pageId="1" pageNumber="166">
(
<figureCitation id="136B21FBFF93063A59F42991FF2CFE4F" box="[160,224,403,429]" captionStart="FIGURE 1" captionStartId="1.[151,260,1997,2023]" captionTargetBox="[317,1264,1530,1972]" captionTargetId="figure-541@1.[317,1271,1530,1972]" captionTargetPageId="1" captionText="FIGURE 1. Tisis siantara n. sp., male holotype (25mm wingspan) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.4772321" httpUri="https://zenodo.org/record/4772321/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BEF3D7EFF93063A59C329B5FF19FE33" blockId="1.[151,691,366,465]" box="[151,213,439,465]" pageId="1" pageNumber="166">LSID</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF93063A59C329FDFBBBFDDF" pageId="1" pageNumber="166" type="materials_examined">
<paragraph id="8BEF3D7EFF93063A59C329FDFBBBFDDF" blockId="1.[151,1437,510,1510]" pageId="1" pageNumber="166">
<emphasis id="B924E16CFF93063A59C329FDFE89FDFA" bold="true" box="[151,325,510,537]" pageId="1" pageNumber="166">Type material.</emphasis>
<materialsCitation id="3B383723FF93063A581829FDFBBBFDDF" ID-GBIF-Occurrence="3128175301" collectingDate="1992-08-22" collectionCode="MGCL" collectorName="J. B. Heppner &amp; E. W. Diehl " country="Indonesia" county="Utara" elevation="1" location="Pematang Siantar" municipality="Medan Prov." pageId="1" pageNumber="166" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B924E16CFF93063A581829FDFE0CFDFB" bold="true" box="[332,448,511,537]" pageId="1" pageNumber="166">
<typeStatus id="54EB83DCFF93063A581829FDFE77FDFB" box="[332,443,511,537]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
.
</emphasis>
<quantity id="4CA8909BFF93063A589329FDFE3FFDFB" box="[455,499,511,537]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="166" unit="m" value="1.0">
<elevation id="007DDA4DFF93063A589329FDFE3FFDFB" box="[455,499,511,537]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="166" unit="m" value="1.0">1 m</elevation>
</quantity>
#,
<quantity id="4CA8909BFF93063A5B5A29FDFD82FDFB" box="[526,590,511,537]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="166" unit="km" value="28.0">28km</quantity>
SW.
<location id="8E8F6BA5FF93063A5BDF29FDFC83FDFB" LSID="urn:lsid:plazi:treatment:03F98C68FF93063959C3296DFAA4F8BE:8E8F6BA5FF93063A5BDF29FDFC83FDFB" box="[651,847,511,537]" country="Indonesia" county="Utara" municipality="Medan Prov." name="Pematang Siantar" pageId="1" pageNumber="166" stateProvince="Sumatera Utara">Pematang Siantar</location>
(
<elevation id="007DDA4DFF93063A5A0B29FCFC68FDFB" box="[863,932,510,537]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="166" unit="m" value="950.0">950m</elevation>
),
<collectingMunicipality id="6B8BA704FF93063A5AE329FDFB86FDFB" box="[951,1098,511,537]" pageId="1" pageNumber="166">Medan Prov.</collectingMunicipality>
,
<collectingRegion id="4994F39CFF93063A5D0029FDFB7DFDFB" box="[1108,1201,511,537]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="166">Sumatra</collectingRegion>
<collectingCounty id="628E45F2FF93063A5DED29FDFB36FDFB" box="[1209,1274,511,537]" pageId="1" pageNumber="166">Utara</collectingCounty>
,
<collectingCountry id="F3477DEEFF93063A5C5129FDFAB9FDFB" box="[1285,1397,511,537]" name="Indonesia" pageId="1" pageNumber="166">Indonesia</collectingCountry>
,
<date id="FFEE1BBEFF93063A5CD429FDFEC5FDDF" pageId="1" pageNumber="166" value="1992-08-22">
<collectingDate id="EFAAE256FF93063A5CD429FDFEC5FDDF" pageId="1" pageNumber="166" value="1992-08-22">22 Aug 1992</collectingDate>
</date>
,
<collectorName id="26A558A8FF93063A58402A21FE62FDDF" box="[276,430,547,573]" pageId="1" pageNumber="166">J. B. Heppner</collectorName>
&amp;
<collectorName id="26A558A8FF93063A58862A21FD99FDDF" box="[466,597,547,573]" pageId="1" pageNumber="166">E. W. Diehl</collectorName>
(genitalia slide JBH 3094). Deposited in
<collectionCode id="ED41A5BBFF93063A5D722A21FBBBFDDF" box="[1062,1143,547,573]" pageId="1" pageNumber="166">MGCL</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF93063A59932A44FECCFD13" pageId="1" pageNumber="166" type="diagnosis">
<paragraph id="8BEF3D7EFF93063A59932A44FECCFD13" blockId="1.[151,1437,510,1510]" pageId="1" pageNumber="166">
<emphasis id="B924E16CFF93063A59932A44FE8EFD82" bold="true" box="[199,322,582,608]" pageId="1" pageNumber="166">Diagnosis.</emphasis>
Diagnostic characters for this new species involve the male genitalia: the male valvae are more distended with a straighter sacculus (very rounded sacculus in
<taxonomicName id="4C5046FDFF93063A5A352A69FBDEFD66" authorityName="Park" authorityYear="2007" box="[865,1042,619,644]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF93063A5A352A69FBDEFD66" box="[865,1042,619,644]" italics="true" pageId="1" pageNumber="166">T. sumatraensis</emphasis>
</taxonomicName>
), and the caudal lobe of juxta are narrow and with small bases (wider and stouter and with large triangular bases in
<taxonomicName id="4C5046FDFF93063A5D6C2A8DFB24FD4A" authorityName="Park" authorityYear="2007" box="[1080,1256,655,680]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF93063A5D6C2A8DFB24FD4A" box="[1080,1256,655,680]" italics="true" pageId="1" pageNumber="166">T. sumatraensis</emphasis>
</taxonomicName>
). The hindwing of the new species has the dark linear line curved and near the dorsal wing margin (straight and medial in
<taxonomicName id="4C5046FDFF93063A5C682AB1FF3CFD12" authorityName="Park" authorityYear="2007" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF93063A5C682AB1FF3CFD12" italics="true" pageId="1" pageNumber="166">T. sumatraensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF93063A59932AF8FB7FFA07" pageId="1" pageNumber="166" type="description">
<paragraph id="8BEF3D7EFF93063A59932AF8FB7FFA07" blockId="1.[151,1437,510,1510]" pageId="1" pageNumber="166">
<emphasis id="B924E16CFF93063A59932AF8FE95FCF6" bold="true" box="[199,345,762,788]" pageId="1" pageNumber="166">Description.</emphasis>
Wingspan
<quantity id="4CA8909BFF93063A588C2AF8FDEFFCF7" box="[472,547,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="166" unit="mm" value="25.0">25mm</quantity>
. Male (
<figureCitation id="136B21FBFF93063A5B2C2AF9FD77FCF7" box="[632,699,763,789]" captionStart="FIGURE 1" captionStartId="1.[151,260,1997,2023]" captionTargetBox="[317,1264,1530,1972]" captionTargetId="figure-541@1.[317,1271,1530,1972]" captionTargetPageId="1" captionText="FIGURE 1. Tisis siantara n. sp., male holotype (25mm wingspan) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.4772321" httpUri="https://zenodo.org/record/4772321/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
).
<emphasis id="B924E16CFF93063A5B802AF9FCDDFCF7" box="[724,785,763,789]" italics="true" pageId="1" pageNumber="166">Head</emphasis>
(
<figureCitation id="136B21FBFF93063A5A742AF9FC44FCF7" box="[800,904,763,789]" captionStart="FIGURES 2–4" captionStartId="2.[151,275,513,539]" captionTargetBox="[155,1436,179,489]" captionTargetId="figure-25@2.[151,1436,179,489]" captionTargetPageId="2" captionText="FIGURES 2–4. Tisis siantara n. sp., male holotype, head close-ups, lateral (2), dorsal (3), and frontal (4) views." figureDoi="http://doi.org/10.5281/zenodo.4772324" httpUri="https://zenodo.org/record/4772324/files/figure.png" pageId="1" pageNumber="166">Figs 2–4</figureCitation>
): Yellowish brown with light orange bristles of chaetosema (chaetosema base black); antenna yellowish brown with wide basal segments brown (more so dorsolaterally), basal joints of flagellum broadly dilated with dark-brown hairs; second segment of labial palpus; long, with long, yellowish-brown scale tufts ventrally in basal 3/5, yellowish brown beyond.
<emphasis id="B924E16CFF93063A5D212B65FB0AFC63" box="[1141,1222,871,897]" italics="true" pageId="1" pageNumber="166">Thorax</emphasis>
: Yellowish brown; tegula golden brown; patagia yellowish brown with long setae; legs yellowish brown with golden setal ends on long scale tufts; hindlegs white-tan.
<emphasis id="B924E16CFF93063A58AE2BADFDAAFC2A" box="[506,614,943,968]" italics="true" pageId="1" pageNumber="166">Forewing</emphasis>
: Dark brown, with median elongated yellow patch and yellow on costa; thin line at base and to costal 1/3 beyond midwing; wing color mixed with some pale scales on brown areas; fringe golden brown; venter pale yellowish brown, with fringe of light orange-brown from tornus and along costal margin.
<emphasis id="B924E16CFF93063A59C32C19FEC9FBD7" box="[151,261,1051,1077]" italics="true" pageId="1" pageNumber="166">Hindwing</emphasis>
: Dark brown linear marking from base to tornus on dorsal portion of wing, curved to apex, margined with golden-yellow medially and on costa, with large hyaline patch centrally; some black scales mixed in near termen; fringe yellow to golden, becoming brown on anal margin; venter pale brown, with yellow-tan on costa and costal fringe, to yellow fringe on termen to bronze-brown on anal margin.
<emphasis id="B924E16CFF93063A5AB12C85FB83FB43" box="[997,1103,1159,1185]" italics="true" pageId="1" pageNumber="166">Abdomen</emphasis>
: Bronze-tan, with lateral line of dark brown; venter white-tan; genital tufts yellow-tan.
<emphasis id="B924E16CFF93063A5A4E2CA9FC73FB27" box="[794,959,1195,1221]" italics="true" pageId="1" pageNumber="166">Male genitalia</emphasis>
(
<figureCitation id="136B21FBFF93063A5A992CA9FBDEFB27" box="[973,1042,1194,1221]" captionStart="FIGURE 5" captionStartId="2.[151,260,1570,1596]" captionTargetBox="[163,1436,562,1546]" captionTargetId="figure-47@2.[151,1436,561,1546]" captionTargetPageId="2" captionText="FIGURE 5. Tisis siantara n. sp.: male genitalia, with details A) aedeagus, B) central genital close-up (slide JBH 3094)." figureDoi="http://doi.org/10.5281/zenodo.4772326" httpUri="https://zenodo.org/record/4772326/files/figure.png" pageId="1" pageNumber="166">Fig. 5</figureCitation>
): Uncus short, pyramidal; tegumen quadratic with rounded dorsal corners; gnathos recurved to acute claw-like point directed downwards; vinculum broadly rounded, with round undifferentiated saccus; juxta a small rounded trapezoidal plate, with long, slender caudal processes meeting centrally as upward approximate stout sinuate projections; valva elongate, with somewhat straight dorsal margin to upcurved diversion midway to rounded cucullus-like termen, acute dorsally, with strong setal-spine margin distally, an inward tooth-like inwardly directed projection (or absent); saccular margin slightly curved to emargination before cucullus, with strong ventral margin basal to median semi-circular cavity, and with a slight dentate or carinate ventral evagination before base; aedeagus elongate and relatively narrow (1/7 as wide as length), with short bulbous phallobase; cornutus as wedge-like form with a small apical hook.
</paragraph>
</subSubSection>
<caption id="DF2F6DF6FF93063A59C32FCFFC33F805" ID-DOI="http://doi.org/10.5281/zenodo.4772321" ID-Zenodo-Dep="4772321" box="[151,1023,1997,2024]" httpUri="https://zenodo.org/record/4772321/files/figure.png" pageId="1" pageNumber="166" startId="1.[151,260,1997,2023]" targetBox="[317,1264,1530,1972]" targetPageId="1">
<paragraph id="8BEF3D7EFF93063A59C32FCFFC33F805" blockId="1.[151,1023,1997,2024]" box="[151,1023,1997,2024]" pageId="1" pageNumber="166">
<emphasis id="B924E16CFF93063A59C32FCFFEEDF805" bold="true" box="[151,289,1997,2023]" pageId="1" pageNumber="166">FIGURE 1.</emphasis>
<taxonomicName id="4C5046FDFF93063A58732FCCFE72F805" authority="Heppner &amp; Park &amp; Bae, 2021" authorityName="Heppner &amp; Park &amp; Bae" authorityYear="2021" box="[295,446,1998,2023]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="siantara" status="sp. nov.">
<emphasis id="B924E16CFF93063A58732FCCFE72F805" box="[295,446,1998,2023]" italics="true" pageId="1" pageNumber="166">Tisis siantara</emphasis>
</taxonomicName>
<emphasis id="B924E16CFF93063A58992FCFFDC7F805" bold="true" box="[461,523,1997,2023]" pageId="1" pageNumber="166">
<taxonomicNameLabel id="A2175C17FF93063A58992FCFFDC7F805" box="[461,523,1997,2023]" pageId="1" pageNumber="166" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (25mm wingspan) (MGCL).
</paragraph>
</caption>
<caption id="DF2F6DF6FF90063959C32A03FA42FDF9" ID-DOI="http://doi.org/10.5281/zenodo.4772324" ID-Zenodo-Dep="4772324" box="[151,1422,513,539]" httpUri="https://zenodo.org/record/4772324/files/figure.png" pageId="2" pageNumber="167" startId="2.[151,275,513,539]" targetBox="[155,1436,179,489]" targetPageId="2">
<paragraph id="8BEF3D7EFF90063959C32A03FA42FDF9" blockId="2.[151,1422,513,539]" box="[151,1422,513,539]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959C32A03FE87FDF9" bold="true" box="[151,331,513,539]" pageId="2" pageNumber="167">FIGURES 2–4.</emphasis>
<taxonomicName id="4C5046FDFF90063958072A00FE26FDF9" authority="Heppner &amp; Park &amp; Bae, 2021" authorityName="Heppner &amp; Park &amp; Bae" authorityYear="2021" box="[339,490,514,539]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="siantara" status="sp. nov.">
<emphasis id="B924E16CFF90063958072A00FE26FDF9" box="[339,490,514,539]" italics="true" pageId="2" pageNumber="167">Tisis siantara</emphasis>
</taxonomicName>
<emphasis id="B924E16CFF90063958AC2A03FDFBFDF9" bold="true" box="[504,567,513,539]" pageId="2" pageNumber="167">
<taxonomicNameLabel id="A2175C17FF90063958AC2A03FDFBFDF9" box="[504,567,513,539]" pageId="2" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype, head close-ups, lateral (2), dorsal (3), and frontal (4) views.
</paragraph>
</caption>
<caption id="DF2F6DF6FF90063959C32E20FF2CF982" ID-DOI="http://doi.org/10.5281/zenodo.4772326" ID-Zenodo-Dep="4772326" httpUri="https://zenodo.org/record/4772326/files/figure.png" pageId="2" pageNumber="167" startId="2.[151,260,1570,1596]" targetBox="[163,1436,562,1546]" targetPageId="2">
<paragraph id="8BEF3D7EFF90063959C32E20FF2CF982" blockId="2.[151,1436,1570,1632]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959C32E20FEEDF9DE" bold="true" box="[151,289,1570,1596]" pageId="2" pageNumber="167">FIGURE 5.</emphasis>
<taxonomicName id="4C5046FDFF900639587D2E21FE0DF9DE" authority="Heppner &amp; Park &amp; Bae, 2021" authorityName="Heppner &amp; Park &amp; Bae" authorityYear="2021" box="[297,449,1571,1596]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="siantara" status="n . sp.">
<emphasis id="B924E16CFF900639587D2E21FE0DF9DE" box="[297,449,1571,1596]" italics="true" pageId="2" pageNumber="167">Tisis siantara</emphasis>
</taxonomicName>
<emphasis id="B924E16CFF900639589D2E20FDDDF9DE" bold="true" box="[457,529,1570,1596]" pageId="2" pageNumber="167">
<taxonomicNameLabel id="A2175C17FF900639589D2E20FDDDF9DE" box="[457,529,1570,1596]" pageId="2" pageNumber="167">n. sp.</taxonomicNameLabel>
</emphasis>
: male genitalia, with details A) aedeagus, B) central genital close-up (slide JBH 3094).
</paragraph>
</caption>
<subSubSection id="C34A6EF5FF90063959932E8CFE6AF94A" box="[199,422,1678,1705]" pageId="2" pageNumber="167" type="biology_ecology">
<paragraph id="8BEF3D7EFF90063959932E8CFE6AF94A" blockId="2.[151,1437,1678,1885]" box="[199,422,1678,1705]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959932E8CFEEDF94A" bold="true" box="[199,289,1678,1704]" pageId="2" pageNumber="167">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF90063959932EB0FC28F92F" box="[199,996,1714,1741]" pageId="2" pageNumber="167" type="distribution">
<paragraph id="8BEF3D7EFF90063959932EB0FC28F92F" blockId="2.[151,1437,1678,1885]" box="[199,996,1714,1741]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959932EB0FE95F92E" bold="true" box="[199,345,1714,1740]" pageId="2" pageNumber="167">Distribution</emphasis>
. Known thus far only from northern
<collectingRegion id="4994F39CFF9006395BAD2EB0FC9AF92E" box="[761,854,1714,1740]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="167">Sumatra</collectingRegion>
(
<collectingCountry id="F3477DEEFF9006395A312EB0FC15F92F" box="[869,985,1714,1741]" name="Indonesia" pageId="2" pageNumber="167">Indonesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF90063959932ED4FBFCF912" box="[199,1072,1750,1777]" pageId="2" pageNumber="167" type="etymology">
<paragraph id="8BEF3D7EFF90063959932ED4FBFCF912" blockId="2.[151,1437,1678,1885]" box="[199,1072,1750,1777]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959932ED4FE84F912" bold="true" box="[199,328,1750,1776]" pageId="2" pageNumber="167">Etymology</emphasis>
. Named after the town (Siantar) near the type-locality in
<collectingRegion id="4994F39CFF9006395A9F2ED4FBE0F912" box="[971,1068,1750,1776]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="167">Sumatra</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6EF5FF90063959932EF8FAA4F8BE" pageId="2" pageNumber="167" type="discussion">
<paragraph id="8BEF3D7EFF90063959932EF8FAA4F8BE" blockId="2.[151,1437,1678,1885]" pageId="2" pageNumber="167">
<emphasis id="B924E16CFF90063959932EF8FEF7F8F6" bold="true" box="[199,315,1786,1812]" pageId="2" pageNumber="167">Remarks.</emphasis>
The new species superficially looks much like
<taxonomicName id="4C5046FDFF9006395A192EF9FC30F8F6" authorityName="Park" authorityYear="2007" box="[845,1020,1787,1812]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF9006395A192EF9FC30F8F6" box="[845,1020,1787,1812]" italics="true" pageId="2" pageNumber="167">T. sumatraensis</emphasis>
</taxonomicName>
, but has the hindwing dark line more curved (straighter in
<taxonomicName id="4C5046FDFF90063958D62F1DFDFEF8DA" authorityName="Park" authorityYear="2007" box="[386,562,1823,1848]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF90063958D62F1DFDFEF8DA" box="[386,562,1823,1848]" italics="true" pageId="2" pageNumber="167">T. sumatraensis</emphasis>
</taxonomicName>
) and closed to the dorsal margin (central in the hindwing of
<taxonomicName id="4C5046FDFF9006395D882F1DFA40F8DA" authorityName="Park" authorityYear="2007" box="[1244,1420,1823,1848]" class="Insecta" family="Lecithoceridae" genus="Tisis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="B924E16CFF9006395D882F1DFA40F8DA" box="[1244,1420,1823,1848]" italics="true" pageId="2" pageNumber="167">T. sumatraensis</emphasis>
</taxonomicName>
). Genital characters are more reliable for species discernment between these two species as shown in diagnosis.
</paragraph>
</subSubSection>
</treatment>
</document>
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